Crambinae

Crambinae is a large subfamily of the lepidopteran family Crambidae, the crambid snout moths. It currently includes over 1,800 species worldwide. The larvae are root feeders or stem borers, mostly on grasses. A few species are pests of sod grasses, maize, sugar cane, rice, and other Poaceae. The monophyly of this group is supported by the structure of the tympanal organs and the phallus attached medially to the juxta, as well as genetic analyses.

Taxonomists' opinions differ as to the correct placement of the Crambidae, some authorities treating them as a subfamily of the family Pyralidae. If this is done, the present group would be demoted to tribe status, as Crambini.

Crambinae taxonomy

• tribe incertae sedis Anaclastis Turner, 1904 Aphrophantis Meyrick, 1933 Argentochiloides Błeszyński, 1961 (= Argentochilo Błeszyński & Collins, 1962) Australargyria Błeszyński, 1970 Autarotis Meyrick, 1886 (= Pogonoptera Turner, 1911) Batiana Walker, 1866 Burmannia Bleszynski, 1965 Catancyla Hampson, 1919 Catharylla Zeller, 1863 Charltona Swinhoe, 1886 Charltoniada Strand, 1918 Conocrambus Hampson, 1919 (= Anaresca Turner, 1947) Corynophora Berg, 1898 (= Halterophora Meyrick, 1897) Cypholomia Meyrick, 1933 Diadexia Turner, 1905 Diploptalis Hampson, 1919 (= Diplotalis Błeszyński, 1963) Diploschistis Meyrick, 1937 Elethyia Ragonot in de Joannis & Ragonot, 1889 Epina Walker, 1866 (= Diatraenopsis Dyar & Heinrich, 1927) Eurhythma Turner, 1904 (= Eurythma Błeszyński & Collins, 1962) Hemiplatytes Barnes & Benjamin, 1924 (= Alamogordia Dyar & Heinrich, 1927) Hemiptocha Dognin, 1905 Idioblasta Warren, 1891 Japonichilo Okano, 1962 Kupea Philpott, 1930 Libuna Moore, 1886 (= Bulina Walker, 1866) Lyndia Savigny, 1816 Maoricrambus Gaskin, 1975 Mestolobes Butler, 1882 (= Promylaea Meyrick, 1899) Metaeuchromius Błeszyński, 1960 (= Pseudeuchromius Błeszyński, 1965) Micrelephas Dognin, 1905 Microchilo Okano, 1962 Micronix Amsel, 1956 Microtalis Turner, 1911 Neargyria Hampson, 1896 Neargyrioides Błeszyński, 1970 Nechilo Błeszyński, 1970 Neobanepa Hampson, 1919 (= Pyralopsis Dognin, 1905) Niveocatharylla Bassi, 1999 Orthomecyna Butler, 1883 Parancyla Hampson, 1919 Paratraea Hampson, 1919 Parerupa Hampson, 1919 (= Coenotalis Hampson, 1919) Protyparcha Meyrick, 1909 Pseudargyria Okano, 1962 Pseudometachilo Błeszyński, 1962 Ptochostola Meyrick, 1882 Roxita Błeszyński, 1963 (= Modestia Błeszyński, 1965) Schoenobiodes Hampson, 1917 (= Stenopydna Roepke, 1943) Sphaerodeltis Meyrick, 1933 Styxon Błeszyński, 1962 Thalamarchis Meyrick, 1897 Tomissa Walker, 1864 Tulla Zimmerman, 1958 Ubida Walker, 1863 (= Crunophila Meyrick, 1882) Zacatecas Błeszyński, 1962
• tribe Ancylolomiini Ragonot, 1889 Ancylolomia Hübner, 1825 (= Acylolomia Hampson, 1919, Jartheza Walker, 1863, Pseudoctenella Strand, 1907, Ctenus Mabille, 1906, Tollia Amsel, 1949) Aurotalis Błeszyński, 1970 Eufernaldia Hulst, 1900 Mesolia Ragonot in de Joannis & Ragonot, 1889 (= Deuterolia Dyar, 1914, Eugrotea Fernald, 1896, Euparolia Dyar, 1914, Masolia Hampson, 1919) Prionapteron Błeszyński, 1965 Prionapteryx Stephens, 1834 (= Alloea Turner, 1947, Calarina Walker, 1866, Hypotomorpha Rebel, 1892, Nuarace Walker, 1863, Naurace Błeszyński & Collins, 1962, Pindicitora Walker, 1863, Pindicitra Shibuya, 1928, Platytesia Strand, 1918, Prionopteryx Zeller, 1863) Prionotalis Hampson, 1919 Pseudoschoenobius Fernald, 1891 Surattha Walker, 1863 Talis Guenée, 1845 (= Drasa Kapur, 1950, Prosmixis Zeller, 1846, Tulis Pagenstecher, 1909) Zovax Błeszyński, 1962
• tribe Argyriini Munroe, 1995 Argyria Hübner, 1818 Urola Walker, 1863 Vaxi Błeszyński, 1962
• tribe Calamotrophini Gaskin, 1988 Calamotropha Zeller, 1863 (= Aurelianus Błeszyński, 1962, Myeza Walker, 1863) Chrysocatharylla Bassi, 1999 Classeya Błeszyński, 1960 Pseudocatharylla Błeszyński, 1961 Pseudoclasseya Błeszyński, 1964
• tribe Chiloini Heinemann, 1865 Chilandrus Błeszyński, 1970 Chilo Zincken, 1817 (= Borer Guenée, 1862, Chilona Sodoffsky, 1837, Chilotraea Kapur, 1950, Diphryx Grote, 1881, Hypiesta Hampson, 1919, Nephalia Turner, 1911, Silveria Dyar, 1925) Chiqua Błeszyński, 1970 Diatraea Guilding, 1828 (= Crambidiatraea Box & Capps, 1955, Diaraetria Grote, 1882, Diatraerupa Schaus, 1913, Diatrea Guilding, 1828, Eodiatraea Box, 1953, Iesta Dyar, 1909, Trinidadia Dyar & Heinrich, 1927, Zeadiatraea Box, 1955) Eschata Walker, 1856 (= Chaerecla Walker, 1865) Gadira Walker, 1866 (= Cryptomima Meyrick, 1882, Scenoploca Meyrick, 1882) Hednota Meyrick, 1886 Leonardo Błeszyński, 1965 Malgasochilo Błeszyński, 1970 Myelobia Herrich-Schäffer, 1854 (= Chilopsis Hampson, 1919, Doratoperas Hampson, 1896, Protaphomia Meyrick, 1936, Xanthopherne Dyar & Heinrich, 1927) Tauroscopa Meyrick, 1888 (= Oressaula Turner, 1913)
• tribe Crambini Latreille, 1810 Agriphila Hübner, 1825 (= Agrophila J. L. R. Agassiz, 1847, Alisa Ganev & Hacker, 1984) Agriphiloides Błeszyński, 1965 Almita B. Landry, 1995 Alphacrambus Bassi, 1995 Amselia Błeszyński, 1959 Angustalius Marion, 1954 (= Bleszynskia Lattin, 1961, Crambopsis Lattin, 1952) Arequipa Walker, 1863 Aureocramboides Błeszyński, 1961 Bassiknysna Kemal & Kocak, 2005 (= Knysna Bassi, 1999) Caffrocrambus Błeszyński, 1961 (= Anomocrambus Błeszyński, 1961, Caffocrambus Bassi, 1994) Catoptria Hübner, 1825 (= Exoria Hübner, 1825, Tetrachila Hübner, 1806) Cervicrambus Błeszyński, 1966 Chrysocrambus Błeszyński, 1957 (= Chrysocramboides Błeszyński, 1957) Chrysoteuchia Hübner, 1825 (= Amphibolia Snellen, 1884, Veronese Błeszyński, 1962) Conocramboides Błeszyński, 1970 Crambixon Błeszyński, 1965 Crambus Fabricius, 1798 (= Argyroteuchia Hübner, 1825, Chilus Billberg, 1820, Palparia Haworth, 1811, Tetrachila Hübner, 1822) Culladia Moore, 1886 (= Araxes Walker, 1863, Crambidion Mabille, 1900, Nirmaladia Rose, 1983) Culladiella Błeszyński, 1970 Dimorphocrambus Gibeaux, 1987 Epichilo Ragonot in de Joannis & Ragonot, 1889 Fernandocrambus Aurivillius, 1922 (= Juania Aurivillius, 1922, Juanita Munroe, 1995) Fissicrambus Błeszyński, 1963 Flavocrambus Błeszyński, 1959 Haplopediasia Błeszyński, 1963 Haploplatytes Błeszyński, 1966 Japonicrambus Okano, 1962 La Błeszyński, 1966 Loxocrambus Forbes, 1920 Mesocrambus Błeszyński, 1957 Mesopediasia Błeszyński, 1963 Metacrambus Błeszyński, 1957 Microcramboides Błeszyński, 1967 Microcrambon Błeszyński, 1970 Microcrambus Błeszyński, 1963 Miraxis Błeszyński, 1962 Miyakea Marumo, 1933 Neocrambus Błeszyński, 1957 Neoculladia Błeszyński, 1967 Neodactria B. Landry, 1995 Neopediasia Okano, 1962 Novocrambus Amsel, 1956 Orocrambus Purdie, 1884 (= Orocrambus Meyrick, 1885) Parapediasia Błeszyński, 1966 (Parapediasia Błeszyński, 1963) Paraplatytes Błeszyński, 1965 Pediasia Hübner, 1825 (= Carvanca Walker, 1856, Oseriates Fazekas, 1991, Pseudopediasia Ganev, 1987) Platytes Guenée, 1845 (= Nagahama Marumo, 1933) Precaffrocrambus Bassi, 2002 Productalius Marion, 1954 Pseudopediasia Błeszyński, 1963 Raphiptera Hampson, 1896 Sebrus Błeszyński, 1970 Sericocrambus Wallengren, 1861 Supercrambus Błeszyński, 1967 Tawhitia Philpott, 1931 (= Velasquez Błeszyński, 1962) Tehama Hulst, 1888 Thaumatopsis Morrison, 1874 (= Propexus Grote, 1880) Thisanotia Hübner, 1825 (= Thinasotia Heinemann, 1865, Thysanotia J. L. R. Agassiz, 1847) Tortriculladia Błeszyński, 1967 Xanthocrambus Błeszyński, 1957 (= Xanthocrambus Błeszyński, 1955)
• tribe Diptychophorini Gaskin, 1972 Cleoeromene Gaskin, 1986 Diptychophora Zeller, 1866 (= Colimaea Dyar, 1925, Colimea Błeszyński, 1966, Mysticomima Meyrick, 1931, Scissolia Barnes & McDunnough, 1914) Gargela Walker, 1864 (= Mixophyla Meyrick, 1887, Angonia Snellen, 1893, Mixophila Hampson, 1896) Glaucocharis Meyrick, 1938 (= Pagmania Amsel, 1961, Pareromene Osthelder, 1941, Ditomoptera Hampson, 1893) Incaeromene Gaskin, 1986 Neoeromene Gaskin, 1986 Peniculimius Schouten, 1994 Steneromene Gaskin, 1986 Tamsica Zimmerman, 1958
• tribe Euchromiusini Léger, Landry & Nuss, 2019 Euchromius Guenée, 1845 (= Eromene Hübner, 1825, Ommatopteryx Kirby, 1897, Pseudoancylolomia Ahmad, Zaidi & Kamaluddin, 1982)
• tribe Haimbachiini B. Landry, 1995 Achilo Amsel, 1957 (= Chilopsis Amsel, 1956) Bissetia Kapur, 1950 (= Girdharia Kapur, 1950) Coniesta Hampson, 1919 Donacoscaptes Zeller, 1877 Eoreuma Ely, 1910 Friedlanderia Agnew, 1987 (= Chiloides Amsel, 1949, Chiloides Błeszyński, 1963) Haimbachia Dyar, 1909 Neogirdharia Song & Chen in Chen, Song & Yuan, 2004 Occidentalia Dyar & Heinrich, 1927 Pseudobissetia Błeszyński, 1959 Thopeutis Hübner, 1818 (= Cephis Ragonot in Staudinger, 1892, Hombergia de Joannis, 1910, Stenochilo Hampson, 1896, Tetrachila Hübner, 1808, Topeutis Hübner, 1825) Xubida Schaus, 1922

Former genera

• Araxates Ragonot in de Joannis & Ragonot, 1889
• Charitopepla Meyrick, 1933
• Neerupa Hampson, 1919
• Loxophantis Meyrick, 1935
• Welaka Hulst, 1888

See also

• List of crambid genera