The Irdin Manha Formation is a geological formation from the Eocene located in Inner Mongolia, China, a few kilometres south of the Mongolian border.

Fossil content

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Mammals

U.S. paleontologists Henry Fairfield Osborn and Roy C. Andrews discovered two premolars on the site in 1923, and assigned the specimen to the new genus Eudinoceras because he believed it to be related to "Dinoceras" (now known as Uintatherium). Within a decade, however, as more complete specimens were recovered, the animal was identified as a Mongolian relative to the North American pantodont Coryphodon. The expedition also lead to the discovery of the only known skull of Andrewsarchus.

Artiodactyls

Artiodactyls reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
Achaenodontidae?An incomplete upper tooth (AMNH 20136).Doubtfully assigned to the family.
Cf. Archaeomeryx, gen indet.Fragment of lower jaw (AMNH 20173).A traguliform.
AndrewsarchusA. mongoliensisAround Telegraph Line Camp.A large skull.A relative of entelodonts formerly thought to be a mesonychid.
ErlianhyusE. primitivusIrdin Manha, Erlian Basin, Nei Mongol.A right upper maxilla with P3–M3 (IVPP V 28275).A basal artiodactyl.
GobiohyusG. orientalisTelegraph Line Camp.Jaw elements.A helohyid also found in the Ulan Shireh Formation.
G. pressidensTelegraph Line Camp.Partial right rami.A helohyid.
G. robustusTelegraph Line Camp.Left ramus (AMNH 20246).A helohyid.
ObotheriumO. parvumIrdin Manha, Erlian Basin, Nei Mongol.Teeth and lower jawsA tapirulid.

Cimolestans

Cimolestans reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
EudinocerasE. mongoliensis2 premolars & jaw elements.A coryphodontid.

Dinoceratans

Dinoceratans reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
GobiatheriumG. mirificum25 miles southwest from Iren Dabasu.Skull, jaws & limb elements.An uintatheriid.

Ferae

Ferae reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
MiacisM. invictusIsolated upper molar (AMNH 20137).A miacid.
PropterodonP. irdinensisJaw fragments.A hyaenodontid.
SarkastodonS. mongoliensisAbout 25 miles southwest of Iren Dabasu.Skull & jaws.An oxyaenid.

Glires

Glires reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
AdvenimusA. burkeiNear Camp Margetts.Jaw elements.A ctenodactyloid rodent.
AsiomysA. dawsoniHuheboerhe.Fragment of right calcaneus (IVPP V24417).An ischyromyid rodent.
ErenlagusE. anielaeHuheboerhe.Teeth.A stem-lagomorph.
GomphosG. shevyrevaeHuheboerhe escarpment.Teeth & foot elements.A mimotonid.
Ischyromyidae genus indet.Species AIrdin Manha escarpment.Right calcaneus (IVPP V24416).Relatively large ischyromyid rodent, calcaneus comparable in size to that of a coypu or Asiatic brush-tailed porcupine.
Species BDaoteyin Obo.Right calcaneus (IVPP V24418).Large ischyromyid rodent with calcaneus matching in length that of a coypu & similar in structure to that of Paramys.
MimolagusM. auroraeIrdin Manha escarpment.Teeth & foot elements.A large mimotonid.
PappocricetodonP. neimongolensisHuheboerhe.Teeth.A cricetid rodent.
P. cf. P. zhongtiaensisHuheboerhe.A left molar.A cricetid rodent.
P. sp.Huheboerhe.A right molar.A cricetid rodent.
ParamyidaeParamyid spp.Teeth & jaws.3 sizes of paramyid rodents, possibly different species.
?Paramyid sp.Left jaw (AMNH 80801).Possible small paramyid rodent.
SimplicimysS. bellusHuheboerhe.Teeth.A ctenodactyloid rodent also known from the Arshanto Formation.
StrenulagusS. solarisIrdin Manha and Huheboerhe localities.Isolated cheek teeth, fragmentary upper incisors (dI2) and postcranial elements.A stem-lagomorph also known from the Khaychin Formation.
TamquammysT. fractusHuheboerhe.Right maxilla (V17798).A ctenodactyloid rodent.
T. wilsoniTeeth.A ctenodactyloid rodent also known from the Arshanto & Nomogen formations.
YongshengomysY. extensusHuheboerhe.Teeth.A ctenodactyloid rodent.
YuomysY. huheboerhensisHuheboerhe.Teeth.A ctenodactyloid rodent.
Y. sp. AHuheboerhe.A right molar (V17805).A ctenodactyloid rodent.
Y. sp. BHuheboerhe.A right molar (V17806).A ctenodactyloid rodent.
Y. sp. CHuheboerhe.A right molar (V17807).A ctenodactyloid rodent.

Mesonychians

Mesonychians reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
Hapalodectes?H. auctusAn upper molar or possibly premolar (AMNH 20130).A hapalodectid.
H. serusAround Telegraph Line Camp.Lower cheek tooth (AMNH 20172).A hapalodectid.
HarpagolestesH. leeiHuheboerhe & Daoteyin Obo.Tooth elements.A mesonychid.
MesonychidaeGen. indet.AMNH 20132.About the size of Harpagolestes uintensis.
Gen. indet.Isolated tooth (AMNH 20133).About the size of Synoplotherium lanius.
MongolonyxM. dolichognathus7 miles west of Camp Margetts.Jaw elements.A large mesonychid.

Panperissodactyla

Panperissodactylas reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
CaenolophusC. proficiensLower jaw & isolated teeth.A rhinocerotoid.
CooperiaC. totadentataTelegraph Line Camp, 23 miles southeast of Iren Dabasu.Front of skull (AMNH 20116).Generic name preoccupied, renamed Forstercooperia.
DesmatotheriumD. fissumUpper jaw fragment.A helaletid.
D. mongolienseIrdin Manha escarpment, Duheminboerhe, Huheboerhe & Chaganboerhe.Parts of 10 individuals.A helaletid also found in the Mergen Formation.
ForstercooperiaF. totadentataTelegraph Line Camp, 23 miles southeast of Iren Dabasu.Front of skull (AMNH 20116).A paraceratheriid, originally named Cooperia.
F. ulanshirehensisIrdin Manha site.Jaw elements.A paraceratheriid also known from the Ulan Shireh Formation.
GnathotitanG. berkeyiIrdin Manha.Jaw elements.A brontothere.
LophialetesL. expeditusJaw elements.A lophialetid.
L. minutusUpper molar.A lophialetid.
MetatelmatheriumM. cristatumCamp Margetts.Skull & lower jaw (AMNH 26411).A brontothere.
M. parvumIrdin Manha.Fragment of left lower jaw (AMNH 20168).A brontothere.
MicrotitanM. mongoliensisJaw elements.A brontothere.
PappacerasP. confluensUpper gray clays & Camp Margetts area.Skull & jaw elements.A paraceratheriid.
ParacolodonP. fissusDuheminboerhe & Daoteyin Obo.Skull elements & teeth.A helaletid.
ProtitanP. bellusSpring Camp, East Mesa, Shara Murun region.Palatal side of skull (AMNH 26104).A brontothere.
P. grangeriSkull elements.A brontothere.
P. minorCamp Margetts.Skull elements.A brontothere.
P. obliquidensIrdin Manha.Part of left maxilla (AMNH 20125).A brontothere.
P. robustusIrdin Manha.Jaw elements & teeth.A brontothere.
RostriamynodonR. grangeriEast of Camp Margetts.Complete skull & mandible (AMNH 107635).An amynodontid.
TeleolophusT. mediusTeeth & jaws.

Primates

Primates reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
TarkopsT. mckennaiHuheboerhe.Incomplete left lower jaw (IVPP V16424).An omomyid.

Reptiles

Birds

Birds reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
ButeoninaeChimney Butte.Left coracoid (AMNH FR 2941).Specimen reassigned to Eogrus? sp.
EogrusE. aeolaChimney Butte.Right metatarsus (AMNH 2936).An eogruid.
E.? sp.Chimney Butte.Left coracoid (AMNH FR 2941).Formerly thought to represent a buteonine, now thought to be an eogruid.
TelecrexT. grangeriChimney Butte, Shara Murun region.An incomplete right femur (AMNH 2942).A guineafowl, formerly thought to be a rail.

Crocodilians

Crocodilians reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
AsiatosuchusA. grangeri25 miles southwest of Iren Dabasu.Portions of at least 2 individuals.A crocodyloid.

Testudines

Testudines reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
"Adocus""A". orientalisTelegraph Line Camp.Anterior half of plastron (AMNH 6356).An adocid.
Amyda?A.? johnsoniTelegraph Line Camp.Two-thirds of a carapace (AMNH 6357) & many carapace fragments (AMNH 6359).A softshell turtle.
TrionychidaeIndeterminateTelegraph Line Camp.Several kinds of sculptured fragments (AMNH 6360).Remains of softshell turtles.

Fish

Fish reported from the Irdin Manha Formation
GenusSpeciesPresenceMaterialNotesImages
PappichthysP. mongoliensisTelegraph Line Camp.Vertebrae.Specimens now believed to represent Siluriformes.
SiluriformesIndeterminateTelegraph Line Camp, Chahar Province.5 vertebrae (AMNH 8535).A catfish likely related to ictalurids. Formerly assigned to Pappichthys mongoliensis.

See also