Iani (after Ianus) is an extinct genus of rhabdodontomorph iguanodontian dinosaur from the Late Cretaceous Cedar Mountain Formation of Utah, United States. The genus contains a single species, I. smithi, known from a partial skeleton including the skull. Its discovery serves as a link between the genus Tenontosaurus and the Rhabdodontidae, with both along with Iani being members of the clade Rhabdodontomorpha.

Discovery and naming

The Iani holotype specimen, NCSM 29373, was discovered in 2014 in sediments of the Cedar Mountain Formation (lower Mussentuchit Member) in Emery County, Utah, United States. The specimen consists of a partial skeleton including a nearly complete, disarticulated skull, cervical, dorsal, sacral, and caudal vertebrae, ribs and haemal arches, partial pectoral and pelvic girdles, and the right arm and leg.

In 2023, Zanno et al. described Iani smithi as a new genus and species of rhabdodontomorph iguanodontian based on these fossil remains. The generic name, "Iani", alludes to Ianus, the Roman god of transitions, referencing the changing biota of the time. The specific name, "smithi", honors Joshua Aaron Smith and his paleontological contributions.

Classification

Life reconstruction of Iani

Zanno et al. (2023) recovered Iani as a basal rhabdodontomorph member of the Iguanodontia. They interpreted it as a transitional taxon between Tenontosaurus and other Late Cretaceous rhabdodontomorphs (the Rhabdodontidae). The cladograms below display the results of their phylogenetic analyses using different datasets:

Topology 1: Poole (2022) dataset Rhabdodontomorpha Muttaburrasaurus Iani Tenontosaurus dossi Tenontosaurus tilletti Rhabdodon Zalmoxes shqiperorum Zalmoxes robustusTopology 2: Dieudonné et al. (2021) dataset Rhabdodontomorpha Tenontosaurus dossi Tenontosaurus tilletti Iani Rhabdodon sp1 Rhabdodon priscus Zalmoxes shqiperorum Zalmoxes robustus Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
RhabdodontomorphaMuttaburrasaurus Iani Tenontosaurus dossi Tenontosaurus tilletti Rhabdodon Zalmoxes shqiperorum Zalmoxes robustus
Muttaburrasaurus
Iani Tenontosaurus dossi Tenontosaurus tilletti Rhabdodon Zalmoxes shqiperorum Zalmoxes robustus
Iani Tenontosaurus dossi Tenontosaurus tilletti
Iani
Tenontosaurus dossi Tenontosaurus tilletti
Tenontosaurus dossi
Tenontosaurus tilletti
Rhabdodon Zalmoxes shqiperorum Zalmoxes robustus
Rhabdodon
Zalmoxes shqiperorum Zalmoxes robustus
Zalmoxes shqiperorum
Zalmoxes robustus
RhabdodontomorphaTenontosaurus dossi Tenontosaurus tilletti Iani Rhabdodon sp1 Rhabdodon priscus Zalmoxes shqiperorum Zalmoxes robustus Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Tenontosaurus dossi Tenontosaurus tilletti
Tenontosaurus dossi
Tenontosaurus tilletti
Iani Rhabdodon sp1 Rhabdodon priscus Zalmoxes shqiperorum Zalmoxes robustus Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Iani
Rhabdodon sp1 Rhabdodon priscus Zalmoxes shqiperorum Zalmoxes robustus Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Rhabdodon sp1 Rhabdodon priscus
Rhabdodon sp1
Rhabdodon priscus
Zalmoxes shqiperorum Zalmoxes robustus Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Zalmoxes shqiperorum Zalmoxes robustus
Zalmoxes shqiperorum
Zalmoxes robustus
Mochlodon vorosi Mochlodon suessi Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Mochlodon vorosi Mochlodon suessi
Mochlodon vorosi
Mochlodon suessi
Transylvanosaurus Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Transylvanosaurus
Foskeia ('Vegagete ornithopod') Muttaburrasaurus Fostoria
Foskeia ('Vegagete ornithopod')
Muttaburrasaurus Fostoria
Muttaburrasaurus
Fostoria
Fostoria