The Chorrillo Formation, also named as Chorillo Formation, is a Maastrichtian (Late Cretaceous Epoch, 72.1–66 million years ago) geologic formation in southern Patagonia, Argentina. The formation is more than 50 metres (160ft) thick and underlies the Calafate Formation and rests on top of the La Irene Formation. It constitutes a significant uppermost Cretaceous continental sedimentary sequence within the Magallanes Basin, located in Santa Cruz Province, Argentina, approximately 30km southwest of El Calafate. Initial investigations in the 1990s laid the groundwork, with detailed paleontological studies emerging since 2019, revealing a rich biotic assemblage. The formation correlates with the lower Dorotea Formation in Chile, sharing dinosaurian ecotypes, while both are clearly different from other northern Patagonian Maastrichtian units (e.g., Allen Formation, Los Alamitos Formation), which are dominated by saltasaurine sauropods and abelisaurids, suggesting potential temporal or environmental distinctions.

Geology

The Chorrillo Formation is embedded within the foreland Magallanes Basin, shaped by Late Cretaceous Andean orogeny driven by subduction along the Pacific margin. Tectonic subsidence, coupled with sediment supply from the rising Patagonian Andes, facilitated deposition. Lithologically, the unit comprises fine- to coarse-grained clastics: mudstones and siltstones dominate floodplain facies, often organic-rich and pedogenically modified; sandstones, arkosic to lithic, form channel fills with cross-bedding; and conglomerates appear as basal lags. Clay mineralogy is dominated by smectite, with minor illite and kaolinite, reflecting volcanic ash alteration. Sedimentologically, it records a meandering fluvial system on a distal floodplain, featuring fining-upward channel sequences, crevasse splays, and overbank deposits. High-frequency paleosol stacking indicates local hydrologic and topographic variability, while low-frequency patterns suggest avulsion-driven progradation. Regional ties extend to the Dorotea Formation in Chile, reflecting basin-wide tectonic influences.

Paleoenvironment

Modern equivalent, Valdivian Rainforest

Paleoenvironmental reconstructions indicate a temperate-warm, seasonally humid climate with mean annual temperatures of 10-18°C and precipitation of 900–1500mm, inferred from paleosols, clay mineralogy, and biotic indicators. Depositional settings encompass low-energy fluvial floodplains with paludal and lacustrine elements, marked by poor drainage, hydromorphic paleosols (e.g., vertisols, histosols), and redoximorphic features such as mottling and iron nodules, suggesting fluctuating water tables and episodic anoxia. Aquatic communities feature Nymphaeaceae, Salviniaceae, Salviniaceae, and Zygnemataceae, indicative of freshwater ponds and swamps, while terrestrial elements include conifers, ferns, and angiosperms in riparian zones. Floral phytozones, derived from palynological data, comprise five assemblages across the section, exhibiting quantitative variations in taxa proportions without major turnover: lower zones are fern-dominated (e.g., Cyatheaceae), transitioning to angiosperm-rich upper zones (e.g., Proteaceae, Nothofagidites), attributed to subtle humidity fluctuations or habitat gradients relative to fluvial channels. These patterns align with Maastrichtian assemblages in southern Gondwanan basins (e.g., La Anita, Lefipán Formation), reflecting angiosperm diversification and Weddellian Province expansion.

Fossils

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Arthropoda

Decapoda

GenusSpeciesLocalityMaterialNotesImages
AstacideaIndeterminate"Locality 4"MPM 21516, three gastrolithsPreviously identified as crushing fish teethExtant example

Insects

GenusSpeciesLocalityMaterialNotesImages
CoelolepidaIndeterminateLa Anita farmMPM-Pal 21835-3:W34/1; MPM-Pal 21835-23:W55/0; MPM-Pal 21835-22:R52/2; MPM-Pal 21835-20:Z47/3; MPM-Pal 21835-22: G55/0; MPM-Pal 2183-21: E42/3A Glossatan Lepidopteran
Diamesinae"Morphotype 3"La Anita farmMPM-Pal 21835-9:E34/4; MPM-Pal 21835-19: V34/4A Chironomidae Nematoceran. Similar to the living genus ParaheptagyiaExtant example
EphemeropteraIndeterminateLa Anita farmMPM-Pal 21835-9:W37/1A Mayfly
LepidopteraIndeterminateLa Anita farmMPM-Pal 21835-10:X30/2; MPM-Pal 21835-7:S27/2
Orthocladiinae"Morphotype 1"La Anita farmMPM-Pal 21835-24: M31/0; MPM-Pal 21835-3: P46/4A Chironomidae Nematoceran
"Morphotype 2"La Anita farmMPM-Pal 21835-25: V26/3
"Morphotype 5"La Anita farmMPM-Pal 21835-9: Q58/3It resembles modern CricotopusCricotopus
Tanypodinae"Morphotype 4"La Anita farmMPM-Pal 21835-2:Y40/3A Chironomidae NematoceranExtant example

Molluscs

Gastropods

GenusSpeciesLocalityMaterialNotesImages
AllopeasA. agnolini"Locality 4"ShellsAn achatinid gastropodExtant specimen
BocourtiaB. leonardodavincii"Locality 4"ShellsA bulimulid gastropodExtant specimen
CoelostemmaC. patagonica"Locality 4"ShellsA holospirid gastropod
HolospiraH. sp."Locality 4"ShellsAn urocoptidae gastropod
LeptinariaCf.L. sp."Locality 4"ShellsAn achatinid gastropodExtant specimen
LilloiconchaL. novasi"Locality 4"ShellsA charopid gastropod
NaesiotusCf.N. sp."Locality 4"ShellsA bulimulid gastropodExtant specimen
PhysaP. sp."Locality 4"ShellsA physidae gastropodExtant specimen
PhysellaCf.P. sp."Locality 4"ShellsA physidae gastropodExtant specimen
PleuroceridaeIndeterminate"Locality 4"ShellsIncertade Sedis
PomaceaP. sp."Locality 4"ShellsAn ampullariidae gastropodExtant specimen
PotamolithusP. fabiani"Locality 4"ShellsA tateid gastropodExtant specimen
StenophysaS. sp."Locality 4"ShellsA physidae gastropod

Bivalves

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesLocalityMaterialNotesImages
Chlamys"C." salamanca"Locality 5"ShellsA Pectinidae bivalveExtant specimen
CubitostreaC. ameghinoi"Locality 5"ShellsAn Ostreidae bivalveExtant specimen
GryphaeostreaG. callophyla"Locality 5"ShellsA Gryphaeidae bivalve

Fishes

GenusSpeciesLocalityMaterialNotesImages
LamniformesIndeterminate"Locality 5"Isolated upper lateral toothSimilar to Cretalamna.
VidalamiinaeIndeterminate"Locality 2"Isolated toothAn amiid fish

Amphibians

GenusSpeciesLocalityMaterialNotesImages
AnuraIndeterminate"Locality 4"MPM 21518, isolated tibiofibula
CalyptocephalellaC. gayi"Locality 2"MPM-PV-22844, proximal right radioulna; MPM-PV-22845, left proximal end of tibiofibula; MPM-PV-22846, proximal half of urostyle.Calyptocephalellidae frogExtant specimen of C. gayi
C. sp.Magallanodon Sitemaxillae MPM-PV-22841 and MPM-PV-22842; ilium MPM-PV-22843A species very small in size, distinct from C. gayi.
CalyptocephalellidaeIndeterminate"Locality 4"MPM 21519, distal end of right humerus with eroded distal margin
PipoideaIndeterminate"Locality 2"Distal end of right humerusResembles, yet is clearly different, from Xenopus.

Plesiosaurs

GenusSpeciesLocalityMaterialNotesImages
ElasmosauridaeIndeterminate"Locality 5"Incomplete cervical centrum and the lateral half of a very fragmentary dorsal vertebraMay belong to the Calafate Formation

Lepidosaurs

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Mosasaurs

GenusSpeciesLocalityMaterialNotesImages
MosasauridaeIndeterminate"Locality 5"Seven teethIts presence suggest marine influences

Rhynchocephalian

GenusSpeciesLocalityMaterialNotesImages
NotosphenosN. finisterre"Monotreme site"Incomplete right maxilla preserving two completeand one partial teeth.A sphenodontid rhynchocephalian

Snakes

GenusSpeciesLocalityMaterialNotesImages
AlethinophidiaIndeterminate"Locality 4"MPM 21522 small vertebra
AnilioideaIndeterminate"Locality 2"MPM-PV-22847, possible mid-precloacal vertebra
ConiophiidaeIndeterminate"Monotreme site"Nearly complete mid-posterior precloacal vertebra; Incomplete posterior precloacal vertebraShares several features with Coniophis.
MadtsoiidaeIndeterminate"Monotreme site"; "Green Valley site"Incomplete mid-posterior precloacal vertebralacking most of the neural arch; Poorly preserved precloacal vertebra preserving the verteb-ral centrum
RionegrophisR. madtsoioides"Locality 2"A partial vertebra.A madtsoiid snake

Turtles

GenusSpeciesLocalityMaterialNotesImages
ChelidaeIndeterminate"Locality 2"MPM 21520, two peripheral and one costal platesLargest plates suggest a maximum carapace length of 40cm.
HydromedusinaeIndeterminate"Locality 2, 4, 5"Left peripheral 3 or 4, bridge peripheral, fragment indet; bridge peripheral, right hypoplastron, buttress, plastral fragment and ?plastral fragmentSimilar to Hydromedusa.Modern Hydromedusa
MeiolaniformesIndeterminate"Locality 4"Distal end of a left humerusSouthernmost record for the clade in South America

Archosaurs

Crocodylomorphs

GenusSpeciesLocalityMaterialNotesImages
KostensuchusK. atroxEstancia La AnitaSkull, jaws, incomplete postcranial skeletonA large-bodied, broad-snouted peirosaurid

Ornithischians

GenusSpeciesLocalityMaterialNotesImages
HadrosauridaeIndeterminateLa Anita FarmTwo incomplete caudal centra
IsasicursorI. santacrucensis"Locality 5"Multiple specimens including teeth, several vertebrae, humerus, thigh bones, shoulder blades, and limbsAn elasmarian ornithopodIsasicursor skeletal
OrnithopodaIndeterminateLa Anita FarmProximal end of a femurA small sized ornithopod
ParankylosauriaIndeterminateLa Anita FarmFive fragmentary caudal vertebrae, incomplete ribs, two incomplete humeri, and nine osteoderms

Sauropods

GenusSpeciesLocalityMaterialNotesImages
FusioolithusF. sp."Locality 2"EggshellsFusioolithidae oogenus
LithostrotiaIndeterminateRío LeonaMACN 5205 (3 middle–posterior caudal centra)
NullotitanN. glaciarisEstancia Alta Vista; "Locality 1, 2, 4, 5"A partial skeleton consisting of a third cervical vertebra, tail vertebrae, a neck rib, ribs, a left shoulder blade, the ends of a right thigh bone, a right shin, a right calf bone, and a right ankle boneA colossosaurian titanosaur
TitanosauriaIndeterminate"Locality 2, 4"Seven isolated teeth

Theropods

GenusSpeciesLocalityMaterialNotes
EnantiornithesIndeterminate"Locality 2, 4"Distal fragment of a pedal phalanx; Two isolated ungual pedal phalanges
KankK. australis"Monotreme site"Fragmentary remains, including vertebrae, teeth, and pedal phalangesA unenlagiid theropod
KookneK. yeutensis"Locality 2"An incomplete right coracoid, lacking the sternal end and the proximal end damagedAn ornithuran bird similar to Anseriformes
MaipM. macrothoraxLa Anita FarmAxis, several dorsal and caudal vertebrae, cervical and dorsal ribs, gastralia, a left coracoid, fragmentary scapula, partial right pubis, and partial metatarsalThe largest megaraptorid theropod of South America and possibly the largest member of this groupM. macrothorax
MegaraptoridaeIndeterminateLa Anita FarmIsolated teeth
NoasauridaeIndeterminate"Locality 4"Isolated right pedal phalanx IV-2?Similar to Velocisaurus and Vespersaurus
PrismatoolithidaeIndeterminate"Locality 4"EggshellsTheropod eggshells
UnenlagiidaeIndeterminate"Locality 4"An isolated dorsal vertebra, pedal ungual of digit II and phalanx III-2Similar to Neuquenraptor
YatenavisY. ieujensisEstancia La AnitaDistal right humerusAn enantiornithine birdYatenavis humerus

Mammals

GenusSpeciesLocalityMaterialNotesImages
MagallanodonM. baikashkenkeRío de Las Chinas Valley and La Anita FarmTentative first upper molariform, anterior end of a left dentary bearing part of its gliriform incisor, and upper mesial incisor from the right sideA ferugliotheriid gondwanathere
M. terrwerr"Locality 2"MPM-PV-23088, isolated right lower molariform; MPM-PV-23089, isolated right lower molariformA ferugliotheriid gondwanathere
MammaliaIndeterminate"Locality 4"Two isolated caudal vertebraeCharacters shared with mammals having long and gracile tails.
MeridiolestidaIndeterminate"Monotreme site"MPM-PV-21707, two fragments of a right dentarySmall sized taxon similar to Reigitherium.
MesungulatoideaIndeterminateLa Anita FarmMPM-PV-21708, fragment of left dentary with five alveoli corresponding with three teeth.Larger than any known Mesozoic meridiolestidan.
OrretheriumO. tzei"Monotreme site"MPM-PV-23514, left upper second molariform.A member of Meridiolestida belonging to the family Mesungulatidae.
PatagorhynchusP. pascuali"Monotreme site"Lower right molar attached to a fragment of the dentaryA ornithorhynchid monotremePatagorhynchus holotype
PatagomaiaP. chainkoLa Anita FarmPartial left acetabulum and ischium; MPM-PV-23515, one fragment of thoracic vertebra, posterior half of one caudal vertebra and 38 indeterminate bones.A large therianPatagomaia holotype

Flora

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Algae

GenusSpeciesAffinity / FamilyImage
BotryococcusB. sp.Botryococcaceae
CymatiosphaeraC. sp.Zygnemataceae
LeiosphaeridiaL. sp.Zygnemataceae
OvoiditesO. parvusZygnemataceae
O. spriggiiZygnemataceae
SchizosporisS. reticulatusZygnemataceae (aff. Spirogyra)
SphaeropleaS. striatocristataSphaeropleaceae

Liverworts

GenusSpeciesAffinity / FamilyImage
AequitriraditesA. spinulosusMarchantiidae
StoverisporitesS. microverrucatusMarchantiidae
TaurocusporitesT. segmentatusMarchantiidae
TriporoletesT. reticulatusMarchantiidae
T. sp.Marchantiidae

Hornworts

GenusSpeciesAffinity / FamilyImage
ForaminisporisF. wonthaggiensisAnthocerotidae
F. sp.Anthocerotidae

Lycopods

GenusSpeciesAffinity / FamilyImage
CamarozonosporitesC. ambigensSelaginellales
CeratosporitesC. equalisSelaginellales
DensoisporitesD. velatusSelaginellales
FoveosporitesF. sp.Selaginellales
RetitriletesR. austroclavatiditesLycopodiales
R. sp. cf. R. douglasiiLycopodiales

Ferns

GenusSpeciesAffinity / FamilyImage
AzollaA. cf. andreisiiSalviniaceae
A. sp.Salviniaceae
BaculatisporitesB. comaumensisOsmundaceae
B. kachaikensisOsmundaceae
B. turbioensisOsmundaceae
cf. CladophlebisC. sp.Polypodiopsida
ClaviferaC. sp.Gleicheniaceae
CyatheaciditesC. annulatusDicksoniaceae (aff. Lophosoria)
CyathiditesC. australisCyatheaceae
C. minorCyatheaceae
C. rafaeliCyatheaceae
DictyophylliditesD. sp. cf. D. concavusDipteridaceae / Matoniaceae
GleicheniiditesG. sp.Gleicheniaceae
LeptolepiditesL. verrucatusBlechnaceae
OrnamentiferaO. sp.Gleicheniaceae
OsmundaciditesO. diaziiOsmundaceae
PeromonolitesP. vellosusBlechnaceae

Non-conifer Seed Plants

GenusSpeciesAffinity / FamilyImage
CycadopitesC. follicularisCycadales, Bennettitales, or Ginkgoales
EphedripitesE. ambiguousEphedrales
EquisetosporitesE. notensisEphedrales
E. sp. cf. E. dudarensisEphedrales
VitreisporitesV. signatusPteridosperms

Conifers

GenusSpeciesAffinity / FamilyImage
AraucariacitesA. australisAraucariaceae
CallialasporitesC. turbatusAraucariaceae
ClassopollisC. sp.Cheirolepidiaceae
CyclusphaeraC. psilataAraucariaceae
DacrycarpitesD. australiensisPodocarpaceae
DilwynitesD. tuberculatusAraucariaceae
GamerroitesG. psilasaccusPodocarpaceae
LygistepollenitesL. floriniiPodocarpaceae
MicrocachryiditesM. antarcticusPodocarpaceae (aff. Microcachrys)Microcachrys
PeripollenitesP. sp.Cupressaceae
PhyllocladiditesP. mawsoniiPodocarpaceae (aff. Lagarostrobos franklinii)Lagarostrobos
PodocarpiditesP. elegansPodocarpaceae
P. elipticusPodocarpaceae
P. exiguusPodocarpaceae
P. herbstiiPodocarpaceae
P. majorPodocarpaceae
P. marwikiiPodocarpaceae
P. otagoensisPodocarpaceae
P. sp. cf. P. langiiPodocarpaceae
P. sp. cf. P. microreticuloidataPodocarpaceae
P. torquatusPodocarpaceae
P. verrucosusPodocarpaceae
PodocarpoxylonP. duseniiPodocarpaceae
TrichotomosulcitesT. subgranulatusPodocarpaceae

Magnoliids

GenusSpeciesAffinity / FamilyImage
ClavatimonocolpitesC. sp.Magnoliids

Monocots

GenusSpeciesAffinity / FamilyImage
ArecipitesA. minutiscabratusArecaceae
A. sp. cf. A. subverrucatusArecaceae
Aff.Eolirion"Morphotype 5"Palmae
LongapertitesL. proxapertitoidesArecaceae
LiliaciditesL. lanceolatusLiliales (aff. Lilium)Lilium
L. variegatusLiliales (aff. Lilium)
L. vermireticulatusLiliales
L. sp. cf. L. kaitangataensisLiliaceae/Amaryllidaceae
Monocotyledonean"Morphotype 6"Monocots
NupharN.? sp. 1NymphaeaceaeExtant specimen
N.? sp. 2Nymphaeaceae
PandaniiditesP. typicusAraceae
ProxapertitesP. operculatusAraceae
SpinozonocolpitesS. hialinusArecaceae

Eudicots

GenusSpeciesAffinity / FamilyImage
BeaupreaB. elegansiformisProtealesBeauprea drawing
Aff. Brachychiton"Morphotype 4"MalvaceaeExtant specimen
ButtiniaB. andreeviiBuxales
ClavamonocolpitesC. sp.?Chloranthaceae
Coccoloba?"Morphotype 1"PolygonaceaeExtant specimen
ConstantinisporisC. jacquei?Chloranthaceae
DicotetraditesD. sp.Ericales
DiporitesD. sp.?Chloranthaceae
DoyleipollenitesD. sp.?Chloranthaceae
EricipitesE. scabratusEricales
Eudicots"Morphotype 3"Eudicots
MyrtaceiditesM. sp. cf. M. mesonesusMyrtales
NothofagiditesN. saraensisNothofagaceae (cf. Nothofagus)Nothofagus
NyssapollenitesN. sp.?Chloranthaceae
PeninsulapollisP. gilliiProteaceae (cf. Beauprea?)
P. truswelliaeProteaceae (cf. Beauprea?)
PolycolpitesP. sp.Pedaliaceae (?cf. Josephinia)
ProteaciditesP. parvusProteaceae (cf. Bellendena montana)Bellendena
P. subscabratusProteaceae
P. sp. cf. P. tenuiexinusProteaceae
P. sp.Proteaceae
PsilatricolporitesP. sp. cf. P. complanatiusTetrameristaceae
P. sp.Tetrameristaceae
RetidiporitesR. camachoiProteaceae
RhoipitesR. baculatusEricales
R. minusculusEricales
R. sp. cf. R. alveolatusEricales
R. sp. 1Ericales
R. sp. 2Ericales
RouseaR. patagonicaEricales
R. sp. cf. R. minusculaEricales
R. sp.Ericales
SyncolporitesS. sp.Ericales
TetracolpitesT. sp.?Chloranthaceae
TriatriopollenitesT. lateflexusProteaceae
TricesticillusT. americanus?Chloranthaceae
TricolpitesT. reticulatusGunneraceae (Gunnera macrophylla)Gunnera
T. sp. cf. T. brevicolpusGunneraceae
T. sp. cf. T. inargutusGunneraceae
T. sp. cf. T. lumbrerensisGunneraceae
T. sp. 1Ranunculaceae (cf. Clematis)Clematis
TricolporitesT. sp. 2Ericales
T. sp. 3Ericales
TricolporoiditesT. sp. cf. T. eximiusEricales
TrioritesT. sp. cf. T. minusculus?Chloranthaceae
TubulifloriditesT. lillieiEricales
VolkheimeritesV. labyrinthus?Chloranthaceae

See also

Bibliography

  • Marenssi, S. A.; Casadío, S.; Santillana, S. N. (September 2003). [Stratigraphy and sedimentology of the Upper Cretaceous-Paleogene units outcropping on the southeastern margin of Lake Viedma, Santa Cruz Province, Argentina]. Revista de la Asociación Geológica Argentina (in Spanish). 58 (3): 403–416.