Bacilli is a taxonomic class of bacteria that includes two orders, Bacillales and Lactobacillales, which contain several well-known pathogens such as Bacillus anthracis (the cause of anthrax). Bacilli are almost exclusively gram-positive bacteria.

The name Bacillus, capitalized and italicized, refers to a specific genus of bacteria. The name Bacilli, capitalized but not italicized, can also refer to a less specific taxonomic group of bacteria that includes two orders, one of which contains the genus Bacillus. When the word is formatted with lowercase and not italicized, 'bacillus', it will most likely be referring to shape and not to the genus at all.

Ambiguity

Several related concepts make use of similar words, and the ambiguity can create considerable confusion. The term "Bacillus" (capitalized and italicized) is also the name of a genus (Bacillus anthracis) that, among many other genera, falls within the class Bacilli.

The word "bacillus" (or its plural "bacilli", with a small b) is also a generic term to describe the morphology of any rod-shaped bacterium. This general term does not mean that the subject is a member of class Bacilli or genus Bacillus. Thus, it does not necessarily imply a similar group of characteristics. Not all members of class Bacilli are rod-shaped (Staphylococcus is spherical), and many other rod-shaped bacteria that do not fall within that class exist (e.g., Clostridium is rod-shaped but very different taxonomically). Moreover, the general term "bacillus" does not necessarily indicate the Gram-positive staining common to class Bacilli. For example, E. coli is a rod-shaped bacterium that can be described as "a bacillus", but it stains Gram-negative and does not belong to the genus Bacillus or the class Bacilli. Some microbiologists have forsaken the general "bacillus" term because of the confusion it can create.

Phylogeny

The currently accepted taxonomy based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and the National Center for Biotechnology Information (NCBI).

16S rRNA based LTP_10_2024120 marker proteins based GTDB 09-RS220
"Bacillia" Hydrogenibacillus Thermicanales Alicyclobacillales (incl. Kyrpidiales, Tumebacillales) Paenibacillales Thermoactinomycetales Novibacillaceae Caldalkalibacillales Caldibacillaceae Calditerricolales Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales "Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales {Bacillota}Mycoplasmatota Mollicutes "Aphodocolales" "Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales] "Caccosomales" Erysipelotrichales Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales] "Bacillia" "Pasteuriales" Kyrpidiales Tumebacillales Alicyclobacillales Desulfuribacillales Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales" Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales {Bacillota}
"Bacillia"Hydrogenibacillus Thermicanales Alicyclobacillales (incl. Kyrpidiales, Tumebacillales) Paenibacillales Thermoactinomycetales Novibacillaceae Caldalkalibacillales Caldibacillaceae Calditerricolales Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales "Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Hydrogenibacillus Thermicanales Alicyclobacillales (incl. Kyrpidiales, Tumebacillales) Paenibacillales
Hydrogenibacillus Thermicanales
Hydrogenibacillus
Thermicanales
Alicyclobacillales (incl. Kyrpidiales, Tumebacillales) Paenibacillales
Alicyclobacillales (incl. Kyrpidiales, Tumebacillales)
Paenibacillales
Thermoactinomycetales Novibacillaceae Caldalkalibacillales Caldibacillaceae Calditerricolales Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales "Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Thermoactinomycetales
Novibacillaceae Caldalkalibacillales Caldibacillaceae Calditerricolales Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales "Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Novibacillaceae Caldalkalibacillales Caldibacillaceae Calditerricolales
Novibacillaceae
Caldalkalibacillales Caldibacillaceae Calditerricolales
Caldalkalibacillales
Caldibacillaceae Calditerricolales
Caldibacillaceae
Calditerricolales
Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales "Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Microaerobacter Desulfuribacillales Tepidibacillales Oxalophagaceae Aneurinibacillales Brevibacillales
Microaerobacter Desulfuribacillales Tepidibacillales
Microaerobacter
Desulfuribacillales Tepidibacillales
Desulfuribacillales
Tepidibacillales
Oxalophagaceae Aneurinibacillales Brevibacillales
Oxalophagaceae
Aneurinibacillales Brevibacillales
Aneurinibacillales
Brevibacillales
"Bacillus thermozeamaize" Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
"Bacillus thermozeamaize"
Bacillales (incl. Planococcaceae) Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Bacillales (incl. Planococcaceae)
Listeriaceae Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Listeriaceae
Exiguobacteriales Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Exiguobacteriales
Staphylococcales Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Staphylococcales
Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales Lactobacillales
Gemellaceae Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales
Gemellaceae
Culicoidibacterales Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales
Culicoidibacterales
Turicibacteraceae Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales
Turicibacteraceae
Haloplasmatales Acholeplasmatales Erysipelotrichales Mycoplasmatales
Haloplasmatales
Acholeplasmatales Erysipelotrichales Mycoplasmatales
Acholeplasmatales
Erysipelotrichales Mycoplasmatales
Erysipelotrichales
Mycoplasmatales
Lactobacillales
{Bacillota}
MycoplasmatotaMollicutes "Aphodocolales" "Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales] "Caccosomales" Erysipelotrichales Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales]
Mollicutes"Aphodocolales" "Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales] "Caccosomales" Erysipelotrichales Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales]
"Aphodocolales"
"Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales] "Caccosomales" Erysipelotrichales Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales]
"Bathyoplasmales" "Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales]
"Bathyoplasmales"
"Izemoplasmatales" Acholeplasmatales [incl.Anaeroplasmatales]
"Izemoplasmatales"
Acholeplasmatales [incl.Anaeroplasmatales]
"Caccosomales" Erysipelotrichales Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales]
"Caccosomales" Erysipelotrichales
"Caccosomales"
Erysipelotrichales
Mycoplasmatales [incl. Entomoplasmatales; Mycoplasmoidales]
"Bacillia""Pasteuriales" Kyrpidiales Tumebacillales Alicyclobacillales Desulfuribacillales Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales" Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
"Pasteuriales"
Kyrpidiales Tumebacillales Alicyclobacillales Desulfuribacillales Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales" Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Kyrpidiales Tumebacillales Alicyclobacillales
Kyrpidiales
Tumebacillales Alicyclobacillales
Tumebacillales
Alicyclobacillales
Desulfuribacillales Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales" Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Desulfuribacillales
Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales" Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Calditerricolales "Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales"
Calditerricolales
"Bacillus thermozeamaize" {Bacillales_F} Thermicanales "Brockiales"
"Bacillus thermozeamaize" {Bacillales_F}
Thermicanales "Brockiales"
Thermicanales
"Brockiales"
Thermoactinomycetales Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Thermoactinomycetales
Paenibacillales Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Paenibacillales
Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Microaerobacter {DSM-22679} Aneurinibacillales Brevibacillales
Microaerobacter {DSM-22679}
Aneurinibacillales Brevibacillales
Aneurinibacillales
Brevibacillales
Tepidibacillales "Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Tepidibacillales
"Rubeoparvulales" Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
"Rubeoparvulales"
Caldalkalibacillales Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Caldalkalibacillales
Exiguobacteriales "Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Exiguobacteriales
"Harrysmithimonadales" "Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
"Harrysmithimonadales"
"Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
"Bacillidae" Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
"Bacillidae"Bacillales_B Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillales_B
Bacillales Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillales
Bacillales_C Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillales_C
Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G Bacillales_H
Bacillales_E Sporolactobacillaceae {Bacillales_K} Bacillales_G
Bacillales_E
Sporolactobacillaceae {Bacillales_K} Bacillales_G
Sporolactobacillaceae {Bacillales_K}
Bacillales_G
Bacillales_H
Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697} Bacillales_D
Bacillus marinisedimentorum {Bacillales_I} Aureibacillaceae {DSM-28697}
Bacillus marinisedimentorum {Bacillales_I}
Aureibacillaceae {DSM-28697}
Bacillales_D
Caryophanaceae {Caryophanales} Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Caryophanaceae {Caryophanales}
Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales Lactobacillales
Staphylococcales Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales
Staphylococcales
Culicoidibacterales Turicibacteraceae {MOL361} Haloplasmatales
Culicoidibacterales
Turicibacteraceae {MOL361} Haloplasmatales
Turicibacteraceae {MOL361}
Haloplasmatales
Lactobacillales
{Bacillota}

See also