Chorrillo Formation
Chorrillo Formation
Stratigraphic range : Maastrichtian PreꞒ Ꞓ O S D C P T J K Pg N
Underlies Calafate Formation
Thickness >50m (160ft)
Lithology
Location
Coordinates 49°54′S 72°30′W/49.9°S 72.5°W/ -49.9; -72.5
Approximate paleocoordinates 51°48′S 62°12′W/51.8°S 62.2°W/ -51.8; -62.2
Country Argentina
Type section
Named by Arbe & Hechem
Year defined 1984
The Chorrillo Formation , also named as Chorillo Formation , is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia , Argentina . The formation is more than 50 metres (160ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation . It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin , located in Santa Cruz Province , Argentina, approximately 30km southwest of El Calafate . Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage. The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units (e.g., Allen Formation , Los Alamitos Formation ), which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.
Geology The Chorrillo Formation is embedded within the foreland Magallanes Basin , shaped by Late Cretaceous Andean orogeny driven by subduction along the Pacific margin. Tectonic subsidence, coupled with sediment supply from the rising Patagonian Andes, facilitated deposition. Lithologically, the unit comprises fine- to coarse-grained clastics: mudstones and siltstones dominate floodplain facies, often organic-rich and pedogenically modified; sandstones, arkosic to lithic, form channel fills with cross-bedding; and conglomerates appear as basal lags. Clay mineralogy is dominated by smectite, with minor illite and kaolinite, reflecting volcanic ash alteration. Sedimentologically, it records a meandering fluvial system on a distal floodplain, featuring fining-upward channel sequences, crevasse splays, and overbank deposits. High-frequency paleosol stacking indicates local hydrologic and topographic variability, while low-frequency patterns suggest avulsion-driven progradation. Regional ties extend to the Dorotea Formation in Chile, reflecting basin-wide tectonic influences.
Paleoenvironment Modern equivalent, Valdivian Rainforest Paleoenvironmental reconstructions indicate a temperate-warm, seasonally humid climate with mean annual temperatures of 10-18°C and precipitation of 900–1500mm, inferred from paleosols, clay mineralogy, and biotic indicators. Depositional settings encompass low-energy fluvial floodplains with paludal and lacustrine elements, marked by poor drainage, hydromorphic paleosols (e.g., vertisols , histosols ), and redoximorphic features such as mottling and iron nodules, suggesting fluctuating water tables and episodic anoxia. Aquatic communities feature Nymphaeaceae , Salviniaceae , Salviniaceae , and Zygnemataceae , indicative of freshwater ponds and swamps, while terrestrial elements include conifers, ferns, and angiosperms in riparian zones. Floral phytozones, derived from palynological data, comprise five assemblages across the section, exhibiting quantitative variations in taxa proportions without major turnover: lower zones are fern-dominated (e.g., Cyatheaceae ), transitioning to angiosperm-rich upper zones (e.g., Proteaceae , Nothofagidites ), attributed to subtle humidity fluctuations or habitat gradients relative to fluvial channels. These patterns align with Maastrichtian assemblages in southern Gondwanan basins (e.g., La Anita, Lefipán Formation ), reflecting angiosperm diversification and Weddellian Province expansion.
Fossils Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Arthropoda Decapoda Genus Species Locality Material Notes Images Astacidea Indeterminate "Locality 4" MPM 21516, three gastroliths Previously identified as crushing fish teeth Extant example
Insects Genus Species Locality Material Notes Images Coelolepida Indeterminate La Anita farm MPM-Pal 21835-3:W34/1; MPM-Pal 21835-23:W55/0; MPM-Pal 21835-22:R52/2; MPM-Pal 21835-20:Z47/3; MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3 A Glossatan Lepidopteran Diamesinae "Morphotype 3" La Anita farm MPM-Pal 21835-9:E34/4; MPM-Pal 21835-19: V34/4 A Chironomidae Nematoceran . Similar to the living genus Paraheptagyia Extant example Ephemeroptera Indeterminate La Anita farm MPM-Pal 21835-9:W37/1 A Mayfly Lepidoptera Indeterminate La Anita farm MPM-Pal 21835-10:X30/2; MPM-Pal 21835-7:S27/2 Orthocladiinae "Morphotype 1" La Anita farm MPM-Pal 21835-24: M31/0; MPM-Pal 21835-3: P46/4 A Chironomidae Nematoceran "Morphotype 2" La Anita farm MPM-Pal 21835-25: V26/3 "Morphotype 5" La Anita farm MPM-Pal 21835-9: Q58/3 It resembles modern Cricotopus Cricotopus Tanypodinae "Morphotype 4" La Anita farm MPM-Pal 21835-2:Y40/3 A Chironomidae Nematoceran Extant example
Molluscs Gastropods Genus Species Locality Material Notes Images Allopeas A. agnolini "Locality 4" Shells An achatinid gastropod Extant specimen Bocourtia B. leonardodavincii "Locality 4" Shells A bulimulid gastropod Extant specimen Coelostemma C. patagonica "Locality 4" Shells A holospirid gastropod Holospira H. sp."Locality 4" Shells An urocoptidae gastropod Leptinaria Cf.L. sp. "Locality 4" Shells An achatinid gastropod Extant specimen Lilloiconcha L. novasi "Locality 4" Shells A charopid gastropod Naesiotus Cf.N. sp. "Locality 4" Shells A bulimulid gastropod Extant specimen Physa P. sp."Locality 4" Shells A physidae gastropod Extant specimen Physella Cf.P. sp. "Locality 4" Shells A physidae gastropod Extant specimen Pleuroceridae Indeterminate "Locality 4" Shells Incertade Sedis Pomacea P. sp."Locality 4" Shells An ampullariidae gastropod Extant specimen Potamolithus P. fabiani "Locality 4" Shells A tateid gastropod Extant specimen Stenophysa S. sp."Locality 4" Shells A physidae gastropod
Bivalves Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Locality Material Notes Images Chlamys "C." salamanca "Locality 5" Shells A Pectinidae bivalve Extant specimen Cubitostrea C. ameghinoi "Locality 5" Shells An Ostreidae bivalve Extant specimen Gryphaeostrea G. callophyla "Locality 5" Shells A Gryphaeidae bivalve
Fishes Genus Species Locality Material Notes Images Lamniformes Indeterminate "Locality 5" Isolated upper lateral tooth Similar to Cretalamna . Vidalamiinae Indeterminate "Locality 2" Isolated tooth An amiid fish
Amphibians Genus Species Locality Material Notes Images Anura Indeterminate "Locality 4" MPM 21518, isolated tibiofibula Calyptocephalella C. gayi "Locality 2" MPM-PV-22844, proximal right radioulna; MPM-PV-22845, left proximal end of tibiofibula; MPM-PV-22846, proximal half of urostyle. Calyptocephalellidae frogExtant specimen of C. gayi C. sp. Magallanodon Sitemaxillae MPM-PV-22841 and MPM-PV-22842; ilium MPM-PV-22843 A species very small in size, distinct from C. gayi . Calyptocephalellidae Indeterminate "Locality 4" MPM 21519, distal end of right humerus with eroded distal margin Pipoidea Indeterminate "Locality 2" Distal end of right humerus Resembles, yet is clearly different, from Xenopus .
Plesiosaurs Genus Species Locality Material Notes Images Elasmosauridae Indeterminate "Locality 5" Incomplete cervical centrum and the lateral half of a very fragmentary dorsal vertebra May belong to the Calafate Formation
Lepidosaurs Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Mosasaurs Genus Species Locality Material Notes Images Mosasauridae Indeterminate "Locality 5" Seven teeth Its presence suggest marine influences
Rhynchocephalian Genus Species Locality Material Notes Images Notosphenos N. finisterre "Monotreme site" Incomplete right maxilla preserving two completeand one partial teeth. A sphenodontid rhynchocephalian
Snakes Genus Species Locality Material Notes Images Alethinophidia Indeterminate "Locality 4" MPM 21522 small vertebra Anilioidea Indeterminate "Locality 2" MPM-PV-22847, possible mid-precloacal vertebra Coniophiidae Indeterminate "Monotreme site" Nearly complete mid-posterior precloacal vertebra; Incomplete posterior precloacal vertebra Shares several features with Coniophis . Madtsoiidae Indeterminate "Monotreme site"; "Green Valley site" Incomplete mid-posterior precloacal vertebralacking most of the neural arch; Poorly preserved precloacal vertebra preserving the verteb-ral centrum Rionegrophis R. madtsoioides "Locality 2" A partial vertebra. A madtsoiid snake
Turtles Genus Species Locality Material Notes Images Chelidae Indeterminate "Locality 2" MPM 21520, two peripheral and one costal plates Largest plates suggest a maximum carapace length of 40cm. Hydromedusinae Indeterminate "Locality 2, 4, 5" Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragment Similar to Hydromedusa . Modern Hydromedusa Meiolaniformes Indeterminate "Locality 4" Distal end of a left humerus Southernmost record for the clade in South America
Archosaurs Crocodylomorphs Genus Species Locality Material Notes Images Kostensuchus K. atrox Estancia La Anita Skull, jaws, incomplete postcranial skeleton A large-bodied, broad-snouted peirosaurid
Ornithischians Genus Species Locality Material Notes Images Hadrosauridae Indeterminate La Anita Farm Two incomplete caudal centra Isasicursor I. santacrucensis "Locality 5" Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbs An elasmarian ornithopod Isasicursor skeletal Ornithopoda Indeterminate La Anita Farm Proximal end of a femur A small sized ornithopod Parankylosauria Indeterminate La Anita Farm Five fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms
Sauropods Genus Species Locality Material Notes Images Fusioolithus F. sp. "Locality 2" Eggshells Fusioolithidae oogenus Lithostrotia Indeterminate Río Leona MACN 5205 (3 middle–posterior caudal centra) Nullotitan N. glaciaris Estancia Alta Vista; "Locality 1, 2, 4, 5" A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle bone A colossosaurian titanosaur Titanosauria Indeterminate "Locality 2, 4" Seven isolated teeth
Theropods Genus Species Locality Material Notes Enantiornithes Indeterminate "Locality 2, 4" Distal fragment of a pedal phalanx; Two isolated ungual pedal phalanges Kank K. australis "Monotreme site" Fragmentary remains, including vertebrae, teeth, and pedal phalanges A unenlagiid theropod Kookne K. yeutensis "Locality 2" An incomplete right coracoid, lacking the sternal end and the proximal end damaged An ornithuran bird similar to Anseriformes Maip M. macrothorax La Anita Farm Axis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsal The largest megaraptorid theropod of South America and possibly the largest member of this group M. macrothorax Megaraptoridae Indeterminate La Anita Farm Isolated teeth Noasauridae Indeterminate "Locality 4" Isolated right pedal phalanx IV-2? Similar to Velocisaurus and Vespersaurus Prismatoolithidae Indeterminate "Locality 4" Eggshells Theropod eggshells Unenlagiidae Indeterminate "Locality 4" An isolated dorsal vertebra, pedal ungual of digit II and phalanx III-2 Similar to Neuquenraptor Yatenavis Y. ieujensis Estancia La Anita Distal right humerus An enantiornithine bird Yatenavis humerus
Mammals Genus Species Locality Material Notes Images Magallanodon M. baikashkenke Río de Las Chinas Valley and La Anita Farm Tentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right side A ferugliotheriid gondwanathere M. terrwerr "Locality 2" MPM-PV-23088, isolated right lower molariform; MPM-PV-23089, isolated right lower molariform A ferugliotheriid gondwanathere Mammalia Indeterminate "Locality 4" Two isolated caudal vertebrae Characters shared with mammals having long and gracile tails. Meridiolestida Indeterminate "Monotreme site" MPM-PV-21707, two fragments of a right dentary Small sized taxon similar to Reigitherium . Mesungulatoidea Indeterminate La Anita Farm MPM-PV-21708, fragment of left dentary with five alveoli corresponding with three teeth. Larger than any known Mesozoic meridiolestidan. Orretherium O. tzei "Monotreme site" MPM-PV-23514, left upper second molariform. A member of Meridiolestida belonging to the family Mesungulatidae . Patagorhynchus P. pascuali "Monotreme site" Lower right molar attached to a fragment of the dentary A ornithorhynchid monotreme Patagorhynchus holotypePatagomaia P. chainko La Anita Farm Partial left acetabulum and ischium; MPM-PV-23515, one fragment of thoracic vertebra, posterior half of one caudal vertebra and 38 indeterminate bones. A large therian Patagomaia holotype
Flora Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Algae Genus Species Affinity / Family Image Botryococcus B. sp. Botryococcaceae Cymatiosphaera C. sp. Zygnemataceae Leiosphaeridia L. sp. Zygnemataceae Ovoidites O. parvus Zygnemataceae O. spriggii Zygnemataceae Schizosporis S. reticulatus Zygnemataceae (aff. Spirogyra )Sphaeroplea S. striatocristata Sphaeropleaceae
Liverworts Genus Species Affinity / Family Image Aequitriradites A. spinulosus Marchantiidae Stoverisporites S. microverrucatus Marchantiidae Taurocusporites T. segmentatus Marchantiidae Triporoletes T. reticulatus Marchantiidae T. sp. Marchantiidae
Hornworts Genus Species Affinity / Family Image Foraminisporis F. wonthaggiensis Anthocerotidae F. sp. Anthocerotidae
Lycopods Genus Species Affinity / Family Image Camarozonosporites C. ambigens Selaginellales Ceratosporites C. equalis Selaginellales Densoisporites D. velatus Selaginellales Foveosporites F. sp. Selaginellales Retitriletes R. austroclavatidites Lycopodiales R. sp. cf. R. douglasii Lycopodiales
Ferns Genus Species Affinity / Family Image Azolla A. cf. andreisii Salviniaceae A. sp. Salviniaceae Baculatisporites B. comaumensis Osmundaceae B. kachaikensis Osmundaceae B. turbioensis Osmundaceae cf. Cladophlebis C. sp. Polypodiopsida Clavifera C. sp. Gleicheniaceae Cyatheacidites C. annulatus Dicksoniaceae (aff. Lophosoria )Cyathidites C. australis Cyatheaceae C. minor Cyatheaceae C. rafaeli Cyatheaceae Dictyophyllidites D. sp. cf. D. concavus Dipteridaceae / Matoniaceae Gleicheniidites G. sp. Gleicheniaceae Leptolepidites L. verrucatus Blechnaceae Ornamentifera O. sp. Gleicheniaceae Osmundacidites O. diazii Osmundaceae Peromonolites P. vellosus Blechnaceae
Non-conifer Seed Plants Genus Species Affinity / Family Image Cycadopites C. follicularis Cycadales , Bennettitales , or Ginkgoales Ephedripites E. ambiguous Ephedrales Equisetosporites E. notensis Ephedrales E. sp. cf. E. dudarensis Ephedrales Vitreisporites V. signatus Pteridosperms
Conifers Genus Species Affinity / Family Image Araucariacites A. australis Araucariaceae Callialasporites C. turbatus Araucariaceae Classopollis C. sp. Cheirolepidiaceae Cyclusphaera C. psilata Araucariaceae Dacrycarpites D. australiensis Podocarpaceae Dilwynites D. tuberculatus Araucariaceae Gamerroites G. psilasaccus Podocarpaceae Lygistepollenites L. florinii Podocarpaceae Microcachryidites M. antarcticus Podocarpaceae (aff. Microcachrys )Microcachrys Peripollenites P. sp. Cupressaceae Phyllocladidites P. mawsonii Podocarpaceae (aff. Lagarostrobos franklinii )Lagarostrobos Podocarpidites P. elegans Podocarpaceae P. elipticus Podocarpaceae P. exiguus Podocarpaceae P. herbstii Podocarpaceae P. major Podocarpaceae P. marwikii Podocarpaceae P. otagoensis Podocarpaceae P. sp. cf. P. langii Podocarpaceae P. sp. cf. P. microreticuloidata Podocarpaceae P. torquatus Podocarpaceae P. verrucosus Podocarpaceae Podocarpoxylon P. dusenii Podocarpaceae Trichotomosulcites T. subgranulatus Podocarpaceae
Magnoliids Genus Species Affinity / Family Image Clavatimonocolpites C. sp. Magnoliids
Monocots Genus Species Affinity / Family Image Arecipites A. minutiscabratus Arecaceae A. sp. cf. A. subverrucatus Arecaceae Aff.Eolirion "Morphotype 5" Palmae Longapertites L. proxapertitoides Arecaceae Liliacidites L. lanceolatus Liliales (aff. Lilium )Lilium L. variegatus Liliales (aff. Lilium )L. vermireticulatus Liliales L. sp. cf. L. kaitangataensis Liliaceae /Amaryllidaceae Monocotyledonean "Morphotype 6" Monocots Nuphar N.? sp. 1 Nymphaeaceae Extant specimen N.? sp. 2 Nymphaeaceae Pandaniidites P. typicus Araceae Proxapertites P. operculatus Araceae Spinozonocolpites S. hialinus Arecaceae
Eudicots Genus Species Affinity / Family Image Beauprea B. elegansiformis Proteales Beauprea drawingAff. Brachychiton "Morphotype 4" Malvaceae Extant specimen Buttinia B. andreevii Buxales Clavamonocolpites C. sp. ?Chloranthaceae Coccoloba ?"Morphotype 1" Polygonaceae Extant specimen Constantinisporis C. jacquei ?Chloranthaceae Dicotetradites D. sp. Ericales Diporites D. sp. ?Chloranthaceae Doyleipollenites D. sp. ?Chloranthaceae Ericipites E. scabratus Ericales Eudicots "Morphotype 3" Eudicots Myrtaceidites M. sp. cf. M. mesonesus Myrtales Nothofagidites N. saraensis Nothofagaceae (cf. Nothofagus )Nothofagus Nyssapollenites N. sp. ?Chloranthaceae Peninsulapollis P. gillii Proteaceae (cf. Beauprea ?)P. truswelliae Proteaceae (cf. Beauprea ?)Polycolpites P. sp. Pedaliaceae (?cf. Josephinia )Proteacidites P. parvus Proteaceae (cf. Bellendena montana )Bellendena P. subscabratus Proteaceae P. sp. cf. P. tenuiexinus Proteaceae P. sp. Proteaceae Psilatricolporites P. sp. cf. P. complanatius Tetrameristaceae P. sp. Tetrameristaceae Retidiporites R. camachoi Proteaceae Rhoipites R. baculatus Ericales R. minusculus Ericales R. sp. cf. R. alveolatus Ericales R. sp. 1 Ericales R. sp. 2 Ericales Rousea R. patagonica Ericales R. sp. cf. R. minuscula Ericales R. sp. Ericales Syncolporites S. sp. Ericales Tetracolpites T. sp. ?Chloranthaceae Triatriopollenites T. lateflexus Proteaceae Tricesticillus T. americanus ?Chloranthaceae Tricolpites T. reticulatus Gunneraceae (Gunnera macrophylla )Gunnera T. sp. cf. T. brevicolpus Gunneraceae T. sp. cf. T. inargutus Gunneraceae T. sp. cf. T. lumbrerensis Gunneraceae T. sp. 1 Ranunculaceae (cf. Clematis )Clematis Tricolporites T. sp. 2 Ericales T. sp. 3 Ericales Tricolporoidites T. sp. cf. T. eximius Ericales Triorites T. sp. cf. T. minusculus ?Chloranthaceae Tubulifloridites T. lilliei Ericales Volkheimerites V. labyrinthus ?Chloranthaceae
See also Bibliography Marenssi, S. A.; Casadío, S.; Santillana, S. N. (September 2003). [Stratigraphy and sedimentology of the Upper Cretaceous-Paleogene units outcropping on the southeastern margin of Lake Viedma, Santa Cruz Province, Argentina]. Revista de la Asociación Geológica Argentina (in Spanish). 58 (3): 403–416.