The Densuș-Ciula Formation is a geological formation in Romania, with strata dating to the Late Cretaceous. It forms part of the Hațeg Island assemblage, with dinosaur remains among the fossils recovered from the formation.

The formation is divided into three members. The lower member contains a high proportion of volcanogenic material and is poorly fossiliferous. The middle member consists of silty mudstones, sandstones, and conglomerates containing volcanogenic clasts, and is richly fossiliferous. The upper member is composed of matrix-supported red conglomerates and contains few fossils.

Fossil content

Indeterminate dromaeosaurid and possible indeterminate troodontid remains present in Județul Hunedoara, Romania. An unnamed theropod is also present.

Amphibians

Amphibians
GenusSpeciesPresenceNotesImages
AlbanerpetonIndeterminateAlbanerpetontid
ParalatoniaP. transylvanicaAlytid
HatzegobatrachusH. grigorescuiBombinatorid

Squamates

Squamates
GenusSpeciesPresenceNotesImages
NidophisN. insularisMadtsoiid snake

Turtles

Turtles
GenusSpeciesPresenceNotesImages
KallokibotionK. bajazidi
DortokidaeIndeterminate

Crocodyliformes

Crocodyliformes
GenusSpeciesPresenceMaterialNotesImages
AllodaposuchusA. precedensA skullA crocodyliform.Allodaposuchus precedens skullLife restoration of Doratodon
DoratodonIndeterminate
SabresuchusS. sympiestodonA neosuchian, formerly thought to be Theriosuchus

Dinosaurs

Dinosaurs of the Densuș-Ciula Formation
GenusSpeciesPresenceMaterialNotesImages
BalaurB. bondocA paravian, possibly a junior synonym of Elopteryx
FerenceratopsF. shqiperorumHunedoara CountyA partial skeleton of a juvenile individual and a left dentaryOriginally "Zalmoxes" shqiperorum but subsequently recognized as a distinct ceratopsian taxon closely related to Ajkaceratops. Only the pelvic material has been referred to Ferenceratops as of the taxon's description in 2026.
KryptohadrosK. kallaiaeHunedoara CountyA partial skull and skeletonA hadrosauroid
TelmatosaurusT. transsylvanicusA skull.A hadrosauromorph.
UriashU. kadiciLocality VI of Kadić.A partial skeleton composed of incomplete vertebrae, metatarsal, femur, and humerus.A titanosaur.
ZalmoxesZ. robustusHunedoara CountyA small rhabdodontid ornithopod

Pterosaurs

Pterosaurs of Densuș-Ciula Formation.
GenusSpeciesPresenceMaterialNotesImages
HatzegopteryxH. thambemaValioaraA preserved robust skull, incomplete left humerus, and unidentified bone fragments.A giant azhdarchid pterosaur.

Mammals

Mammals
GenusSpeciesPresenceNotesImages
HaininaIndeterminateTwo species

See also