Enantiornis is a genus of Enantiornithes. The type and only currently accepted species E. leali is from the Late Cretaceous Lecho Formation at El Brete, Argentina. It was described from specimen PVL-4035, a coracoid, proximal scapula and proximal humerus found close to each other and suspected to represent the left shoulder of a single individual.

Description

The genus and the larger group it belongs to, get their name from the reversed scapula-coracoid connection they possess compared to modern birds and the hesperornithids that were their contemporaries: Enanti "opposite", ornis is "bird".

Another left shoulder and wing, almost complete and found associated in one lump of rock, as well as a few isolated bones were also assigned to this species mainly based on size. It is among the largest enantiornithines discovered to date, with a length in life of around 78.5 cm (30.9 in), hip height of 34 cm (13 in), weight of 6.75 kg (14.9 lb), and wingspan comparable to herring gulls, around 1.2 m (3 ft 11 in). Its ecological niche resembled that of a mid-sized vulture or eagle.

Taxonomy and phylogeny

E. leali was possibly fairly closely related to Avisaurus, another genus of probably carnivorous enantiornithines, though its exact relationship is unclear. It was placed in a family of its own, Enantiornithidae. Other species from Asia that were previously placed in this genus are now split off. The former Enantiornis martini is now placed in Incolornis, while the former Enantiornis walkeri is now tentatively assigned to Explorornis. The reason for this is that these species were described when the diversity of enantiornithines was underestimated.

As no hindlimb elements are known from Enantiornis, it might include one of the El Brete enantiornithines known only from leg bones, namely Lectavis, Soroavisaurus or Yungavolucris. However, these apparently were all smaller than Enantiornis. Hindlimb material tentatively assigned to Martinavis and Elbretornis seems somewhat and a lot too small, respectively, to represent Enantiornis.

The cladogram below is from Wang et al., 2022:

EnantiornithesCruralispennia Protopteryx Elsornis Flexomornis Longipterygidae Iberomesornis bh bh Longirostravis Rapaxavis Euenantiornithes c e Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Cruralispennia
Protopteryx Elsornis Flexomornis Longipterygidae Iberomesornis bh bh Longirostravis Rapaxavis Euenantiornithes c e Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Protopteryx
Elsornis Flexomornis Longipterygidae Iberomesornis bh bh Longirostravis Rapaxavis Euenantiornithes c e Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Elsornis Flexomornis
Elsornis
Flexomornis
Longipterygidae Iberomesornis bh bh Longirostravis Rapaxavis Euenantiornithes c e Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
LongipterygidaeIberomesornis bh bh Longirostravis Rapaxavis
Iberomesornis
bhbh Longirostravis Rapaxavis
bhLongirostravis
Rapaxavis
Euenantiornithesc e Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
ce Sinornis e Enantiornis Halimornis Concornis Neuquenornis c b Eoalulavis Liaoningornis Gobipteryx Hebeiornis
eSinornis
eEnantiornis Halimornis Concornis Neuquenornis
Enantiornis
Halimornis Concornis Neuquenornis
Halimornis
Concornis Neuquenornis
Concornis
Neuquenornis
cb Eoalulavis Liaoningornis Gobipteryx Hebeiornis
bEoalulavis Liaoningornis
Eoalulavis
Liaoningornis
Gobipteryx Hebeiornis
Gobipteryx
Hebeiornis
Eocathayornis Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Eocathayornis
Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Qiliania Avisauridae Intiornis Soroavisaurus Avisaurus Gettyia n Mirarce e
Qiliania
AvisauridaeIntiornis Soroavisaurus Avisaurus Gettyia n Mirarce
Intiornis Soroavisaurus
Intiornis
Soroavisaurus
Avisaurus Gettyia n Mirarce
Avisaurus
Gettyia n Mirarce
Gettyia
nMirarce
e
Feitianius fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Feitianius
fn Longusunguis fn n Dunhuangia fn Musivavis Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
fn Longusunguis fn n Dunhuangia fn Musivavis
fnLongusunguis
fnn Dunhuangia fn Musivavis
nDunhuangia
fnMusivavis
Pengornithidae Pengornis m Yuanchuavis Eopengornis Parapengornis m Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
PengornithidaePengornis m Yuanchuavis Eopengornis Parapengornis
Pengornis
mYuanchuavis Eopengornis Parapengornis
Yuanchuavis
Eopengornis Parapengornis
Eopengornis
Parapengornis
m
Grabauornis Parvavis Cratoavis Gretcheniao Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Grabauornis Parvavis Cratoavis Gretcheniao
Grabauornis
Parvavis Cratoavis Gretcheniao
Parvavis
Cratoavis Gretcheniao
Cratoavis
Gretcheniao
Huoshanornis Shengjingornis Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Huoshanornis Shengjingornis
Huoshanornis
Shengjingornis
Zhouornis Bohaiornithidae Bohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Zhouornis
BohaiornithidaeBohaiornis ij Eoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Bohaiornis
ijEoenantiornis Fortunguavis Mystiornis Shenqiornis Sulcavis
Eoenantiornis
Fortunguavis Mystiornis Shenqiornis Sulcavis
Fortunguavis
Mystiornis Shenqiornis Sulcavis
Mystiornis
Shenqiornis Sulcavis
Shenqiornis
Sulcavis
l

Key to letters:

b = Boluochia c = Cathayornis e = Enantiophoenix f = Houornis h = Longipteryx i = Parabohaiornis j = Pterygornis l = Vorona m = Yuanjiawaornis n = Yungavolucris