This article lists the genera of the Archaea and also shows taxa of archaea that do not contain genera. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI) unless otherwise noted.
Phylogeny
For the GTDB phylogeny, National Center for Biotechnology Information (NCBI) taxonomy was initially used to decorate the genome tree via tax2tree. The 16S rRNA-based Greengenes taxonomy is used to supplement the taxonomy particularly in regions of the tree with no cultured representatives. List of Prokaryotic names with Standing in Nomenclature (LPSN) is used as the primary taxonomic authority for establishing naming priorities. Taxonomic ranks are normalised using Phylorank and the taxonomy manually curated to remove polyphyletic groups.
Cladogram was taken from the GTDB release 11-RS232 (15th April 2026).
Phylum Microcaldota
Class Microcaldia
Order Microcaldales
- Family Microcaldaceae Sakai et al. 2023 Microcaldus Sakai et al. 2023
Phylum "Undinarchaeota"
| Phylogeny of "Undinarchaeota" |
| "Undinarchaeia" "Undinarchaeales" "Naiadarchaeaceae" "Ca. Naiadarchaeum" "Undinarchaeaceae" "Ca. Undinarchaeum" |
| "Undinarchaeia" | "Undinarchaeales" "Naiadarchaeaceae" "Ca. Naiadarchaeum" "Undinarchaeaceae" "Ca. Undinarchaeum" |
| "Undinarchaeales" | "Naiadarchaeaceae" "Ca. Naiadarchaeum" "Undinarchaeaceae" "Ca. Undinarchaeum" |
| "Naiadarchaeaceae" | "Ca. Naiadarchaeum" |
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| "Undinarchaeaceae" | "Ca. Undinarchaeum" |
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Class "Undinarchaeia"
Order "Undinarchaeales"
- Family "Naiadarchaeaceae" Dombrowski et al. 2020 "Candidatus Naiadarchaeum" Dombrowski et al. 2020
- Family "Undinarchaeaceae" Dombrowski et al. 2020 "Candidatus Undinarchaeum" Dombrowski et al. 2020
Phylum "Huberarchaeaota"
Class "Huberarchaeia"
Order "Huberarchaeales"
- Family "Huberarchaeaceae" Rinke et al. 2020 "Candidatus Huberarchaeum" corrig. Schwank et al. 2019
Phylum " Aenigmatarchaeota "
| Phylogeny of "Aenigmatarchaeota" |
| "Aenigmatarchaeia" "Aenigmatarchaeaceae" "Ca. Aenigmatarchaeum" PWEA01 "Ca. Haloaenigmatarchaeum" |
| "Aenigmatarchaeia" | "Aenigmatarchaeaceae" "Ca. Aenigmatarchaeum" PWEA01 "Ca. Haloaenigmatarchaeum" |
| "Aenigmatarchaeaceae" | "Ca. Aenigmatarchaeum" |
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| PWEA01 | "Ca. Haloaenigmatarchaeum" |
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Class "Aenigmatarchaeia"
Order "Aenigmatarchaeales"
- Family "Aenigmatarchaeaceae" corrig. Rinke et al. 2020 "Candidatus Aenigmatarchaeum" corrig. Rinke et al. 2013
Order PWEA01
- Family "Haloaenigmatarchaeaceae" Gutiérrez-Preciado et al. 2024 [PWEA01] "Candidatus Haloaenigmatarchaeum" Gutiérrez-Preciado et al. 2024
Phylum "Terrarchaeota"
| Phylogeny of "Terrarchaeota" |
| "Terrarchaeia" "Terrarchaeales" "Marinivulcanaceae" "Ca. Marinivulcanus" "Terrarchaeaceae" "Ca. Terrarchaeum" |
| "Terrarchaeia" | "Terrarchaeales" "Marinivulcanaceae" "Ca. Marinivulcanus" "Terrarchaeaceae" "Ca. Terrarchaeum" |
| "Terrarchaeales" | "Marinivulcanaceae" "Ca. Marinivulcanus" "Terrarchaeaceae" "Ca. Terrarchaeum" |
| "Marinivulcanaceae" | "Ca. Marinivulcanus" |
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| "Terrarchaeaceae" | "Ca. Terrarchaeum" |
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Class "Terrarchaeia"
Order "Terrarchaeales"
- Family "Marinivulcanaceae" Hamm et al. 2026 [WAPT01] "Candidatus Marinivulcanus" Hamm et al. 2026 [WAPT01]
- Family "Terrarchaeaceae" Hamm et al. 2026 ["Caliditerrarchaeaceae" Hamm et al. 2025; GLR71] "Candidatus Terrarchaeum" Hamm et al. 2026 ["Candidatus Caliditerrarchaeum" Hamm et al. 2025; GLR71]
Phylum " Nanohalarchaeota "
| Phylogeny of "Nanohalarchaeota" |
| "Nanohalobiia" "Nanohalobiales" "Nanohalalkaliarchaeaceae" "Ca. Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Ca. Asbonarchaeum" "Ca. Nanoanaerosalina" "Nanohalobiaceae" "Ca. Nanosalinicola" "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Nanohalobiia" | "Nanohalobiales" "Nanohalalkaliarchaeaceae" "Ca. Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Ca. Asbonarchaeum" "Ca. Nanoanaerosalina" "Nanohalobiaceae" "Ca. Nanosalinicola" "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Nanohalobiales" | "Nanohalalkaliarchaeaceae" "Ca. Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Ca. Asbonarchaeum" "Ca. Nanoanaerosalina" "Nanohalobiaceae" "Ca. Nanosalinicola" "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Nanohalalkaliarchaeaceae" "Ca. Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Ca. Asbonarchaeum" "Ca. Nanoanaerosalina" |
| "Nanohalalkaliarchaeaceae" | "Ca. Nanohalalkaliarchaeum" |
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| "Nanoanaerosalinaceae" | "Ca. Asbonarchaeum" "Ca. Nanoanaerosalina" |
| "Ca. Asbonarchaeum" |
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| "Ca. Nanoanaerosalina" |
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| "Nanohalobiaceae" "Ca. Nanosalinicola" "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Nanohalobiaceae" | "Ca. Nanosalinicola" "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Ca. Nanosalinicola" |
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| "Ca. Nanohalococcus" "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Ca. Nanohalococcus" |
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| "Ca. Nanosalina" "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Ca. Nanosalina" |
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| "Ca. Nanohalobium" "Ca. Nanohalovita" |
| "Ca. Nanohalobium" |
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| "Ca. Nanohalovita" |
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Class "Nanohalarchaeia"
Order "Nanohalarchaeales"
- Family "Nanohalarchaeaceae" ?"Candidatus Nanohalarchaeum" corrig. Hamm et al. 2019 ?"Candidatus Nanopetraeus" corrig. Crits‐Christoph et al. 2016
Order "Nanohydrothermales"
- Family "Nanohydrothermaceae" Xie et al. 2022 ?"Candidatus Nanohydrothermus" Xie et al. 2022
Order "Nucleotidisoterales"
- Family "Nanosalenecaceae" Xie et al. 2022 ?"Candidatus Nanosalenecus" Xie et al. 2022
- Family "Nucleotidisoteraceae" Xie et al. 2022 ?"Candidatus Nucleotidisoter" Xie et al. 2022
- Family "Nucleotidivindicaceae" Xie et al. 2022 ?"Nucleotidivindex" Xie et al. 2022
Class "Nanohalobiia"
Order "Nanohalobiales"
- Family "Nanohalalkaliarchaeaceae" La Cono et al. 2020 "Candidatus Nanohalalkaliarchaeum" Ghai et al. 2011
- Family "Nanoanaerosalinaceae" Zhao et al. 2022 "Candidatus Asbonarchaeum" Baker et al. 2024 "Candidatus Nanoanaerosalina" Zhao et al. 2022
- Family "Nanohalobiaceae" La Cono et al. 2020 ?"Candidatus Haloredivivus" Ghai et al. 2011 "Candidatus Nanohalobium" La Cono et al. 2020 "Candidatus Nanohalococcus" Reva et al. 2023 "Candidatus Nanosalina" Narasingarao et al. 2012 "Candidatus Nanosalinicola" corrig. Narasingarao et al. 2012 "Candidatus Nanohalovita" Reva et al. 2023
Phylum Nanobdellota
| Phylogeny of Nanobdellota |
| Nanobdellia "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" "Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella |
| Nanobdellia | "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" "Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella |
| "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" |
| "Tiddalikarchaeales" | "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" |
| "Tiddalikarchaeaceae" | "Ca. Tiddalikarchaeum" |
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| "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" |
| "JAPDLS01" | "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" |
| "Haiyanarchaeaceae" | "Ca. Haiyanarchaeum" |
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| "Jingweiarchaeales" | "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" |
| "Jingweiarchaeaceae" | "Ca. Jingweiarchaeum" |
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| "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "Parvarchaeales" | "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "Parvarchaeaceae" | "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "Ca. Rehaiarchaeum" |
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| "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum" |
| "Ca. Acidifodinimicrobium" |
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| "Ca. Parvarchaeum" |
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| "Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella |
| "Pacearchaeales" "Woesearchaeales" |
| "Pacearchaeales" |
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| "Woesearchaeales" |
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| Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella |
| Nanobdellales | "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella |
| "Nanoarchaeaceae" | "Nanoarchaeum" |
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| Nanobdellaceae | "Ca. Nanoclepta" Nanobdella |
| "Ca. Nanoclepta" |
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| Nanobdella |
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Class Nanobdellia
Order "Pacearchaeales"
Order "Woesearchaeales"
Order Nanobdellales
- Family "Nanoarchaeaceae" Huber et al. 2011 "Nanoarchaeum" Huber et al. 2002
- Family Nanobdellaceae Kato et al. 2022 ["Nanopusillaceae" Huber et al. 2011] Nanobdella Kato et al. 2022 ["Ca. Nanobsidianus" Castelle et al. 2015; "Ca. Nanopusillus" Wurch et al. 2016] "Candidatus Nanoclepta" St. John et al. 2019
Order "Tiddalikarchaeales"
- Family "Tiddalikarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Tiddalikarchaeum" Vázquez-Campos et al. 2021
Order JAPDLS01
- Family "Haiyanarchaeaceae" Rao et al. 2023 "Candidatus Haiyanarchaeum" Rao et al. 2023 ["Ca. Tinosarcha" Pallen, Rodriguez-R & Alikhan 2022]
Order "Jingweiarchaeales"
- Family "Jingweiarchaeaceae" Rao et al. 2023 "Candidatus Jingweiarchaeum" Rao et al. 2023
Order "Parvarchaeales"
- Family "Parvarchaeaceae" Rinke et al. 2020 "Candidatus Acidifodinimicrobium" Luo et al. 2020 "Candidatus Parvarchaeum" Baker et al. 2010 (ARMAN 4 & 5) "Candidatus Rehaiarchaeum" Rao et al. 2023
Phylum " Altarchaeota "
Class "Altarchaeia"
Order "Altarchaeales"
- Family "Altarchaeaceae" corrig. Probst et al. 2014 "Candidatus Altarchaeum" corrig. Probst et al. 2014
Phylum " Iainarchaeota "
| Phylogeny of "Iainarchaeota" |
| "Iainarchaeia" "Forterreales" "Forterreaceae" "Ca. Forterrea" "Iainarchaeales" "Iainarchaeaceae" "Ca. Iainarchaeum" |
| "Iainarchaeia" | "Forterreales" "Forterreaceae" "Ca. Forterrea" "Iainarchaeales" "Iainarchaeaceae" "Ca. Iainarchaeum" |
| "Forterreales" | "Forterreaceae" "Ca. Forterrea" |
| "Forterreaceae" | "Ca. Forterrea" |
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| "Iainarchaeales" | "Iainarchaeaceae" "Ca. Iainarchaeum" |
| "Iainarchaeaceae" | "Ca. Iainarchaeum" |
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Class "Iainarchaeia"
Order "Forterreales"
- Family "Forterreaceae" Pallen, Rodriguez-R & Alikhan 2022 [CSSED10-239] "Candidatus Forterrea" Probst & Banfield 2017
Order "Iainarchaeales"
- Family "Iainarchaeaceae" Rinke et al. 2020 "Candidatus Iainarchaeum" Rinke et al. 2013
Phylum " Micrarchaeota "
| Phylogeny of "Micrarchaeota" |
| "Micrarchaeia" "Norongarragalinales" "Norongarragalinaceae" "Ca. Norongarragalina" "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Burarchaeales" "Gugararchaeaceae" "Ca. Gugararchaeum" "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Micrarchaeia" | "Norongarragalinales" "Norongarragalinaceae" "Ca. Norongarragalina" "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Burarchaeales" "Gugararchaeaceae" "Ca. Gugararchaeum" "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Norongarragalinales" "Norongarragalinaceae" "Ca. Norongarragalina" |
| "Norongarragalinales" | "Norongarragalinaceae" "Ca. Norongarragalina" |
| "Norongarragalinaceae" | "Ca. Norongarragalina" |
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| "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Burarchaeales" "Gugararchaeaceae" "Ca. Gugararchaeum" "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" |
| "Micrarchaeales" | "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" |
| "Micrarchaeaceae" | "Ca. Mancarchaeum" "Ca. Micrarchaeum" |
| "Ca. Mancarchaeum" |
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| "Ca. Micrarchaeum" |
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| "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Burarchaeales" "Gugararchaeaceae" "Ca. Gugararchaeum" "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Anstonellales" | "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" |
| "Anstonellaceae" | "Ca. Anstonella" |
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| "Bilamarchaeaceae" | "Ca. Bilamarchaeum" |
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| "Burarchaeales" | "Gugararchaeaceae" "Ca. Gugararchaeum" "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Gugararchaeaceae" "Ca. Gugararchaeum" |
| "Gugararchaeaceae" | "Ca. Gugararchaeum" |
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| "Burarchaeaceae" "Ca. Burarchaeum" "Fermentimicrarchaeaceae" "Ca. Fermentimicrarchaeum" |
| "Burarchaeaceae" | "Ca. Burarchaeum" |
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| "Fermentimicrarchaeaceae" | "Ca. Fermentimicrarchaeum" |
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Class "Micrarchaeia"
Order "Norongarragalinales"
- Family "Norongarragalinaceae" Vázquez-Campos et al. 2021 "Candidatus Norongarragalina" Vázquez-Campos et al. 2021
Order "Micrarchaeales"
- Family "Micrarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Mancarchaeum" Golyshina et al. 2017 "Candidatus Micrarchaeum" Baker et al. 2010
Order "Anstonellales"
- Family "Anstonellaceae" Vázquez-Campos et al. 2021 "Candidatus Anstonella" Vázquez-Campos et al. 2021
- Family "Bilamarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Bilamarchaeum" Vázquez-Campos et al. 2021
Order "Burarchaeales"
- Family "Gugararchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Gugararchaeum" Vázquez-Campos et al. 2021
- Family "Burarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Burarchaeum" Vázquez-Campos et al. 2021
- Family "Fermentimicrarchaeaceae" Kadnikov et al. 2020 (Sv326) "Candidatus Fermentimicrarchaeum" Kadnikov et al. 2020
Phylum Promethearchaeota
Taxonomy of this group is based on LPSN, and also these works:
| Phylogeny of Promethearchaeota |
| "Asgardarchaeia" "Asgardarchaeales" "Thorarchaeia" "Thorarchaeales" "Atabeyarchaeia" "Atabeyarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Asgardarchaeia" "Asgardarchaeales" "Thorarchaeia" "Thorarchaeales" "Atabeyarchaeia" "Atabeyarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Asgardarchaeia" "Asgardarchaeales" "Thorarchaeia" "Thorarchaeales" |
| "Asgardarchaeia" | "Asgardarchaeales" |
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| "Thorarchaeia" | "Thorarchaeales" |
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| "Atabeyarchaeia" "Atabeyarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Atabeyarchaeia" "Atabeyarchaeales" |
| "Atabeyarchaeia" | "Atabeyarchaeales" |
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| "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales |
| "Baldrarchaeia" "Baldrarchaeales" "Jordiarchaeia" "Jordiarchaeales" |
| "Baldrarchaeia" | "Baldrarchaeales" |
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| "Jordiarchaeia" | "Jordiarchaeales" |
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| "Odinarchaeia" "Odinarchaeales" "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales |
| "Odinarchaeia" "Odinarchaeales" |
| "Odinarchaeia" | "Odinarchaeales" |
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| "Hermodarchaeia" "Hermodarchaeales" Promethearchaeia "Helarchaeales" Promethearchaeales |
| "Hermodarchaeia" | "Hermodarchaeales" |
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| Promethearchaeia | "Helarchaeales" Promethearchaeales |
| "Helarchaeales" |
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| Promethearchaeales |
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| "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Borrarchaeia" "Borrarchaeales" "Sifarchaeales" |
| "Borrarchaeia" | "Borrarchaeales" "Sifarchaeales" |
| "Borrarchaeales" |
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| "Sifarchaeales" |
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| "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Wukongarchaeia" "Wukongarchaeales" |
| "Wukongarchaeia" | "Wukongarchaeales" |
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| "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Njordarchaeia" | "Njordarchaeales" |
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| "Heimdallarchaeia" | "Hodarchaeales" "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Hodarchaeales" |
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| "Gerdarchaeales" (JABLTI01) "Heimdallarchaeales" |
| "Gerdarchaeales" (JABLTI01) |
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| "Heimdallarchaeales" |
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Promethearchaeota incertae sedis
- ?"Friggarchaeota" Caceres 2019
- ?"Gefionarchaeota" Caceres 2019
- ?"Idunnarchaeota" Caceres 2019
- ?"Kexuearchaeota" Cai et al. 2021
- ?"Sleipnirarchaeota" Kioukis et al. 2025
- ?"Candidatus Balderarchaeum" Xie et al. 2022
- ?"Candidatus Ranarchaeum" Appler et al. 2026
- ?"Candidatus Sukunaarchaeum" Harada et al. 2025
- ?"Candidatus Tyrarchaeum" Xie et al. 2022
Class "Asgardarchaeia"
Order "Asgardarchaeales"
- Family "Asgardarchaeaceae" Eme et al. 2023 ex Tamarit et al. 2024 "Candidatus Asgardarchaeum" Eme et al. 2023 ex Tamarit et al. 2024
Class " Thorarchaeia "
Order "Thorarchaeales"
- Family "Thorarchaeaceae" (MBG-B)
Class "Atabeyarchaeia"
Order "Atabeyarchaeales"
- Family "Atabeyarchaeaceae" Valentin-Alvaradoet al. 2024 "Candidatus Atabeyarchaeum" Valentin-Alvarado et al. 2024
Class "Baldrarchaeia"
Order "Baldrarchaeales"
- Family "Baldrarchaeaceae" corrig. Liu et al. 2021 "Candidatus Baldrarchaeum" Liu et al. 2021
Class "Jordarchaeia"
Order "Jordarchaeales"
- Family "Freyrarchaeaceae" Xie et al. 2022 "Candidatus Freyrarchaeum" Xie et al. 2022
- Family "Jordarchaeaceae" Sun et al. 2021 "Candidatus Jordarchaeum" Sun et al. 2021 ["Ca. Freyarchaeum" (sic) Valentin-Alvarado et al. 2024]
Class "Odinarchaeia"
Order "Odinarchaeales"
- Family "Odinarchaeaceae" Tamarit et al. 2022 "Candidatus Odinarchaeum" Tamarit et al. 2022
Order " Yangjianarchaeales "
Class "Hermodarchaeia"
Order "Hermodarchaeales"
- Family "Hermodarchaeaceae" corrig. Liu et al. 2021 "Candidatus Hermodarchaeum" Liu et al. 2021
Class Promethearchaeia
Order "Helarchaeales"
Order Promethearchaeales
| Phylogeny of "Promethearchaeales" |
| "Sigynarchaeaceae" "Ca. Sigynarchaeum" Promethearchaeaceae Prometheoarchaeum "Ca. Harpocratesius" "Ca. Lokiarchaeum" |
| "Sigynarchaeaceae" "Ca. Sigynarchaeum" Promethearchaeaceae Prometheoarchaeum "Ca. Harpocratesius" "Ca. Lokiarchaeum" |
| "Sigynarchaeaceae" | "Ca. Sigynarchaeum" |
|
| Promethearchaeaceae | Prometheoarchaeum "Ca. Harpocratesius" "Ca. Lokiarchaeum" |
| Prometheoarchaeum |
|
| "Ca. Harpocratesius" "Ca. Lokiarchaeum" |
| "Ca. Harpocratesius" |
|
| "Ca. Lokiarchaeum" |
|
|
|
|
- Family "Sigynarchaeaceae" Xie et al. 2022 "Candidatus Sigynarchaeum" Xie et al. 2022
- Family Promethearchaeaceae Imachi et al. 2024 "Candidatus Harpocratesius" Wu et al. 2022 "Candidatus Lokiarchaeum" corrig. Spang et al. 2015 (MBGB, DSAG) ?Nerearchaeum Nobs et al. 2026 Promethearchaeum Imachi et al. 2024
Class "Borrarchaeia"
Order "Sifarchaeales"
- Family "Sifarchaeaceae" Sun et al. 2021 "Candidatus Sifarchaeum" corrig. Farag, Zhao & Biddle 2020
Order "Borrarchaeales"
- Family "Borrarchaeaceae" Liu et al. 2021 "Candidatus Borrarchaeum" Liu et al. 2021
Class "Wukongarchaeia"
Order "Wukongarchaeales"
- Family "Wukongarchaeaceae" Liu et al. 2021 "Candidatus Wukongarchaeum" Liu et al. 2021
Class "Njordarchaeia"
Order "Njordarchaeales"
- Family "Panguiarchaeaceae" Qu et al. 2023 "Panguiarchaeum" Qu et al. 2023
- Family "Njordarchaeaceae" Xie et al. 2022 "Candidatus Njordarchaeum" Xie et al. 2022
Class "Heimdallarchaeia"
- ?"Candidatus Yibarchaeum" MacLeod et al. 2025
Order " Wenzhongarchaeales "
Order "Hodarchaeales"
- Family "Hodarchaeaceae" Liu et al. 2021 "Candidatus Hodarchaeum" Liu et al. 2021
- Family "Margulisarchaeaceae" Imachi et al. 2025 ?"Candidatus Flexarchaeum" Imachi et al. 2025 ?"Candidatus Margulisarchaeum" Imachi et al. 2025
Order "Gerdarchaeales"
Order "Heimdallarchaeales"
- Family "Kariarchaeaceae" Liu et al. 2021 "Candidatus Kariarchaeum" Liu et al. 2021
- Family "Heimdallarchaeaceae" Wu et al. 2022 "Candidatus Heimdallarchaeum" Wu et al. 2022
Phylum " Korarchaeota "
| Phylogeny of "Korarchaeota" |
| "Korarchaeia" "Korarchaeaceae" "Ca. Korarchaeum" "Ca. Methanodesulfokora" |
| "Korarchaeia" | "Korarchaeaceae" "Ca. Korarchaeum" "Ca. Methanodesulfokora" |
| "Korarchaeaceae" | "Ca. Korarchaeum" "Ca. Methanodesulfokora" |
| "Ca. Korarchaeum" |
|
| "Ca. Methanodesulfokora" |
|
|
|
Class "Korarchaeia"
Order "Korarchaeales"
- Family "Korarchaeaceae" Rinke et al. 2020 "Candidatus Korarchaeum" Elkins et al. 2008 "Candidatus Methanodesulfokora" McKay et al. 2019
Phylum Thermoproteota
| Phylogeny of Thermoproteota |
| "BAT" CALKCG01 "Bifangarchaeales" "Bathyarchaeia" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" Nitrososphaeria_A "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales "Sulfobacteria" Methanosuratincolia "Methanonezhaarchaeales" "Culexarchaeales" Methanosuratincolales "Thermoproteia" "Gearchaeales" Thermofilales Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales |
| "BAT" CALKCG01 "Bifangarchaeales" "Bathyarchaeia" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" Nitrososphaeria_A "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales "Sulfobacteria" Methanosuratincolia "Methanonezhaarchaeales" "Culexarchaeales" Methanosuratincolales "Thermoproteia" "Gearchaeales" Thermofilales Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales |
| "BAT" | CALKCG01 "Bifangarchaeales" "Bathyarchaeia" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" Nitrososphaeria_A "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales |
| CALKCG01 "Bifangarchaeales" "Bathyarchaeia" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" |
| CALKCG01 | "Bifangarchaeales" |
|
| "Bathyarchaeia" | "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" |
| "Xuanwuarculales" "Hecatellales" "Houtuarculales" |
| "Xuanwuarculales" |
|
| "Hecatellales" "Houtuarculales" |
| "Hecatellales" |
|
| "Houtuarculales" |
|
|
|
| "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" |
| "Wuzhiqiibiales" |
|
| "Zhuquarculales" "Bathyarchaeales" |
| "Zhuquarculales" |
|
| "Bathyarchaeales" |
|
|
|
|
|
| Nitrososphaeria_A "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales |
| Nitrososphaeria_A | "Caldarchaeales" |
|
| Nitrososphaeria | "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales |
| "Geothermarchaeales" |
|
| PSMU01 Conexivisphaerales Nitrososphaerales |
| PSMU01 |
|
| Conexivisphaerales Nitrososphaerales |
| Conexivisphaerales |
|
| Nitrososphaerales |
|
|
|
|
|
|
| "Sulfobacteria" | Methanosuratincolia "Methanonezhaarchaeales" "Culexarchaeales" Methanosuratincolales "Thermoproteia" "Gearchaeales" Thermofilales Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales |
| Methanosuratincolia "Methanonezhaarchaeales" "Culexarchaeales" Methanosuratincolales "Thermoproteia" "Gearchaeales" Thermofilales Thermoproteales |
| Methanosuratincolia | "Methanonezhaarchaeales" "Culexarchaeales" Methanosuratincolales |
| "Methanonezhaarchaeales" "Culexarchaeales" |
| "Methanonezhaarchaeales" |
|
| "Culexarchaeales" |
|
|
| Methanosuratincolales |
|
|
| "Thermoproteia" | "Gearchaeales" Thermofilales Thermoproteales |
| "Gearchaeales" Thermofilales Thermoproteales |
| "Gearchaeales" |
|
| Thermofilales Thermoproteales |
| Thermofilales |
|
| Thermoproteales |
|
|
|
|
|
| "Sulfolobia" "Tardisphaerales" Sulfolobales |
| "Sulfolobia" | "Tardisphaerales" Sulfolobales |
| "Tardisphaerales" |
|
| Sulfolobales |
|
|
|
|
|
Thermoproteota Incertae sedis
"Brockiarchaeota"
Class CALKCG01
Order "Bifangarchaeales"
- Family "Bifangicolaceae" Hou et al. 2023 ?"Candidatus Bifangicola" Hou et al. 2023 [3300025546_26]
- Family "Bifangarchaeaceae" Hou et al. 2023 [B24] "Candidatus Bifangarchaeum" corrig. Hou et al. 2023 [DTBU01 sp011374155]
- Family "Bifangistellaceae" Hou et al. 2023 [DRYS01] "Candidatus Bifangistella" Hou et al. 2023 [DRYS01 sp011367275]
Class " Bathyarchaeia "
Order "Xuanwuarculales"
- Family "Mazuisomaceae" Hou et al. 2023 ?"Candidatus Mazuisoma" Hou et al. 2023
- Family "Mazuicellaceae" Hou et al. 2023 ?"Candidatus Mazuibellus" Hou et al. 2023 ?"Candidatus Mazuicella" Hou et al. 2023
- Family "Mazuousiaecae" Hou et al. 2023 ?"Candidatus Mazuarcula" Hou et al. 2023 ?"Candidatus Mazuousia" Hou et al. 2023
- Family "Xuanwuarculaceae" Hou et al. 2023 "Candidatus Xuanwuarcula" Hou et al. 2023 [RBG-16-48-13 sp001775995]
Order "Hecatellales"
| Phylogeny of "Hecatellales" |
| "Hecatellaceae" "Ca. Hecatella" "Ca. Jinwuarcula" "Ca. Jinwuicola" "Ca. Jinwuimorpha" |
| "Hecatellaceae" | "Ca. Hecatella" "Ca. Jinwuarcula" "Ca. Jinwuicola" "Ca. Jinwuimorpha" |
| "Ca. Hecatella" "Ca. Jinwuarcula" |
| "Ca. Hecatella" |
|
| "Ca. Jinwuarcula" |
|
|
| "Ca. Jinwuicola" "Ca. Jinwuimorpha" |
| "Ca. Jinwuicola" |
|
| "Ca. Jinwuimorpha" |
|
|
|
- Family "Hecatellaceae" Rinke et al. 2020 ["Jinwuarculaceae" Hou et al. 2023; "Jinwuicolaceae" Hou et al. 2023; "Jinwuimonadaceae" Hou et al. 2023; "Jinwuiprofundaceae" Hou et al. 2023; "Jinwuousiaceae" Hou et al. 2023] "Candidatus Hecatella" Adam et al. 2022 "Candidatus Jinwuarcula" Hou et al. 2023 [B25] "Candidatus Jinwuicola" Hou et al. 2023 [EX4484-218 sp002254975] ?"Candidatus Jinwuimonas" Hou et al. 2023 "Candidatus Jinwuimorpha" Hou et al. 2023 [EX4484-218 sp003661965] ?"Candidatus Jinwuiprofundus" Hou et al. 2023 ?"Candidatus Jinwuousia" Hou et al. 2023 [JdFR-11 sp002011035]
Order "Houtuarculales"
| Phylogeny of "Houtuarculales" |
| "Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus" "Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Ca. Houtuarcula" |
| "Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus" "Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Ca. Houtuarcula" |
| "Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus" |
| "Crenohoutuithermaceae" | "Ca. Crenohoutuithermus" |
|
| "Fontihoutuithermaceae" | "Ca. Fontihoutuithermus" |
|
|
| "Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Ca. Houtuarcula" |
| "Aphrohoutuibellaceae" | "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" |
| "Ca. Aphrohoutuibellus" |
|
| "Ca. Hydrohoutuibellus" |
|
|
| "Houtuousiaceae" | "Ca. Houtuousia" "Ca. Houtuarcula" |
| "Ca. Houtuousia" |
|
| "Ca. Houtuarcula" |
|
|
|
|
- Family "Houtuithermaceae" Hou et al. 2023 ?"Candidatus Houtuithermus" Hou et al. 2023
- Family "Marihoutuimonadaceae" Hou et al. 2023 ?"Candidatus Houtuibellus" Hou et al. 2023 ?"Candidatus Marihoutuimonas" Hou et al. 2023
- Family "Crenohoutuithermaceae" Hou et al. 2023 "Candidatus Crenohoutuithermus" Hou et al. 2023 [DTDX01]
- Family "Fontihoutuithermaceae" Hou et al. 2023 ["Fluxihoutuithermaceae" Hou et al. 2023; DTGE01] ?"Candidatus Fluxihoutuithermus" Hou et al. 2023 [DTOY01 sp011358085] "Candidatus Fontihoutuithermus" Hou et al. 2023 [DTGE01 sp011389385]
- Family "Aphrohoutuibellaceae" Hou et al. 2023 [RBG-13-38-9] "Candidatus Aphrohoutuibellus" Hou et al. 2023 [RBG-13-38-9] "Candidatus Hydrohoutuibellus" Hou et al. 2023 [DTMT01 sp011355205]
- Family "Houtuousiaceae" Hou et al. 2023 ["Houtuarculaceae" Hou et al. 2023; "Limnohoutuiplasmataceae" corrig. Hou et al. 2023; 40CM-2-53-6] "Candidatus Houtuousia" Hou et al. 2023 [40CM-2-53-6] "Candidatus Houtuarcula" Hou et al. 2023 [FEN-987] ?"Candidatus Limnohoutuiplasma" Hou et al. 2023
Order "Wuzhiqiibiales"
| Phylogeny of "Wuzhiqiibiales" |
| "Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha" "Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium" |
| "Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha" "Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium" |
| "Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha" |
| "Wuzhiqiimorphaceae" | "Ca. Wuzhiqiimorpha" |
|
|
| "Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium" |
| "Wuzhiqiimonadaceae" | "Ca. Wuzhiqiimonas" |
|
| "Wuzhiqiibiaceae" | "Ca. Wuzhiqiibium" |
|
|
|
- Family "Wuzhiqiithermaceae" Hou et al. 2023 ?"Candidatus Wuzhiqiithermus" Hou et al. 2023
- Family "Wuzhiqiitorraceae" Hou et al. 2023 ?"Candidatus Wuzhiqiitorris" Hou et al. 2023
- Family "Wuzhiqiimorphaceae" Hou et al. 2023 [PIYN01] "Candidatus Wuzhiqiimorpha" Hou et al. 2023 [PIYN01 sp003601775]
- Family "Wuzhiqiimonadaceae" Hou et al. 2023 [B23] "Candidatus Wuzhiqiimonas" Hou et al. 2023 [B23]
- Family "Wuzhiqiibiaceae" Hou et al. 2023 "Candidatus Wuzhiqiibium" Hou et al. 2023 [TCS64]
Order "Zhuquarculales"
- Family "Zhuquarculaceae" Hou et al. 2023 "Candidatus Zhuquarcula" Hou et al. 2023 [EX4484-135]
- Family "Zhuquomorphaceae" Hou et al. 2023 [B24-G1] "Candidatus Zhuquomorpha" Hou et al. 2023
Order "Bathyarchaeales"
| Phylogeny of "Bathyarchaeales" |
| "Pontizhurongicolaceae" "Ca. Pontizhurongicola" "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Pontizhurongicolaceae" "Ca. Pontizhurongicola" "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Pontizhurongicolaceae" "Ca. Pontizhurongicola" |
| "Pontizhurongicolaceae" | "Ca. Pontizhurongicola" |
|
|
| "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" |
| "Hanbarchaeaceae" | "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" |
| "Ca. Hanbarchaeum" |
|
| "Ca. Fufeiimorpha" "Ca. Zhurongicella" |
| "Ca. Fufeiimorpha" |
|
| "Ca. Zhurongicella" |
|
|
|
| "Marizhurongicellaceae" | "Ca. Marizhurongibium" "Ca. Marizhurongicella" |
| "Ca. Marizhurongibium" |
|
| "Ca. Marizhurongicella" |
|
|
|
| "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Hanbopyraceae" "Ca. Hanbopyrus" |
| "Hanbopyraceae" | "Ca. Hanbopyrus" |
|
|
| "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" |
| "Jingweiiplasmataceae" | "Ca. Jingweiiplasma" |
|
| "Bathyarchaeaceae" | "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" |
| "Ca. Baizosoma" |
|
| "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" |
| "Ca. Halozhurongicella" |
|
| "Ca. Baizobellus" "Ca. Bathyarchaeum" |
| "Ca. Baizobellus" |
|
| "Ca. Bathyarchaeum" |
|
|
|
|
|
| "Baizomorphaceae" "Ca. Baizomorpha" "Bathycorpusculaceae" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Baizomorphaceae" | "Ca. Baizomorpha" |
|
| "Bathycorpusculaceae" | "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" |
| "Ca. Buyanhuyuitorris" |
|
| "Ca. Buyanhuyuicella" |
|
|
| "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Ca. Hanbocola" "Ca. Hanbothermus" |
| "Ca. Hanbocola" |
|
| "Ca. Hanbothermus" |
|
|
| "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Ca. Yanziithermus" |
|
| "Ca. Yuguotorris" "Ca. Bathycorpusculum" |
| "Ca. Yuguotorris" |
|
| "Ca. Bathycorpusculum" |
|
|
|
|
|
|
|
|
|
|
- Family "Actozhurongousiaceae" Hou et al. 2023 ?"Candidatus Actozhurongousia" Hou et al. 2023
- Family "Yuguousiaceae" Hou et al. 2023 ?"Candidatus Yuguousia" Hou et al. 2023
- Family "Pontizhurongicolaceae" Hou et al. 2023 "Candidatus Pontizhurongicola" Hou et al. 2023 [B24-2 sp002490245]
- Family "Hanbarchaeaceae" Hou et al. 2023 ["Zhurongicellaceae" Hou et al. 2023; DTEX01] "Candidatus Fufeiimorpha" Hou et al. 2023 [PIYB01] "Candidatus Hanbarchaeum" Hou et al. 2023 [DTEX01] "Candidatus Zhurongicella" Hou et al. 2023 [B63]
- Family "Marizhurongicellaceae" Hou et al. 2023 [B26-1] ?"Candidatus Aoguangisoma" Hou et al. 2023 "Candidatus Marizhurongibium" Hou et al. 2023 [B17-G17 sp003662645] "Candidatus Marizhurongicella" Hou et al. 2023 [B26-1] ?"Candidatus Yanziimonas" Hou et al. 2023 ?"Candidatus Zhurongibium" Hou et al. 2023
- Family "Hanbopyraceae" Hou et al. 2023 "Candidatus Hanbopyrus" Hou et al. 2023 [WUQV01 sp009889585]
- Family "Jingweiiplasmataceae" Hou et al. 2023 [SOJC01] "Candidatus Jingweiiplasma" Hou et al. 2023 [SOJC01]
- Family "Bathyarchaeaceae" Khomyakova et al. 2023 ["Baizosomaceae" Hou et al. 2023; "Baizostellaceae" Hou et al. 2023; "Halozhurongicellaceae" Hou et al. 2023] "Candidatus Baizobellus" Hou et al. 2023 [SOJZ01] ?"Candidatus Baizosediminiarchaeum" Yu et al. 2023 [BIN-L-1] "Candidatus Baizosoma" Hou et al. 2023 [BA1 sp001399805] ?"Candidatus Baizostella" Hou et al. 2023 "Candidatus Bathyarchaeum" Khomyakova et al. 2023 "Candidatus Halozhurongicella" Hou et al. 2023 [B1-G15]
- Family "Baizomorphaceae" Hou et al. 2023 [UBA233] "Candidatus Baizomorpha" Hou et al. 2023
- Family "Bathycorpusculaceae" Loh & Brune 2023 ["Aoshunithermaceae" Hou et al. 2023; "Baizomonadaceae" Hou et al. 2023; "Buyanhuyuicellaceae" Hou et al. 2023; "Hanbothermaceae" Hou et al. 2023; "Yanziithermaceae" Hou et al. 2023; "Yuguotorraceae" corrig. Hou et al. 2023] ?"Candidatus Aoshunithermus" Hou et al. 2023 ?"Candidatus Baizomonas" Hou et al. 2023 "Candidatus Bathycorpusculum" Loh & Brune 2023 ["Ca. Hafrusarcha" Pallen, Rodriguez-R & Alikhan 2022] "Candidatus Buyanhuyuicella" Hou et al. 2023 [SOJA01] "Candidatus Buyanhuyuitorris" Hou et al. 2023 [PIYA01] "Candidatus Hanbocola" Hou et al. 2023 [DSZD01] "Candidatus Hanbothermus" Hou et al. 2023 [DRVP01] "Candidatus Yanziithermus" Hou et al. 2023 [JdFR-07] "Candidatus Yuguotorris" Hou et al. 2023 [DSLH01]
Class Nitrososphaeria_A
Order "Caldarchaeales"
| Phylogeny of "Caldarchaeales" |
| "Calditenuaceae" "Ca. Calditenuis" "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum" "Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Calditenuaceae" "Ca. Calditenuis" "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum" "Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Calditenuaceae" "Ca. Calditenuis" |
| "Calditenuaceae" | "Ca. Calditenuis" |
|
|
| "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum" "Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Caldarchaeaceae" | "Ca. Caldarchaeum" "Ca. Pelearchaeum" |
| "Ca. Caldarchaeum" |
|
| "Ca. Pelearchaeum" |
|
|
| "Wolframiiraptoraceae" | "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Ca. Terraquivivens" |
|
| "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Ca. Geocrenenecus" |
|
| "Ca. Benthortus" "Ca. Wolframiiraptor" |
| "Ca. Benthortus" |
|
| "Ca. Wolframiiraptor" |
|
|
|
|
|
|
- Family "Calditenuaceae" Balbay et al. 2023 "Candidatus Calditenuis" Beam et al. 2016
- Family "Caldarchaeaceae" Rinke et al. 2020 "Candidatus Caldarchaeum" corrig. Nunoura et al. 2011 "Candidatus Pelearchaeum" Balbay et al. 2023 ["Ca. Cadrafarcha" Pallen, Rodriguez-R & Alikhan 2022]
- Family "Wolframiiraptoraceae" Buessecker et al. 2022 "Candidatus Benthortus" Buessecker et al. 2022 "Candidatus Geocrenenecus" Buessecker et al. 2022 "Candidatus Terraquivivens" Buessecker et al. 2022 "Candidatus Wolframiiraptor" Buessecker et al. 2022
Class Nitrososphaeria
Nitrososphaeria incertae sedis
- ?"Candidatus Cenoporarchaeum" corrig. Zhang et al. 2019
- ?"Candidatus Gigantothauma" corrig. Muller et al. 2010
- ?"Candidatus Nitrosodeserticola" Hwang et al. 2021
Order "Geothermarchaeales"
- Family "Geothermarchaeaceae" Adam et al. 2022 "Candidatus Geothermarchaeum" Adam et al. 2022 "Candidatus Scotarchaeum" Adam et al. 2022
Order PSMU01
- Family PSMU01 "Candidatus Australarchaeum" Herbold et al. 2024
Order Conexivisphaerales
- Family Conexivisphaeraceae Kato et al. 2021 (THSC Group) Conexivisphaera Kato et al. 2021
- Family "Thermosulfuridaceae" Chang et al. 2026 "Acidarchaeum" Chang et al. 2026 [JAJZYL01] ?"Thermosulfuris" Chang et al. 2026
Order Nitrososphaerales
| Phylogeny of Nitrososphaerales |
| "Gagatemarchaeaceae" "Ca. Gagatemarchaeum" "Nitrosomiraceae" "Ca. Nitrosomirus" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosobungeria" Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Gagatemarchaeaceae" "Ca. Gagatemarchaeum" "Nitrosomiraceae" "Ca. Nitrosomirus" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosobungeria" Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Gagatemarchaeaceae" "Ca. Gagatemarchaeum" |
| "Gagatemarchaeaceae" | "Ca. Gagatemarchaeum" |
|
|
| "Nitrosomiraceae" "Ca. Nitrosomirus" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosobungeria" Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Nitrosomiraceae" "Ca. Nitrosomirus" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" |
| "Nitrosomiraceae" | "Ca. Nitrosomirus" |
|
| "Nitrosocaldaceae" | "Ca. Nitrosothermus" "Ca. Nitrosocaldus" |
| "Ca. Nitrosothermus" |
|
| "Ca. Nitrosocaldus" |
|
|
|
| Nitrososphaeraceae "Ca. Nitrosobungeria" Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| Nitrososphaeraceae | "Ca. Nitrosobungeria" Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" |
| "Ca. Nitrosobungeria" |
|
| Nitrososphaera_A "Ca. Nitrosopolaris" Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" |
| Nitrososphaera_A "Ca. Nitrosopolaris" |
| Nitrososphaera_A |
|
| "Ca. Nitrosopolaris" |
|
|
| Nitrososphaera "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" |
| Nitrososphaera |
|
| "Ca. Nitrosomicrobium" "Ca. Nitrosocosmicus" |
| "Ca. Nitrosomicrobium" |
|
| "Ca. Nitrosocosmicus" |
|
|
|
|
|
| Nitrosopumilaceae | "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Nitrosotalea" |
|
| "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Nitrosotenuis" |
|
| "Ca. Nitrosopelagicus" "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Nitrosopelagicus" |
|
| "Ca. Nitrosoabyssus" "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Nitrosoabyssus" |
|
| "Ca. Cenarchaeum" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Cenarchaeum" |
|
| Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus |
| Nitrosarchaeum |
|
| "Ca. Nitrosomaritimum" Nitrosopumilus |
| "Ca. Nitrosomaritimum" |
|
| Nitrosopumilus |
|
|
|
|
|
|
|
|
|
|
|
- Family "Methylarchaceae" Hua et al. 2019 ?"Candidatus Methylarchaeum" Hua et al. 2019 ?"Candidatus Methanotowutia" Ou et al. 2022
- Family "Gagatemarchaeaceae" Sheridan et al. 2023 [UBA183] "Candidatus Gagatemarchaeum" Sheridan et al. 2023 [Bog-1369; GCA_003164815.1] ?"Candidatus Subgagatemarchaeum" Sheridan et al. 2023 [Fn1; IMG2558309099]
- Family "Nitrosomiraceae" Zheng et al. 2024 [UBA213] "Candidatus Nitrosomirus" Zheng et al. 2024
- Family "Nitrosocaldaceae" Qin et al. 2016 "Candidatus Nitrosocaldus" de la Torre et al. 2008 "Candidatus Nitrosothermus" Luo et al. 2021
- Family Nitrososphaeraceae Stieglmeier et al. 2014 "Candidatus Nitrosobungeria" Tan et al. 2026 [TH5893] "Candidatus Nitrosocosmicus" corrig. Lehtovirta-Morley et al. 2016 "Candidatus Nitrosomicrobium" Tan et al. 2026 [TA-21] "Candidatus Nitrosopolaris" Pessi, Rutanen & Hultman 2022 ["Ca. Flebogarcha" Pallen, Rodriguez-R & Alikhan 2022] ?"Candidatus Nitrososappho" Singleton et al. 2025 Nitrososphaera Stieglmeier et al. 2014
- Family Nitrosopumilaceae Qin et al. 2017 "Ca. Cenarchaeum" DeLong & Preston 1996 Nitrosarchaeum corrig. Jung et al. 2018 "Candidatus Nitrosoabyssus" Garritano et al. 2024 ?"Candidatus Nitrosokoinonia" Glasl et al. 2023 "Candidatus Nitrosomaritimum" Zhao et al. 2024 "Candidatus Nitrosopelagicus" Santoro et al. 2015 Nitrosopumilus Qin et al. 2017 ["Ca. Nitrosomarinus" Ahlgren et al. 2017] ?"Candidatus Nitrosospongia" Moeller et al. 2019 "Candidatus Nitrosotalea" Lehtovirta 2011 "Candidatus Nitrosotenuis" Lebedeva et al. 2013
Class Methanosuratincolia
Order "Methanohydrogenicales"
- Family "Methanohydrogenicaceae" Berghuis et al. 2019 ?"Candidatus Methanohydrogenicus" Berghuis et al. 2019
Order "Methanomethylarchaeales"
- Family "Methanomethylarchaeceae" corrig. Hua et al. 2019 ?"Candidatus Methanomethylarchaeum" corrig. Hua et al. 2019
Order "Methanomethylovorales"
- Family "Methanomethylovoraceae" Hua et al. 2019 ?"Candidatus Methanomethylovorus" Hua et al. 2019
Order "Methanonezhaarchaeales"
- Family "Methanohydrogenotrophicaceae" Hua et al. 2019 ?"Candidatus Methanogearchaeum" corrig. Hua et al. 2019 ?"Candidatus Methanohydrogenotrophicum" Hua et al. 2019
- Family "Methanonezhaarchaeceae" Kohtz, Nupp & Hatzenpichler 2025 [WYZ-LMO8] "Candidatus Methanonezhaarchaeum" Kohtz, Nupp & Hatzenpichler 2025 [WYZ-LMO8]
- Family "Muzhaarchaeaceae" [B40-G2: B40-G2]
Order "Culexarchaeales"
| Phylogeny of "Culexarchaeales" |
| "Culexarchaeaceae" "Ca. Culexarchaeum" B10‑G16 "Ca. Culexmicrobium_B" "Culexmicrobiaceae" "Ca. Culexmicrobium" |
| "Culexarchaeaceae" "Ca. Culexarchaeum" B10‑G16 "Ca. Culexmicrobium_B" "Culexmicrobiaceae" "Ca. Culexmicrobium" |
| "Culexarchaeaceae" "Ca. Culexarchaeum" |
| "Culexarchaeaceae" | "Ca. Culexarchaeum" |
|
|
| B10‑G16 "Ca. Culexmicrobium_B" "Culexmicrobiaceae" "Ca. Culexmicrobium" |
| B10‑G16 | "Ca. Culexmicrobium_B" |
|
| "Culexmicrobiaceae" | "Ca. Culexmicrobium" |
|
|
|
- Family "Culexarchaeaceae" Kohtz et al. 2022 ["Methanomediaceae" Berghuis et al. 2019] "Candidatus Culexarchaeum" Kohtz et al. 2022 ["Ca. Methanomedium" Berghuis et al. 2019]
- Family "Culexmicrobiaceae" Kohtz et al. 2022 "Candidatus Culexmicrobium" Kohtz et al. 2022
- Family B10-G16 "Candidatus Culexmicrobium_B"
Order Methanosuratincolales
- Family Methanosuratincolaceae Wu et al. 2025 ?"Candidatus Methanoluiimicrobium" Wu et al. 2024 "Candidatus Methanomethylicus" Vanwonterghem et al. 2016 ?"Candidatus Methanostamsia" Wu et al. 2024 Methanosuratinicola Wu et al. 2025
Class Thermoproteia
Order "Gearchaeales"
- Family "Gearchaeaceae" corrig. ?Geoarchaeum (sic) Hunt et al. 2016
Order Thermofilales
- Family Thermofilaceae Burggraf, Huber & Stetter 1997 Infirmifilum Zayulina et al. 2021 Thermofilum Zillig & Gierl 1983
Order Thermoproteales
- Family Thermocladiaceae Chuvochina et al. 2024 Caldivirga Itoh et al. 1999 Thermocladium Itoh, Suzuki & Nakase 1998 Vulcanisaeta Itoh, Suzuki & Nakase 2002
- Family Thermoproteaceae Zillig & Stetter 1982 Pyrobaculum Huber, Kristjansson & Stetter 1988 Thermoproteus Zillig & Stetter 1982
Class "Sulfolobia"
Order "Tardisphaerales"
- Family "Marsarchaeaceae" (sic) (NAG2, NCGII)
- Family "Tardisphaerace" Prokofeva et al. 2025 ?"Tardisphaera" Prokofeva et al. 2025
Order Sulfolobales
- Family Desulfurococcaceae Zillig & Stetter 1983 Desulfurococcus Zillig & Stetter 1983 Staphylothermus Stetter & Fiala 1986 ?Sulfophobococcus Hensel et al. 1997 Thermogladius Osburn & Amend 2011 ex Kochetkova et al. 2016 Thermosphaera Huber et al. 1998 ?"Candidatus Zestomicrobium" Dzhuraeva, Bobodzhanova & Birkeland 2024
- Family Fervidicoccaceae Perevalova et al. 2010 Fervidicoccus Perevalova et al. 2010
- Family Pyrodictiaceae Burggraf, Huber & Stetter 1997 ?Geogemma Kashefi, Holmes & Lovley 2006 Hyperthermus Zillig, Holz & Wunderl 1991 Pyrodictium Stetter, König & Stackebrandt 1984 "Pyrofollis" Miyazaki et al. 2023 Pyrolobus Blochl et al. 1999
- Family Acidilobaceae Prokofeva et al. 2009 Aeropyrum Sako et al. 1996 Acidilobus Prokofeva et al. 2000 Caldisphaera Itoh et al. 2003 ["Ca. Hestiella" St-John and Reysenbach 2024] "Candidatus Calypsonella" St John and Reysenbach 2024 ?Stetteria Jochimsen, Peinemann-Simon & Thomm 1998 Thermodiscus Stetter 2003 "Candidatus Tiamatella" St John and Reysenbach 2024
- Family Ignisphaeraceae RChuvochina et al. 2024 Ignisphaera Niederberger et al. 2006
- Family "Zestosphaeraceae" Pallen, Rodriguez-R & Alikhan 2022 Zestosphaera St. John et al. 2021
- Family Ignicoccaceae Chuvochina et al. 2024 Ignicoccus Huber, Burggraf & Stetter 2000
- Family Sulfolobaceae Stetter 1989 Acidianus Segerer et al. 1986 ["Sulfosphaerellus" Zhong et al. 1982; Desulfurolobus Zillig & Böck 1987] "Candidatus Aramenus" Servín-Garcidueñas & Martínez-Romero 2014 ?"Caldariella" de Rosa, Gambacorta & Bu'lock 1975 Metallosphaera Huber et al. 1989 Saccharolobus Sakai & Kurosawa 2018 Stygiolobus Segerer et al. 1991 Sulfodiicoccus Sakai & Kurosawa 2017 Sulfolobus Brock et al. 1972 Sulfuracidifex Itoh et al. 2020 Sulfurisphaera Kurosawa et al. 1998 ?Sulfurococcus Golovacheva, Vale’kho & Troitskii 1995
Phylum AGJL01
Class " Methanococcia "
Order Methanococcales
- Family Methanocaldococcaceae Whitman 2002 Methanocaldococcus Whitman 2002
- Family Methanococcaceae Balch & Wolfe 1981 Methanococcus Kluyver & van Niel 1936 Methanofervidicoccus Sakai et al. 2019 Methanothermococcus Whitman 2002 Methanotorris Whitman 2002 ["Methanoignis" Boone, Whitman & Rouviére 1993]
Phylum JAADCQ01
Class " Methanopyria "
Order Methanopyrales
- Family Methanopyraceae Huber & Stetter 2002 Methanopyrus Kurr et al. 1992
Phylum "Hydrothermarchaeota"
Class "Hydrothermarchaeia"
Order "Hydrothermarchaeales"
- Family "Hydrothermarchaeaceae" Chuvochina et al. 2019 "Candidatus Hydrothermarchaeum" Chuvochina et al. 2019
- Family BMS3B "Candidatus Pyrohabitans" Adam et al. 2022
Phylum " Hadarchaeota "
Class "Persephonarchaeia"
Class "Hadarchaeia"
Order "Hadarchaeales"
- Family "Cerberiarchaeaceae" Benito Merino et al. 2024 ?"Candidatus Cerberiarchaeum" Benito Merino et al. 2024 ?"Candidatus Melinoarchaeum" Yu et al. 2024
- Family "Hadarchaeaceae" Chuvochina et al. 2019 ex Chuvochina et al. 2023 "Candidatus Hadarchaeum" Chuvochina et al. 2019 ?"Candidatus Hadesarchaeum" Hua et al. 2019 ?"Candidatus Methanourarchaeum" Hua et al. 2019
Phylum Methanobacteriota_B
Class "Theionarchaeia"
Class " Thermococcia "
Order "Methanofastidiosales"
- Family "Methanofastidiosaceae" Rinke et al. 2020 "Candidatus Methanofastidiosum" Nobu et al. 2016
Order Thermococcales
- Family Thermococcaceae Zillig et al. 1988 Palaeococcus Takai et al. 2000 Pyrococcus Fiala & Stetter 1986 Thermococcus Zillig 1983 ["Caldococcus" Svetlichny et al. 1987]
Phylum Methanobacteriota
| Phylogeny of Methanobacteriota |
| "Methanobacteriia" DSM‑2088 Methanothermaceae Methanothermus Methanobacteriales DSM‑23052 Methanothermobacter_A Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| "Methanobacteriia" | DSM‑2088 Methanothermaceae Methanothermus Methanobacteriales DSM‑23052 Methanothermobacter_A Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| DSM‑2088 | Methanothermaceae Methanothermus |
| Methanothermaceae | Methanothermus |
|
|
| Methanobacteriales | DSM‑23052 Methanothermobacter_A Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| DSM‑23052 Methanothermobacter_A |
| DSM‑23052 | Methanothermobacter_A |
|
|
| Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| Methanothermobacteraceae | Methanothermobacter |
|
| Methanobacteriaceae | "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter |
| "Methanacia" "Methanoflexus" "Methanovirga" |
| "Methanacia" |
|
| "Methanoflexus" "Methanovirga" |
| "Methanoflexus" |
|
| "Methanovirga" |
|
|
|
| "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter |
| "Methanorudis" "Methanobaculum" "Methanobinarius" |
| "Methanorudis" |
|
| "Methanobaculum" "Methanobinarius" |
| "Methanobaculum" |
|
| "Methanobinarius" |
|
|
|
| "Methanocatella" Methanobrevibacter_A "Methanarmilla" Methanobrevibacter |
| "Methanocatella" |
|
| Methanobrevibacter_A "Methanarmilla" Methanobrevibacter |
| Methanobrevibacter_A |
|
| "Methanarmilla" Methanobrevibacter |
| "Methanarmilla" |
|
| Methanobrevibacter |
|
|
|
|
|
|
| Methanobacterium_E Methanobacterium_F Methanobacterium_D Methanosphaera Methanobacterium |
| Methanobacterium_E Methanobacterium_F |
| Methanobacterium_E |
|
| Methanobacterium_F |
|
|
| Methanobacterium_D Methanosphaera Methanobacterium |
| Methanobacterium_D |
|
| Methanosphaera Methanobacterium |
| Methanosphaera |
|
| Methanobacterium |
|
|
|
|
|
|
|
|
Class " Methanobacteriia "
Order DSM-2088
- Family Methanothermaceae Stetter et al. 1982 Methanothermus Stetter et al. 1982
Order Methanobacteriales
- Family Methanothermobacteraceae Chuvochina et al. 2024 Methanothermobacter Wasserfallen et al. 2000
- Family Methanobacteriaceae Barker 1956 "Methanacia" Protasov & Brune 2023 "Methanarmilla" Protasov & Brune 2023 Methanobacterium Kluyver & van Niel 1936 "Methanobaculum" Protasov & Brune 2023 "Methanobinarius" Protasov & Brune 2023 Methanobrevibacter Balch & Wolfe 1981 "Methanocatella" Protasov & Brune 2023 "Methanoflexus" Protasov & Brune 2023 "Methanorudis" Protasov & Brune 2023 Methanosphaera Miller & Wolin 1985 "Methanovirga" Protasov & Brune 2023
Phylum Thermoplasmatota
| Phylogeny of Thermoplasmatota |
| "Izemarchaea" "Thermoprofundales" "Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Izemarchaea" "Thermoprofundales" "Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Izemarchaea" "Thermoprofundales" |
| "Izemarchaea" | "Thermoprofundales" |
|
|
| "Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Poseidoniia" MGIII "Poseidoniales" |
| "Poseidoniia" | MGIII "Poseidoniales" |
| MGIII |
|
| "Poseidoniales" |
|
|
|
| "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Penumbrarchaeia" | "Penumbrarchaeales" |
|
| "Thermoplasmatia" | "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Proteinoplasmatales" "Yaplasmatales" "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales |
| "Proteinoplasmatales" "Yaplasmatales" |
| "Proteinoplasmatales" |
|
| "Yaplasmatales" |
|
|
| "Sysuiplasmatales" "Gimiplasmatales" Methanomassiliicoccales |
| "Sysuiplasmatales" |
|
| "Gimiplasmatales" Methanomassiliicoccales |
| "Gimiplasmatales" |
|
| Methanomassiliicoccales |
|
|
|
|
| "Halarchaeoplasmatales" "Lutacidiplasmatales" "Thermoplasmatales" |
| "Halarchaeoplasmatales" |
|
| "Lutacidiplasmatales" "Thermoplasmatales" |
| "Lutacidiplasmatales" |
|
| "Thermoplasmatales" |
|
|
|
|
|
|
|
Class "Izemarchaea"
Order "Thermoprofundales"
Class "Poseidoniia"
Order MGIII
Order "Poseidoniales"
- Family "Poseidoniaceae" Rinke et al. 2019 "Candidatus Poseidonia" Rinke et al. 2019
- Family "Thalassarchaeaceae" Rinke et al. 2019 "Candidatus Thalassarchaeum" corrig. Martin-Cuadrado et al. 2015
Class "Penumbrarchaeia"
Order "Penumbrarchaeales"
- Family "Penumbrarchaeaceae" Maeke et al. 2025 "Candidatus Penumbrarchaeum" Maeke et al. 2025
Class " Thermoplasmatia "
Order "Angelarchaeales"
Order "Luniplasmatales"
- Family "Lunaplasmataceae" (sic) Zinke et al. 2021 ?"Candidatus Lunaplasma" (sic) Zinke et al. 2021
Order "Proteinoplasmatales"
Order "Yaplasmatales"
- (RBG-16-68-12) [incl. "Detorarchales" Pallen & Alikhan 2021; "Limitiplasmatales" corrig. Zheng et al. 2022]
Order "Sysuiplasmatales"
- Family "Sysuiplasmataceae" Yuan et al. 2021 "Candidatus Sysuiplasma" Yuan et al. 2021
Order "Gimiplasmatales"
- Family "Gimiplasmataceae" Hu et al. 2020 "Candidatus Gimiplasma" Hu et al. 2020 (UA10834)
Order Methanomassiliicoccales
| Phylogeny of Methanomassiliicoccales |
| Methanomassiliicoccaceae Methanomassiliicoccus Methanomethylophilaceae "Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma" |
| Methanomassiliicoccaceae Methanomassiliicoccus Methanomethylophilaceae "Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma" |
| Methanomassiliicoccaceae | Methanomassiliicoccus |
|
| Methanomethylophilaceae | "Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma" |
| "Ca. Methanomicula" |
|
| "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma" |
| "Methanarcanum" |
|
| "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma" |
| "Methanoprimaticola" |
|
| Methanomethylophilus "Methanogranum" "Methanoplasma" |
| Methanomethylophilus |
|
| "Methanogranum" "Methanoplasma" |
| "Methanogranum" |
|
| "Methanoplasma" |
|
|
|
|
|
|
|
- Family "Methanomixtatrophicaceae" Hua et al. 2019 ?"Candidatus Methanomixtatrophicum" Hua et al. 2019
- Family Methanomassiliicoccaceae Iino et al. 2013 Methanomassiliicoccus Dridi et al. 2012
- Family Methanomethylophilaceae Gaci et al. 2014 ex Borrel et al. 2024 "Methanarcanum" Chibani et al. 2023 "Methanogranum" Iino et al. 2023 Methanomethylophilus Borrel et al. 2024 "Candidatus Methanomicula" Protasov & Brune 2023 ["Ca. Cribefarcha" Pallen, Rodriguez-R & Alikhan 2022] "Methanoplasma" Lang & Brune 2023 "Methanoprimaticola" Chibani et al. 2023 ?"Candidatus Methanospyradousia" Gilroy et al. 2021
Order "Halarchaeoplasmatales"
| Phylogeny of "Halarchaeoplasmatales" |
| "Halarchaeoplasmataceae" "Ca. Natronoplasma" "Ca. Saliniplasma" "Ca. Haladaptiplasma" "Ca. Halarchaeoplasma" |
| "Halarchaeoplasmataceae" | "Ca. Natronoplasma" "Ca. Saliniplasma" "Ca. Haladaptiplasma" "Ca. Halarchaeoplasma" |
| "Ca. Natronoplasma" |
|
| "Ca. Saliniplasma" "Ca. Haladaptiplasma" "Ca. Halarchaeoplasma" |
| "Ca. Saliniplasma" |
|
| "Ca. Haladaptiplasma" "Ca. Halarchaeoplasma" |
| "Ca. Haladaptiplasma" |
|
| "Ca. Halarchaeoplasma" |
|
|
|
|
- Family "Halarchaeoplasmataceae" Zhou et al. 2023 (PWKY01) "Candidatus Haladaptiplasma" corrig. Zhou et al. 2022 "Candidatus Halarchaeoplasma" Zhou et al. 2023 ["Ca. Haloplasma" Zhou et al. 2022 non Antunes et al. 2008] "Candidatus Natronoplasma" Zhou et al. 2022 "Candidatus Saliniplasma" Zhou et al. 2022
Order "Lutacidiplasmatales"
- Family "Lutacidiplasmataceae" Sheridan et al. 2022 "Candidatus Lutacidiplasma" Sheridan et al. 2022 (UBA184)
Order Thermoplasmatales
- Family "Aciduliprofundaceae" Rinke et al. 2020 "Aciduliprofundum" Reysenbach et al. 2006 (DHVE2 group)
- Family Thermoplasmataceae Reysenbach 2002 Acidiplasma Golyshina et al. 2009 ?"Candidatus Carboxiplasma" Bulaev et al. 2023 Cuniculiplasma corrig. Golyshina et al. 2016 Ferroplasma Golyshina et al. 2000 ?Oxyplasma Golyshina et al. 2024 Picrophilus Schleper et al. 1996 ?"Candidatus Scheffleriplasma" Krause et al. 2022 Thermogymnomonas Itoh, Yoshikawa & Takashina 2007 Thermoplasma Darland et al. 1970
Phylum Halobacteriota
| Phylogeny of Halobacteriota |
| Methanonatronarchaeia Methanonatronarchaeales "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales Bog‑38 "Methanoflorentales" Methanocellia Methanocellales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halobacteriales |
| Methanonatronarchaeia Methanonatronarchaeales "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales Bog‑38 "Methanoflorentales" Methanocellia Methanocellales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halobacteriales |
| Methanonatronarchaeia Methanonatronarchaeales |
| Methanonatronarchaeia | Methanonatronarchaeales |
|
|
| "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales Bog‑38 "Methanoflorentales" Methanocellia Methanocellales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halobacteriales |
| "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales Bog‑38 "Methanoflorentales" Methanocellia Methanocellales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales |
| "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales |
| "Methanoliparia" | "Methanoliparales" |
|
| "Archaeoglobia" | "Mnemosynellales" Archaeoglobales |
| "Mnemosynellales" |
|
| Archaeoglobales |
|
|
|
| Bog‑38 "Methanoflorentales" Methanocellia Methanocellales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales |
| Bog‑38 "Methanoflorentales" Methanocellia Methanocellales |
| Bog‑38 | "Methanoflorentales" |
|
| Methanocellia | Methanocellales |
|
|
| "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanosarcinia Methanosarcinales_A Methanotrichales Methanosarcinales |
| "Syntropharchaeia" | "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" |
| "Aerarchaeales" |
|
| "Syntropharchaeales" "Alkanophagales" |
| "Syntropharchaeales" |
|
| "Alkanophagales" |
|
|
|
| Methanosarcinia | Methanosarcinales_A Methanotrichales Methanosarcinales |
| Methanosarcinales_A Methanotrichales |
| Methanosarcinales_A |
|
| Methanotrichales |
|
|
| Methanosarcinales |
|
|
|
|
|
| "Methanomicrobia" Methanomicrobiales Halobacteria Halobacteriales |
| "Methanomicrobia" | Methanomicrobiales |
|
| Halobacteria | Halobacteriales |
|
|
|
|
Halobacteriota incertae sedis
- ?"Candidatus Methanoinsularis" Ou et al. 2022
- ?"Candidatus Methanoporticola" Ou et al. 2022
Class Methanonatronarchaeia
Order Methanonatronarchaeales
- Family Methanonatronarchaeaceae Sorokin et al. 2018 "Candidatus Methanohalarchaeum" Sorokin et al. 2018 Methanonatronarchaeum Sorokin et al. 2018
Class "Methanoliparia"
Order "Methanoliparales"
- Family "Methanoliparaceae" Borrel et al. 2019 "Candidatus Methanoliparum" Borrel et al. 2019 "Candidatus Methanolliviera" Borrel et al. 2019
Class " Archaeoglobia "
Order "Mnemosynellales"
- Family "Mnemosynellaceae" Adam et al. 2022 "Candidatus Mnemosynella" Adam et al. 2022
Order Archaeoglobales
- Family "Allopolytropaceae" corrig. Boyd et al. 2019 ?"Candidatus Allopolytropus" corrig. Boyd et al. 2019 ["Ca. Polytropus" Boyd et al. 2019 non Kirsch 1866]
- Family Archaeoglobaceae Huber & Stetter 2002 Archaeoglobus Stetter 1988 Ferroglobus Hafenbradl et al. 1997 Geoglobus Kashefi et al. 2002 ?"Candidatus Methanoglobus" Buessecker et al. 2023 ?"Candidatus Methanomixotrophus" corrig. Liu et al. 2021 ?"Candidatus Methanoproducendum" Hua et al. 2019
Class Bog-38
Order "Methanoflorentales"
- Family "Methanoflorentaceae" Mondav et al. 2014 "Candidatus Methanoflorens" Mondav et al. 2014
Class Methanocellia
Order Methanocellales
| Phylogeny of Methanocellales |
| Methanocellaceae Methanooceanicella Methanocella_A Methanocella |
| Methanocellaceae | Methanooceanicella Methanocella_A Methanocella |
| Methanooceanicella Methanocella_A |
| Methanooceanicella |
|
| Methanocella_A |
|
|
| Methanocella |
|
|
- Family Methanocellaceae Sakai et al. 2008 Methanocella Sakai et al. 2008 Methanooceanicella Zhang et al. 2024
Class "Syntropharchaeia"
Order "Aerarchaeales"
- Family YL-1 "Candidatus Methanodefluvium" Leu et al. 2025
- Family "Aerarchaeaceae" Leu et al. 2025 [CG-2] "Candidatus Aerarchaeum" Leu et al. 2025 (GG-2) "Candidatus Aerovita" Leu et al. 2025 (CM-1, CG-1, MW-1)
Order "Syntropharchaeales"
- Family "Syntropharchaeaceae" (ANME-2 cluster) "Candidatus Syntropharchaeum" corrig. Laso-Perez et al. 2016
Order "Alkanophagales"
| Phylogeny of "Alkanophagales" |
| "Alkanophagaceae" "Ca. Alkanophaga" "Methanospirantaceae" "Ca. Methanophaga" "Ca. Methanospirare" "Ca. Methanoxibalbensis" |
| "Alkanophagaceae" "Ca. Alkanophaga" "Methanospirantaceae" "Ca. Methanophaga" "Ca. Methanospirare" "Ca. Methanoxibalbensis" |
| "Alkanophagaceae" | "Ca. Alkanophaga" |
|
| "Methanospirantaceae" | "Ca. Methanophaga" "Ca. Methanospirare" "Ca. Methanoxibalbensis" |
| "Ca. Methanophaga" |
|
| "Ca. Methanospirare" "Ca. Methanoxibalbensis" |
| "Ca. Methanospirare" |
|
| "Ca. Methanoxibalbensis" |
|
|
|
|
- Family?"Veteromethanophagaceae" Vulcano et al. 2022
- Family "Alkanophagaceae" [B39-G2] "Candidatus Alkanophaga" Zehnle et al. 2023
- Family "Methanospirantaceae" corrig. Laso-Pérez et al. 2022 ?"Candidatus Methanoalium" Chadwick et al. 2022 "Candidatus Methanophaga" Chadwick et al. 2022 "Candidatus Methanospirans" corrig. Laso-Pérez et al. 2022 "Candidatus Methanoxibalbella" corrig. Laso-Pérez et al. 2022
Order Methanosarcinales_A
Family Methermicoccaceae Cheng et al. 2007
- Methermicoccus Cheng et al. 2007
Class Methanosarcinia
Order Methanotrichales
- Family Methanotrichaceae Akinyemi et al. 2021 "Candidatus Methanocrinis" Khomyakova et al. 2023 Methanothrix Huser, Wuhrmann & Zehnder 1983 [Methanosaeta Patel & Sprott 1990]
Order Methanosarcinales
| Phylogeny of Methanosarcinales |
| "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Argarchaeum" "Methanoperedentaceae" "Ca. Methanoperedens" "Methanogastraceae" "Ca. Methanogaster" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Argarchaeum" "Methanoperedentaceae" "Ca. Methanoperedens" "Methanogastraceae" "Ca. Methanogaster" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Argarchaeum" "Methanoperedentaceae" "Ca. Methanoperedens" |
| "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" |
| "Methanocomedentaceae" | "Ca. Methanocomedens" "Ca. Methanomarinus" |
| "Ca. Methanocomedens" |
|
| "Ca. Methanomarinus" |
|
|
|
| "Ethanoperedentaceae" "Ca. Argarchaeum" "Methanoperedentaceae" "Ca. Methanoperedens" |
| "Ethanoperedentaceae" | "Ca. Argarchaeum" |
|
| "Methanoperedentaceae" | "Ca. Methanoperedens" |
|
|
|
| "Methanogastraceae" "Ca. Methanogaster" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| "Methanogastraceae" | "Ca. Methanogaster" |
|
| Methanosarcinaceae | "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina |
| "Methanofrustulum" Methanolapillus Methanimicrococcus |
| "Methanofrustulum" |
|
| Methanolapillus Methanimicrococcus |
| Methanolapillus |
|
| Methanimicrococcus |
|
|
|
| Methanosarcina |
|
|
| Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| Methanohalobium Methanosalsum |
| Methanohalobium |
|
| Methanosalsum |
|
|
| Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus |
| Methanococcoides Methanohalophilus |
| Methanococcoides |
|
| Methanohalophilus |
|
|
| Methanomethylovorans Methanolobus |
| Methanomethylovorans |
|
| Methanolobus |
|
|
|
|
|
|
|
- Family "Methanocomedentaceae" corrig. Chadwick et al. 2022 "Candidatus Methanocomedens" Chadwick et al. 2022 (ANME-2a) "Candidatus Methanomarinus" Chadwick et al. 2022 (ANME-2b)
- Family "Ethanoperedentaceae" (sic) Pallen, Rodriguez-R & Alikhan 2022 (EX4572-44) "Ca. Argarchaeum" corrig. Chen et al. 2019 ["Candidatus Ethaniperedens" corrig. Hahn et al. 2020]
- Family "Methanoperedentaceae" corrig. Haroon et al. 2013 "Candidatus Methanoperedens" Haroon et al. 2013 (ANME-2d, AAA)
- Family "Methanogastraceae" corrig. Chadwick et al. 2022 (ANME-2c) "Candidatus Methanogaster" Chadwick et al. 2022
- Family Methanosarcinaceae Balch & Wolfe 1981 ?Halomethanococcus Yu & Kawamura 1988 Methanimicrococcus corrig. Sprenger et al. 2000 Methanococcoides Sowers & Ferry 1985 "Methanofrustulum" Protasov & Brune 2023 Methanohalobium Zhilina & Zavarzin 1988 Methanohalophilus Paterek & Smith 1988 Methanolapillus Protasov & Brune 2025 Methanolobus König & Stetter 1983 Methanomethylovorans Lomans et al. 2004 Methanosalsum Boone & Baker 2002 Methanosarcina Kluyver & van Niel 1936 ?"Candidatus Methanovorans" Chadwick et al. 2022 (ANME-3)
Class " Methanomicrobia "
Order Methanomicrobiales
- Family Methanofollaceae Chuvochina et al. 2024 Methanofollis Zellner et al. 1999
- Family Methanocorpusculaceae Zellner et al. 1989 Methanocalculus Ollivier et al. 1998 Methanocorpusculum Zellner et al. 1988 ["Methanorbis" Protasov & Brune 2023]
- Family Methanospirillaceae Boone et al. 2002 "Methanofilum" Protasov & Brune 2023 Methanolinea Imachi et al. 2008 Methanoregula Brauer et al. 2011 Methanosphaerula Cadillo-Quiroz et al. 2009 Methanospirillum Ferry, Smith & Wolfe 1974
- Family Methanoculleaceae Chuvochina et al. 2024 Methanoculleus Maestrojun et al. 1990
- Family Methanomicrobiaceae Balch & Wolfe 1981 Methanochimaera Zhou et al. 2024 Methanogenium Romesser et al. 1981 Methanolacinia Zellner et al. 1990 Methanomicrobium Balch & Wolfe 1981 [Methanoeremita Zhou et al. 2024] Methanoplanus Wildgruber et al. 1984 Methanovulcanius Chien et al. 2023
Class "Ordosarchaeia"
Order "Ordosarchaeales"
- Family "Ordosarchaeaceae" Zhao et al. 2024 ?"Candidatus Ordosarchaeum" Zhao et al. 2024
Class Halobacteria
Order Halobacteriales
| Phylogeny of Halobacteriales |
| "Afararchaeaceae" "Ca. Afararchaeum" Halorutilaceae Halorutilus "Hikarchaeaceae" "Ca. Chewarchaeum" "Ca. Hikarchaeum" "Salinarchaeaceae" Salinarchaeum Halalkalicoccaceae Natronorarus Halalkalicoccus Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| "Afararchaeaceae" "Ca. Afararchaeum" Halorutilaceae Halorutilus "Hikarchaeaceae" "Ca. Chewarchaeum" "Ca. Hikarchaeum" "Salinarchaeaceae" Salinarchaeum Halalkalicoccaceae Natronorarus Halalkalicoccus Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| "Afararchaeaceae" "Ca. Afararchaeum" Halorutilaceae Halorutilus |
| "Afararchaeaceae" | "Ca. Afararchaeum" |
|
| Halorutilaceae | Halorutilus |
|
|
| "Hikarchaeaceae" "Ca. Chewarchaeum" "Ca. Hikarchaeum" "Salinarchaeaceae" Salinarchaeum Halalkalicoccaceae Natronorarus Halalkalicoccus Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| "Hikarchaeaceae" "Ca. Chewarchaeum" "Ca. Hikarchaeum" |
| "Hikarchaeaceae" | "Ca. Chewarchaeum" "Ca. Hikarchaeum" |
| "Ca. Chewarchaeum" |
|
| "Ca. Hikarchaeum" |
|
|
|
| "Salinarchaeaceae" Salinarchaeum Halalkalicoccaceae Natronorarus Halalkalicoccus Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| "Salinarchaeaceae" Salinarchaeum |
| "Salinarchaeaceae" | Salinarchaeum |
|
|
| Halalkalicoccaceae Natronorarus Halalkalicoccus Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| Halalkalicoccaceae Natronorarus Halalkalicoccus |
| Halalkalicoccaceae | Natronorarus Halalkalicoccus |
| Natronorarus |
|
| Halalkalicoccus |
|
|
|
| Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium |
| Haladaptataceae | Halomicrococcus Haladaptatus Halorussus |
| Halomicrococcus Haladaptatus |
| Halomicrococcus |
|
| Haladaptatus |
|
|
| Halorussus |
|
|
| Halobacteriaceae | Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium |
| Halanaeroarchaeum Halodesulfurarchaeum |
| Halanaeroarchaeum |
|
| Halodesulfurarchaeum |
|
|
| Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium |
| Halospeciosus Salarchaeum Halocalculus Halarchaeum |
| Halospeciosus Salarchaeum |
| Halospeciosus |
|
| Salarchaeum |
|
|
| Halocalculus Halarchaeum |
| Halocalculus |
|
| Halarchaeum |
|
|
|
| Halobacterium |
|
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|
|
| Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd |
| Natronoarchaeaceae Halostella Natranaeroarchaeum Natronoarchaeum |
| Natronoarchaeaceae | Halostella Natranaeroarchaeum Natronoarchaeum |
| Halostella |
|
| Natranaeroarchaeum Natronoarchaeum |
| Natranaeroarchaeum |
|
| Natronoarchaeum |
|
|
|
|
| Natrialbaceae cont'd |
|
|
| Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd |
| Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula |
| Halococcaceae | Halococcus |
|
| Haloarculaceae | Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula |
| Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus |
| Halocatena Halomarina |
| Halocatena |
|
| Halomarina |
|
|
| Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus |
| Natronomonas |
|
| Halorarius Halosegnis Salinirubellus Haloglomus |
| Halorarius Halosegnis |
| Halorarius |
|
| Halosegnis |
|
|
| Salinirubellus Haloglomus |
| Salinirubellus |
|
| Haloglomus |
|
|
|
|
|
| Halorientalis Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula |
| Halorientalis |
|
| Halococcoides Halapricum Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula |
| Halococcoides Halapricum Halorhabdus |
| Halococcoides |
|
| Halapricum Halorhabdus |
| Halapricum |
|
| Halorhabdus |
|
|
|
| Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula |
| Salinirussus Salinibaculum Halovenus Halosimplex |
| Salinirussus Salinibaculum Halovenus |
| Salinirussus |
|
| Salinibaculum Halovenus |
| Salinibaculum |
|
| Halovenus |
|
|
|
| Halosimplex |
|
|
| Halomicrobium Haloarcula |
| Halomicrobium |
|
| Haloarcula |
|
|
|
|
|
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|
| Haloferacaceae cont'd |
|
|
|
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|
|
|
|
- ?"Haloalcalophilium" Lizama et al. 2000a
- Family "Abyssiniarchaeaceae" Gutiérrez-Preciado et al. 2024 ?"Candidatus Abyssiniarchaeum" Gutiérrez-Preciado et al. 2024 (DAL-8mg-83m_91C4R)
- Family "Karumarchaeaceae" Gutiérrez-Preciado et al. 2024 ?"Candidatus Karumarchaeum" Gutiérrez-Preciado et al. 2024 (DAL-8mg-83m_91C4R)
- Family "Afararchaeaceae" Baker et al. 2024 "Candidatus Afararchaeum" Baker et al. 2024
- Family Halorutilaceae Durán-Viseras et al. 2023 Halorutilus Durán-Viseras et al. 2023
- Family "Hikarchaeaceae" Martijn et al. 2020 (MG-IV) "Candidatus Chewarchaeum" Baker et al. 2024 "Candidatus Hikarchaeum" Martijn et al. 2020
- Family "Salinarchaeaceae" Salinarchaeum Cui et al. 2014
- Family Halalkalicoccaceae Mao et al. 2025 Halalkalicoccus Xue et al. 2005 Natronorarus Mao et al. 2025
- Family Haladaptataceae Rinke et al. 2020 Haladaptatus Savage et al. 2007 Halomicrococcus Chen et al. 2020 Halorussus Cui et al. 2014
- Family Halobacteriaceae Halanaeroarchaeum Sorokin et al. 2016 Halarchaeum Minegishi et al. 2010 Halobacterium (Elazari-Volcani 1940) Elazari-Volcani 1957 ["Halobacter" Anderson 1954] Halocalculus Minegishi et al. 2015 Halodesulfurarchaeum Sorokin et al. 2017 Halospeciosus Li et al. 2024 Salarchaeum Shimane et al. 2011
- Family Natronoarchaeaceae Sorokin et al. 2023 Halostella Song et al. 2016 Natranaeroarchaeum Sorokin et al. 2023 Natronoarchaeum Shimane et al. 2010
| Phylogeny of Natrialbaceae |
| Halorubellus Haloarchaeobius Halovivax Natronobeatus Natribaculum Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halorubellus Haloarchaeobius Halovivax Natronobeatus Natribaculum Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halorubellus Haloarchaeobius |
| Halorubellus |
|
| Haloarchaeobius |
|
|
| Halovivax Natronobeatus Natribaculum Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halovivax |
|
| Natronobeatus Natribaculum Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natronobeatus |
|
| Natribaculum Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natribaculum |
|
| Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natronobiforma Saliphagus Halomontanus Natrononativus Natronosalvus |
| Natronobiforma Saliphagus Halomontanus |
| Natronobiforma Saliphagus |
| Natronobiforma |
|
| Saliphagus |
|
|
| Halomontanus |
|
|
| Natrononativus Natronosalvus |
| Natrononativus |
|
| Natronosalvus |
|
|
|
| Salinilacihabitans Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Salinilacihabitans |
|
| Halovalidus Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halovalidus |
|
| Halostagnicola Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halostagnicola |
|
| Natronococcus Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natronococcus |
|
| Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Halopiger_A Natrarchaeobaculum Halopiger_B Halosolutus |
| Halopiger_A Natrarchaeobaculum |
| Halopiger_A |
|
| Natrarchaeobaculum |
|
|
| Halopiger_B Halosolutus |
| Halopiger_B |
|
| Halosolutus |
|
|
|
| Natrarchaeobius Salinadaptatus Natrialba Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natrarchaeobius Salinadaptatus Natrialba |
| Natrarchaeobius |
|
| Salinadaptatus Natrialba |
| Salinadaptatus |
|
| Natrialba |
|
|
|
| Natronolimnobius Halopiger Natronobacterium Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natronolimnobius Halopiger Natronobacterium |
| Natronolimnobius Halopiger |
| Natronolimnobius |
|
| Halopiger |
|
|
| Natronobacterium |
|
|
| Natronolimnohabitans Haloterrigena Natronorubrum Natrinema |
| Natronolimnohabitans Haloterrigena Natronorubrum |
| Natronolimnohabitans Haloterrigena |
| Natronolimnohabitans |
|
| Haloterrigena |
|
|
| Natronorubrum |
|
|
| Natrinema |
|
|
|
|
|
|
|
|
|
|
|
|
|
|
- Family Natrialbaceae Gupta et al. 2015 Haloarchaeobius Makhdoumi-Kakhki et al. 2012 Halomontanus Mao et al. 2024 [Natronoglomus Sorokin et al. 2025] Halopiger Gutierrez et al. 2007 Halorubellus Cui et al. 2014 Halosolutus Sun et al. 2022 Halostagnicola Castillo et al. 2006 Haloterrigena Ventosa et al. 1999 Halovalidus Dong et al. 2025 Halovivax Castillo et al. 2006 Natrarchaeobaculum Sorokin et al. 2020 Natrarchaeobius corrig. Sorokin et al. 2020 Natrialba Kamekura & Dyall-Smith 1996 Natribaculum Liu et al. 2015 [Halovarius Mehrshad et al. 2015] Natronobacterium Tindall et al. 1984 [Halobiforma Hezayen et al. 2002] Natronobiforma Sorokin et al. 2019 Natronococcus Tindall et al. 1984 Natronolimnobius Itoh et al. 2005 Natronolimnohabitans Sorokin et al. 2020 Natrinema McGenity et al. 1998 Natronobeatus Li et al. 2023 Natrononativus Li et al. 2023 Natronorubrum Xu et al. 1999 Natronosalvus Tan et al. 2023 Salinadaptatus Xue et al. 2020 Salinilacihabitans Li et al. 2023 Saliphagus Yin et al. 2017
- Family Halococcaceae Gupta et al. 2016 Halococcus Schoop 1935
- Family Haloarculaceae Halapricum Song et al. 2014 ["Halarchaeoglobus" Sorokin et al. 2021] Haloarcula Torreblanca et al. 1986 [Halomicroarcula Echigo et al. 2013] Halocatena Verma et al. 2020 [Actinarchaeum corrig. Tang et al. 2024] Halococcoides Sorokin et al. 2019 Haloglomus Duran-Viseras, Sanchez-Porro & Ventosa 2020 Halomarina Inoue et al. 2011 Halomicroarcula Echigo et al. 2013 Halomicrobium Oren et al. 2002 [Halosiccatus Mehrshad et al. 2016] Halorientalis Cui et al. 2011 Halorarius Sun et al. 2023 Halorhabdus Waino et al. 2000 Halosegnis Duran-Viseras et al. 2022 [Sala Song et al. 2023 non Ross 1937 non Walker 1867; Salella Deshmukh & Oren 2023] Halosimplex Vreeland et al. 2003 Halovenus Makhdoumi-Kakhki et al. 2012 Natronomonas Kamekura et al. 1997 Salinibaculum Han & Cui 2020 Salinirubellus Hou et al. 2018 Salinirussus Cui et al. 2017
| Phylogeny of Haloferacaceae |
| Salinirubrum Haloplanus Salinirarus Haloprofundus Halegenticoccus Halobium Halogranum Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halolamina "Ca. Halobonum" Halorarum Halobaculum Halalkalirubrum Natronocalculus Halohasta Halonotius Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Salinirubrum Haloplanus Salinirarus Haloprofundus Halegenticoccus Halobium Halogranum Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halolamina "Ca. Halobonum" Halorarum Halobaculum Halalkalirubrum Natronocalculus Halohasta Halonotius Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Salinirubrum |
|
| Haloplanus Salinirarus Haloprofundus Halegenticoccus Halobium Halogranum Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halolamina "Ca. Halobonum" Halorarum Halobaculum Halalkalirubrum Natronocalculus Halohasta Halonotius Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Haloplanus Salinirarus Haloprofundus Halegenticoccus Halobium Halogranum Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum |
| Haloplanus Salinirarus |
| Haloplanus |
|
| Salinirarus |
|
|
| Haloprofundus Halegenticoccus Halobium Halogranum Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum |
| Haloprofundus Halegenticoccus Halobium Halogranum |
| Haloprofundus |
|
| Halegenticoccus Halobium Halogranum |
| Halegenticoccus |
|
| Halobium Halogranum |
| Halobium |
|
| Halogranum |
|
|
|
|
| Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum |
| Salinigranum |
|
| Haloferax Halopelagius Halogeometricum Haloquadratum |
| Haloferax |
|
| Halopelagius Halogeometricum Haloquadratum |
| Halopelagius |
|
| Halogeometricum Haloquadratum |
| Halogeometricum |
|
| Haloquadratum |
|
|
|
|
|
|
|
| Halolamina "Ca. Halobonum" Halorarum Halobaculum Halalkalirubrum Natronocalculus Halohasta Halonotius Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Halolamina "Ca. Halobonum" Halorarum Halobaculum |
| Halolamina |
|
| "Ca. Halobonum" Halorarum Halobaculum |
| "Ca. Halobonum" |
|
| Halorarum Halobaculum |
| Halorarum |
|
| Halobaculum |
|
|
|
|
| Halalkalirubrum Natronocalculus Halohasta Halonotius Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Halalkalirubrum Natronocalculus Halohasta Halonotius |
| Halalkalirubrum Natronocalculus |
| Halalkalirubrum |
|
| Natronocalculus |
|
|
| Halohasta Halonotius |
| Halohasta |
|
| Halonotius |
|
|
|
| Halalkaliarchaeum Halopenitus Haloparvum Halorubrum |
| Halalkaliarchaeum |
|
| Halopenitus Haloparvum Halorubrum |
| Halopenitus |
|
| Haloparvum Halorubrum |
| Haloparvum |
|
| Halorubrum |
|
|
|
|
|
|
|
|
- Family Haloferacaceae Gupta et al. 2015 Halalkaliarchaeum Sorokin et al. 2019 Halalkalirubrum Zuo et al. 2021 ?"Candidatus Halectosymbiota" corrig. Filker et al. 2014 Halegenticoccus Liu et al. 2022 Halobaculum Oren et al. 1995 Halobium Mori et al. 2016 "Candidatus Halobonum" Ugalde et al. 2013 Haloferax Torreblanca et al. 1986 Halogeometricum Montalvo-Rodriguez et al. 1998 [Halosarcina Savage et al. 2008] Halogranum Cui et al. 2010 Halohasta Mou et al. 2013 Halolamina Cui et al. 2011 Halonotius Burns et al. 2010 ?"Halonovum" Hwang et al. 2025 Haloparvum Chen et al. 2016 Halopelagius Cui et al. 2010 Halopenitus Amoozegar et al. 2012 Haloplanus Elevi Bardavid et al. 2007 Haloprofundus Zhang et al. 2017 Haloquadratum Burns et al. 2007 [Halobellus Cui et al. 2011] Halorubrum McGenity & Grant 1996 [Halorubrobacterium Kamekura & Dyall-Smith 1996] ?Halosubterraneus Ding et al. 2026 ?Halotolerantifilum Fisher et al. 2026 Natronocalculus Sorokin et al. 2023 Salinigranum Cui & Zhang 2014 Salinirarus Zhang et al. 2025 Salinirubrum corrig. Cui & Qiu 2014
See also