Lemmysuchus is a genus of machimosaurid thalattosuchian from the Middle Jurassic Callovian of England and France. Like many other teleosauroids from Europe, it has had a convoluted taxonomic history.

History

Skull and other elements of the holotype specimen

"Steneosaurus" obtusidens was first described from the Callovian-aged Oxford Clay Formation of Cambridgeshire, southeastern England on the basis of a partial skeleton (NHMUK PV R 3168). Characters cited as diagnostic for "S." obtusidens included: short rostrum (preorbital length 52 percent of total skull length); teeth blunt and rounded at the tips; armor adorned with elongate pit ornamentation.

The validity of "Steneosaurus" obtusidens was disputed in a 1987 paper reviewing the type specimens of nominal teleosauroid species from the Oxford Clay. Characters used by Charles William Andrews to distinguish "S." obtusidens from other species were dismissed as variable within specimens of Steneosaurus edwardsi, and "S." obtusidens was sunk as a junior synonym of Steneosaurus durobrivensis (=S. edwardsi). Some authors considered this taxon a probable junior synonym of Machimosaurus hughii based on a subsequently discovered specimen found in Callovian-age exposures in Calvados, Lower Normandy, France, though they stressed that a taxonomic review of blunt-snouted teleosaurids was needed. This synonymy was accepted in a 2009 paper regarding thalattosuchian morphometrics without comment.

In 2013, a new specimen of Machimosaurus was described from Late Jurassic (Kimmeridgian deposits in southern Germany (later named Machimosaurus buffetauti), and it became clear that variation within blunt-toothed teleosauroids was taxonomically significant enough for "Steneosaurus" obtusidens to be recognized as generically distinct from Machimosaurus. Other cladistic and comparative studies agreed with this assessment, recovering "S." obtusidens in a clade with Machimosaurus and Steneosaurus edwardsi.

In 2017, the species was moved to its own genus, Lemmysuchus, referring to Ian Fraser Kilmister, better known as "Lemmy", of the band Motörhead, and the Greek word for crocodile, soukhos.

Classification

Lemmysuchus is a member of the derived group of machimosaurids called Machimosaurini, which includes the largest and most robust members. It is closely related to other large machimosaurids such as Yvridiosuchus and Machimosaurus.

The cladogram below is from an analysis conducted by Johnson and colleagues in 2020.

MachimosauridaeMacrospondylus bollensis Clovesuurdameredeor stephani Charitomenosuchus leedsi Seldsienean megistorhynchus Deslongchampsina larteti Machimosaurinae Proexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Macrospondylus bollensis
Clovesuurdameredeor stephani Charitomenosuchus leedsi Seldsienean megistorhynchus Deslongchampsina larteti Machimosaurinae Proexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Clovesuurdameredeor stephani
Charitomenosuchus leedsi Seldsienean megistorhynchus Deslongchampsina larteti Machimosaurinae Proexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Charitomenosuchus leedsi
Seldsienean megistorhynchus Deslongchampsina larteti Machimosaurinae Proexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Seldsienean megistorhynchus
Deslongchampsina larteti Machimosaurinae Proexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Deslongchampsina larteti
MachimosaurinaeProexochokefalos hebertii cf. Proexochokefalos bouchardi Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Proexochokefalos hebertii cf. Proexochokefalos bouchardi
Proexochokefalos hebertii
cf. Proexochokefalos bouchardi
Neosteneosaurus edwardsi Andrianavoay baroni Machimosaurini Lemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Neosteneosaurus edwardsi
Andrianavoay baroni
MachimosauriniLemmysuchus obtusidens Yvridiosuchus boutilieri Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Lemmysuchus obtusidens
Yvridiosuchus boutilieri
Machimosaurus buffetauti Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Machimosaurus buffetauti
Machimosaurus mosae Machimosaurus rex Machimosaurus hugii
Machimosaurus mosae
Machimosaurus rex Machimosaurus hugii
Machimosaurus rex
Machimosaurus hugii

Ecology

Lemmysuchus lived during a time when much of Europe was covered by shallow seas and was an island archipelago. It had very large muscle attachment sites and robust jaws, meaning it had a powerful bite. Coupled with its straight and blunt dentition, Lemmysuchus was well adapted for feeding on large and armored prey items such as turtles and large fish.

Lemmysuchus is estimated to have a body length of roughly 5.8 meters (19 feet) based on known remains.