Archaeothyris is an extinct genus of ophiacodontid synapsid that lived during the Late Carboniferous and is known from Nova Scotia. Dated to 306 million years ago, Archaeothyris, along with a more poorly known synapsid called Echinerpeton, are the oldest undisputed synapsids known. The name means ancient window (Greek), and refers to the opening in the skull, the temporal fenestra, which indicates this is an early synapsid. Protoclepsydrops also from Nova Scotia is slightly older but is known by very fragmentary materials.

Life restoration of Archaeothyris florensis

Description

Archaeothyris was more advanced than the early sauropsids, having strong jaws that could open wider than those of the early reptiles. While its sharp teeth were all of the same size & shape, it did possess a pair of enlarged canines, suggesting that it was a carnivore.

Archaeothyris's legs were articulated laterally at its pelvis and shoulders, which gave it a sprawling stance.[citation needed] The first toe is smaller than the second.[citation needed]

Classification

Archaeothyris belonged to the family Ophiacodontidae, a group of early pelycosaurs that evolved early in the Late Carboniferous. It was one of the earliest and most basal synapsids (the group which includes mammals).

Below is a cladogram modified from the analysis of Benson (2012):

Tseajaia campi Limnoscelis paludis Amniota Captorhinus spp. Protorothyris archeri Synapsida Caseasauria Ianthodon schultzei Edaphosauridae Sphenacodontia Varanopidae Ophiacodontidae Archaeothyris florensis Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Tseajaia campi
Limnoscelis paludis
AmniotaCaptorhinus spp. Protorothyris archeri Synapsida Caseasauria Ianthodon schultzei Edaphosauridae Sphenacodontia Varanopidae Ophiacodontidae Archaeothyris florensis Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Captorhinus spp. Protorothyris archeri
Captorhinus spp.
Protorothyris archeri
SynapsidaCaseasauria Ianthodon schultzei Edaphosauridae Sphenacodontia Varanopidae Ophiacodontidae Archaeothyris florensis Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Caseasauria Ianthodon schultzei Edaphosauridae Sphenacodontia
Caseasauria
Ianthodon schultzei Edaphosauridae Sphenacodontia
Ianthodon schultzei
Edaphosauridae Sphenacodontia
Edaphosauridae
Sphenacodontia
Varanopidae Ophiacodontidae Archaeothyris florensis Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Varanopidae
OphiacodontidaeArchaeothyris florensis Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Archaeothyris florensis
Varanosaurus acutirostris Ophiacodon spp. Stereophallodon ciscoensis
Varanosaurus acutirostris
Ophiacodon spp. Stereophallodon ciscoensis
Ophiacodon spp.
Stereophallodon ciscoensis

Discovery and paleoecology

Fossils of Archaeothyris were first described in 1972 from the Joggins Fossil Cliffs, the same locality in which the early reptiles Hylonomus and Petrolacosaurus (both of which resemble Archaeothyris) were found.

Archaeothyris lived in what is now Nova Scotia, about 306 million years ago in the Carboniferous Period (Pennsylvanian). Nova Scotia at this time was a coal forest. The "trees" (actually giant club mosses) were very tall, some, such as Lepidodendron, up to 50 metres (164 ft) tall. Archaeothyris and the other early amniotes lived in the moist vegetation on the forest ground, together with the more terrestrially adapted labyrinthodont amphibians.

See also

Further reading

  • Kemp, T. S. (2005). The Origin & Evolution of Mammals. Oxford: Oxford University Press. ISBN 978-0-19-850761-1.

External links

  • - includes description of important transitional genera from reptile to mammal (includes a little information about Archaeothyris)