List of Archaea genera

This article lists the genera of the Archaea and also shows taxa of archaea that do not contain genera. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI) unless otherwise noted.

Phylogeny

For the GTDB phylogeny, National Center for Biotechnology Information (NCBI) taxonomy was initially used to decorate the genome tree via tax2tree. The 16S rRNA-based Greengenes taxonomy is used to supplement the taxonomy particularly in regions of the tree with no cultured representatives. List of Prokaryotic names with Standing in Nomenclature (LPSN) is used as the primary taxonomic authority for establishing naming priorities. Taxonomic ranks are normalised using Phylorank and the taxonomy manually curated to remove polyphyletic groups.

Cladogram was taken from the GTDB release 10-RS226 (16 April 2025).

Archaea Nanobdellati "Altarchaeota" "Iainarchaeota" "Micrarchaeota" "Undinarchaeota" "Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota "Hadarchaeota" "Methanobacteriota B" "Hydrothermarchaeota" Methanobacteriota Thermoplasmatota Halobacteriota Korarchaeota" Thermoproteota Promethearchaeati Promethearchaeota	Thermoproteati Methanobacteriati
Archaea	Nanobdellati "Altarchaeota" "Iainarchaeota" "Micrarchaeota" "Undinarchaeota" "Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota "Hadarchaeota" "Methanobacteriota B" "Hydrothermarchaeota" Methanobacteriota Thermoplasmatota Halobacteriota Korarchaeota" Thermoproteota Promethearchaeati Promethearchaeota
	Nanobdellati "Altarchaeota" "Iainarchaeota" "Micrarchaeota" "Undinarchaeota" "Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota
Nanobdellati	"Altarchaeota" "Iainarchaeota" "Micrarchaeota" "Undinarchaeota" "Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota
	"Altarchaeota" "Iainarchaeota" "Micrarchaeota"
	"Altarchaeota"

	"Iainarchaeota" "Micrarchaeota"
	"Iainarchaeota"

	"Micrarchaeota"



	"Undinarchaeota" "Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota
	"Undinarchaeota"
	"Undinarchaeota"


	"Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota" Nanobdellota
	"Huberarchaeaota" "Aenigmarchaeota" "Nanohalarchaeota"
	"Huberarchaeaota"

	"Aenigmarchaeota" "Nanohalarchaeota"
	"Aenigmarchaeota"

	"Nanohalarchaeota"



	Nanobdellota





	"Hadarchaeota" "Methanobacteriota B" "Hydrothermarchaeota" Methanobacteriota Thermoplasmatota Halobacteriota Korarchaeota" Thermoproteota Promethearchaeati Promethearchaeota
	"Hadarchaeota" "Methanobacteriota B" "Hydrothermarchaeota" Methanobacteriota
	"Hadarchaeota" "Methanobacteriota B"
	"Hadarchaeota"

	"Methanobacteriota B"


	"Hydrothermarchaeota" Methanobacteriota
	"Hydrothermarchaeota"

	Methanobacteriota



	Thermoplasmatota Halobacteriota Korarchaeota" Thermoproteota Promethearchaeati Promethearchaeota
	Thermoplasmatota Halobacteriota
	Thermoplasmatota

	Halobacteriota


	Korarchaeota" Thermoproteota Promethearchaeati Promethearchaeota
	Korarchaeota"

	Thermoproteota Promethearchaeati Promethearchaeota
	Thermoproteota

Promethearchaeati	Promethearchaeota

Phylum Microcaldota

Class Microcaldia

Order Microcaldales

Family Microcaldaceae Sakai et al. 2023 Microcaldus Sakai et al. 2023

Phylum " Altarchaeota "

Class "Altarchaeia"

Order "Altarchaeales"

Family "Altarchaeaceae" corrig. Probst et al. 2014 "Candidatus Altarchaeum" corrig. Probst et al. 2014

Phylum " Iainarchaeota "

Class "Iainarchaeia"

Order "Forterreales"

Family "Forterreaceae" Pallen, Rodriguez-R & Alikhan 2022 [CSSED10-239] "Candidatus Forterrea" Probst & Banfield 2017

Order "Iainarchaeales"

Family "Iainarchaeaceae" Rinke et al. 2020 "Candidatus Iainarchaeum" Rinke et al. 2013

Phylum " Micrarchaeota "

Phylogeny of "Micrarchaeota"
"Norongarragalinales" "Ca. Norongarragalina" "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Gugararchaeales" "Ca. Gugararchaeum" "Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
	"Norongarragalinales" "Ca. Norongarragalina" "Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Gugararchaeales" "Ca. Gugararchaeum" "Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
	"Norongarragalinales" "Ca. Norongarragalina"
"Norongarragalinales"	"Ca. Norongarragalina"


	"Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum" "Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Gugararchaeales" "Ca. Gugararchaeum" "Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
	"Micrarchaeales" "Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum"
"Micrarchaeales"	"Micrarchaeaceae" "Ca. Mancarchaeum" "Ca. Micrarchaeum"
"Micrarchaeaceae"	"Ca. Mancarchaeum" "Ca. Micrarchaeum"
	"Ca. Mancarchaeum"

	"Ca. Micrarchaeum"




	"Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum" "Gugararchaeales" "Ca. Gugararchaeum" "Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
	"Anstonellales" "Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum"
"Anstonellales"	"Anstonellaceae" "Ca. Anstonella" "Bilamarchaeaceae" "Ca. Bilamarchaeum"
"Anstonellaceae"	"Ca. Anstonella"

"Bilamarchaeaceae"	"Ca. Bilamarchaeum"



	"Gugararchaeales" "Ca. Gugararchaeum" "Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
	"Gugararchaeales" "Ca. Gugararchaeum"
"Gugararchaeales"	"Ca. Gugararchaeum"


	"Burarchaeales" "Ca. Burarchaeum" "Fermentimicrarchaeales" "Ca. Fermentimicrarchaeum"
"Burarchaeales"	"Ca. Burarchaeum"

"Fermentimicrarchaeales"	"Ca. Fermentimicrarchaeum"

Class "Micrarchaeia"

Order "Norongarragalinales"

Family "Norongarragalinaceae" Vázquez-Campos et al. 2021 "Candidatus Norongarragalina" Vázquez-Campos et al. 2021

Order "Micrarchaeales"

Family "Micrarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Mancarchaeum" Golyshina et al. 2017 "Candidatus Micrarchaeum" Baker et al. 2010

Order "Anstonellales"

Family "Anstonellaceae" Vázquez-Campos et al. 2021 "Candidatus Anstonella" Vázquez-Campos et al. 2021
Family "Bilamarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Bilamarchaeum" Vázquez-Campos et al. 2021

Order "Gugararchaeales"

Family "Gugararchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Gugararchaeum" Vázquez-Campos et al. 2021

Order "Burarchaeales"

Family "Burarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Burarchaeum" Vázquez-Campos et al. 2021

Order "Fermentimicrarchaeales"

Family "Fermentimicrarchaeaceae" Kadnikov et al. 2020 (Sv326) "Candidatus Fermentimicrarchaeum" Kadnikov et al. 2020

Phylum "Undinarchaeota"

Class "Undinarchaeia"

Order "Undinarchaeales"

Family "Naiadarchaeaceae" Dombrowski et al. 2020 "Candidatus Naiadarchaeum" Dombrowski et al. 2020
Family "Undinarchaeaceae" Dombrowski et al. 2020 "Candidatus Undinarchaeum" Dombrowski et al. 2020

Phylum "Huberarchaeaota"

Class "Huberarchaeia"

Order "Huberarchaeales"

Family "Huberarchaeaceae" Rinke et al. 2020 "Candidatus Huberarchaeum" corrig. Schwank et al. 2019

Phylum " Aenigmatarchaeota "

Class "Aenigmatarchaeia"

Order "Aenigmatarchaeales"

Family "Aenigmatarchaeaceae" corrig. Rinke et al. 2020 "Candidatus Aenigmatarchaeum" corrig. Rinke et al. 2013

Phylum " Nanohalarchaeota "

Class "Nanohalarchaeia"

Order "Nanohalarchaeales"

Family "Nanohalarchaeaceae" ?"Candidatus Nanohalarchaeum" corrig. Hamm et al. 2019 ?"Candidatus Nanopetraeus" corrig. Crits‐Christoph et al. 2016

Order "Nanohydrothermales"

Family "Nanohydrothermaceae" Xie et al. 2022 ?"Candidatus Nanohydrothermus" Xie et al. 2022

Order "Nucleotidisoterales"

Family "Nanosalenecaceae" Xie et al. 2022 ?"Candidatus Nanosalenecus" Xie et al. 2022
Family "Nucleotidisoteraceae" Xie et al. 2022 ?"Candidatus Nucleotidisoter" Xie et al. 2022
Family "Nucleotidivindicaceae" Xie et al. 2022 ?"Nucleotidivindex" Xie et al. 2022

Class "Nanohalobiia"

Order "Nanohalobiales"

Phylogeny of "Nanohalobiales"
"Nanohalalkaliarchaeaceae" "Candidatus Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Candidatus Asbonarchaeum" "Candidatus Nanoanaerosalina" "Nanohalobiaceae" "Candidatus Nanosalinicola" "Candidatus Nanohalococcus" "Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Nanohalalkaliarchaeaceae" "Candidatus Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Candidatus Asbonarchaeum" "Candidatus Nanoanaerosalina" "Nanohalobiaceae" "Candidatus Nanosalinicola" "Candidatus Nanohalococcus" "Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Nanohalalkaliarchaeaceae" "Candidatus Nanohalalkaliarchaeum" "Nanoanaerosalinaceae" "Candidatus Asbonarchaeum" "Candidatus Nanoanaerosalina"
"Nanohalalkaliarchaeaceae"	"Candidatus Nanohalalkaliarchaeum"

"Nanoanaerosalinaceae"	"Candidatus Asbonarchaeum" "Candidatus Nanoanaerosalina"
	"Candidatus Asbonarchaeum"

	"Candidatus Nanoanaerosalina"



	"Nanohalobiaceae" "Candidatus Nanosalinicola" "Candidatus Nanohalococcus" "Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
"Nanohalobiaceae"	"Candidatus Nanosalinicola" "Candidatus Nanohalococcus" "Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Candidatus Nanosalinicola"

	"Candidatus Nanohalococcus" "Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Candidatus Nanohalococcus"

	"Candidatus Nanohalobium" "Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Candidatus Nanohalobium"

	"Candidatus Nanohalovita" "Candidatus Nanosalina"
	"Candidatus Nanohalovita"

	"Candidatus Nanosalina"

Family "Nanoanaerosalinaceae" Zhao et al. 2022 "Candidatus Asbonarchaeum" Baker et al. 2024 "Candidatus Nanoanaerosalina" Zhao et al. 2022
Family "Nanohalalkaliarchaeaceae" La Cono et al. 2020 "Candidatus Nanohalalkaliarchaeum" Ghai et al. 2011
Family "Nanohalobiaceae" La Cono et al. 2020 ?"Candidatus Haloredivivus" Ghai et al. 2011 "Candidatus Nanohalobium" La Cono et al. 2020 "Candidatus Nanohalococcus" Reva et al. 2023 "Candidatus Nanosalina" Narasingarao et al. 2012 "Candidatus Nanosalinicola" corrig. Narasingarao et al. 2012 "Candidatus Nanohalovita" Reva et al. 2023

Phylum Nanobdellota

Phylogeny of Nanobdellota
"Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella "Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Pacearchaeales" "Woesearchaeales" Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella
	"Pacearchaeales" "Woesearchaeales"
	"Pacearchaeales"

	"Woesearchaeales"


	Nanobdellales "Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella
Nanobdellales	"Nanoarchaeaceae" "Nanoarchaeum" Nanobdellaceae "Ca. Nanoclepta" Nanobdella
"Nanoarchaeaceae"	"Nanoarchaeum"

Nanobdellaceae	"Ca. Nanoclepta" Nanobdella
	"Ca. Nanoclepta"

	Nanobdella





	"Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum" "Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Tiddalikarchaeales" "Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum"
"Tiddalikarchaeales"	"Tiddalikarchaeaceae" "Ca. Tiddalikarchaeum"
"Tiddalikarchaeaceae"	"Ca. Tiddalikarchaeum"



	"Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum" "JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Jingweiarchaeales" "Jingweiarchaeaceae" "Ca. Jingweiarchaeum"
"Jingweiarchaeales"	"Jingweiarchaeaceae" "Ca. Jingweiarchaeum"
"Jingweiarchaeaceae"	"Ca. Jingweiarchaeum"



	"JAPDLS01" "Haiyanarchaeaceae" "Ca. Haiyanarchaeum" "Parvarchaeales" "Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
"JAPDLS01"	"Haiyanarchaeaceae" "Ca. Haiyanarchaeum"
"Haiyanarchaeaceae"	"Ca. Haiyanarchaeum"


"Parvarchaeales"	"Parvarchaeaceae" "Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
"Parvarchaeaceae"	"Ca. Rehaiarchaeum" "Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Ca. Rehaiarchaeum"

	"Ca. Acidifodinimicrobium" "Ca. Parvarchaeum"
	"Ca. Acidifodinimicrobium"

	"Ca. Parvarchaeum"

Class Nanobdellia

Order "Pacearchaeales"

Order "Woesearchaeales"

Order Nanobdellales

Family "Nanoarchaeaceae" Huber et al. 2011 "Nanoarchaeum" Huber et al. 2002
Family Nanobdellaceae Kato et al. 2022 ["Nanopusillaceae" Huber et al. 2011] Nanobdella Kato et al. 2022 ["Ca. Nanobsidianus" Castelle et al. 2015; "Ca. Nanopusillus" Wurch et al. 2016] "Candidatus Nanoclepta" St. John et al. 2019

Order "Tiddalikarchaeales"

Family "Tiddalikarchaeaceae" Vázquez-Campos et al. 2021 "Candidatus Tiddalikarchaeum" Vázquez-Campos et al. 2021

Order "Jingweiarchaeales"

Family "Jingweiarchaeaceae" Rao et al. 2023 "Candidatus Jingweiarchaeum" Rao et al. 2023

Order JAPDLS01

Family "Haiyanarchaeaceae" Rao et al. 2023 "Candidatus Haiyanarchaeum" Rao et al. 2023 ["Ca. Tinosarcha" Pallen, Rodriguez-R & Alikhan 2022]

Order "Parvarchaeales"

Family "Parvarchaeaceae" Rinke et al. 2020 "Candidatus Acidifodinimicrobium" Luo et al. 2020 "Candidatus Parvarchaeum" Baker et al. 2010 (ARMAN 4 & 5) "Candidatus Rehaiarchaeum" Rao et al. 2023

Phylum Promethearchaeota

Taxonomy of this group is based on LPSN, and also these works:

Phylogeny of "Asgardarchaeota"
"Asgardarchaeia" "Asgardarchaeales" "Atabeyarchaeia" "Atabeyarchaeales" "Sifarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales" "Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Asgardarchaeia" "Asgardarchaeales" "Atabeyarchaeia" "Atabeyarchaeales" "Sifarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales" "Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Asgardarchaeia" "Asgardarchaeales"
"Asgardarchaeia"	"Asgardarchaeales"


	"Atabeyarchaeia" "Atabeyarchaeales" "Sifarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales" "Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Atabeyarchaeia" "Atabeyarchaeales"
"Atabeyarchaeia"	"Atabeyarchaeales"


	"Sifarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales" "Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Sifarchaeia" "Borrarchaeales" "Sifarchaeales" "Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales"
	"Sifarchaeia" "Borrarchaeales" "Sifarchaeales"
"Sifarchaeia"	"Borrarchaeales" "Sifarchaeales"
	"Borrarchaeales"

	"Sifarchaeales"



	"Wukongarchaeia" "Wukongarchaeales" "Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales"
	"Wukongarchaeia" "Wukongarchaeales"
"Wukongarchaeia"	"Wukongarchaeales"


	"Njordarchaeia" "Njordarchaeales" "Heimdallarchaeia" "Hodarchaeales" "Heimdallarchaeales"
"Njordarchaeia"	"Njordarchaeales"

"Heimdallarchaeia"	"Hodarchaeales" "Heimdallarchaeales"
	"Hodarchaeales"

	"Heimdallarchaeales"





	"Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales" "Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Odinarchaeia" "Odinarchaeales" "Jordiarchaeia" "Jordiarchaeales"
"Odinarchaeia"	"Odinarchaeales"

"Jordiarchaeia"	"Jordiarchaeales"


	"Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales" "Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
	"Baldrarchaeia" "Baldrarchaeales" "Thorarchaeia" "Thorarchaeales"
"Baldrarchaeia"	"Baldrarchaeales"

"Thorarchaeia"	"Thorarchaeales"


	"Hermodarchaeia" "Hermodarchaeales" "Promethearchaeia" "Helarchaeales" "Promethearchaeales"
"Hermodarchaeia"	"Hermodarchaeales"

"Promethearchaeia"	"Helarchaeales" "Promethearchaeales"
	"Helarchaeales"

	"Promethearchaeales"

Promethearchaeota incertae sedis

?"Friggarchaeota" Caceres 2019
?"Gefionarchaeota" Caceres 2019
?"Idunnarchaeota" Caceres 2019
?"Sleipnirarchaeota" Kioukis et al. 2025
?"Candidatus Balderarchaeum" Xie et al. 2022
?"Candidatus Sukunaarchaeum" Harada et al. 2025
?"Candidatus Tyrarchaeum" Xie et al. 2022

Class "Asgardarchaeia"

Order "Asgardarchaeales"

Family "Asgardarchaeaceae" Eme et al. 2023 ex Tamarit et al. 2024 "Candidatus Asgardarchaeum" Eme et al. 2023 ex Tamarit et al. 2024

Class "Atabeyarchaeia"

Order "Atabeyarchaeales"

Family "Atabeyarchaeaceae" Valentin-Alvaradoet al. 2024 "Candidatus Atabeyarchaeum" Valentin-Alvarado et al. 2024

Class "Sifarchaeia"

Order "Sifarchaeales"

Family "Sifarchaeaceae" Sun et al. 2021 "Candidatus Sifarchaeum" corrig. Farag, Zhao & Biddle 2020

Order "Borrarchaeales"

Family "Borrarchaeaceae" Liu et al. 2021 "Candidatus Borrarchaeum" Liu et al. 2021

Class "Wukongarchaeia"

Order "Wukongarchaeales"

Family "Wukongarchaeaceae" Liu et al. 2021 "Candidatus Wukongarchaeum" Liu et al. 2021

Class "Njordarchaeia"

Order "Njordarchaeales"

Family "Panguiarchaeaceae" Qu et al. 2023 "Panguiarchaeum" Qu et al. 2023
Family "Njordarchaeaceae" Xie et al. 2022 "Candidatus Njordarchaeum" Xie et al. 2022

Class "Heimdallarchaeia"

Order " Wenzhongarchaeales "

Order "Hodarchaeales"

Family "Hodarchaeaceae" Liu et al. 2021 "Candidatus Hodarchaeum" Liu et al. 2021
Family "Margulisarchaeaceae" Imachi et al. 2025 ?"Candidatus Flexarchaeum" Imachi et al. 2025 ?"Candidatus Margulisarchaeum" Imachi et al. 2025

Order "Gerdarchaeales"

Order "Heimdallarchaeales"

Family "Kariarchaeaceae" Liu et al. 2021 "Candidatus Kariarchaeum" Liu et al. 2021
Family "Heimdallarchaeaceae" Wu et al. 2022 "Candidatus Heimdallarchaeum" Wu et al. 2022

Class "Odinarchaeia"

Order "Odinarchaeales"

Family "Odinarchaeaceae" Tamarit et al. 2022 "Candidatus Odinarchaeum" Tamarit et al. 2022

Order " Yangjianarchaeales "

Class "Jordarchaeia"

Order "Jordarchaeales"

Family "Freyrarchaeaceae" Xie et al. 2022 "Candidatus Freyrarchaeum" Xie et al. 2022
Family "Jordarchaeaceae" Sun et al. 2021 "Candidatus Jordarchaeum" Sun et al. 2021 ["Ca. Freyarchaeum" (sic) Valentin-Alvarado et al. 2024]

Class "Baldrarchaeia"

Order "Baldrarchaeales"

Family "Baldrarchaeaceae" corrig. Liu et al. 2021 "Candidatus Baldrarchaeum" Liu et al. 2021

Class " Thorarchaeia "

Order "Thorarchaeales"

Family "Thorarchaeaceae" (MBG-B)

Class "Hermodarchaeia"

Order "Hermodarchaeales"

Family "Hermodarchaeaceae" corrig. Liu et al. 2021 "Candidatus Hermodarchaeum" Liu et al. 2021

Class Promethearchaeia

Order "Helarchaeales"

Order Promethearchaeales

Phylogeny of "Promethearchaeales"
"Sigynarchaeaceae" "Ca. Sigynarchaeum" Promethearchaeaceae "Ca. Prometheoarchaeum" "Ca. Harpocratesius" "Ca. Lokiarchaeum"
	"Sigynarchaeaceae" "Ca. Sigynarchaeum" Promethearchaeaceae "Ca. Prometheoarchaeum" "Ca. Harpocratesius" "Ca. Lokiarchaeum"
"Sigynarchaeaceae"	"Ca. Sigynarchaeum"

Promethearchaeaceae	"Ca. Prometheoarchaeum" "Ca. Harpocratesius" "Ca. Lokiarchaeum"
	"Ca. Prometheoarchaeum"

	"Ca. Harpocratesius" "Ca. Lokiarchaeum"
	"Ca. Harpocratesius"

	"Ca. Lokiarchaeum"

Family Sigynarchaeaceae Xie et al. 2022 "Candidatus Sigynarchaeum" Xie et al. 2022
Family Promethearchaeaceae Imachi et al. 2024 "Candidatus Harpocratesius" Wu et al. 2022 "Candidatus Lokiarchaeum" corrig. Spang et al. 2015 (MBGB, DSAG) Promethearchaeum Imachi et al. 2024

Phylum " Korarchaeota "

Class "Korarchaeia"

Order "Korarchaeales"

Family "Korarchaeaceae" Rinke et al. 2020 "Candidatus Korarchaeum" Elkins et al. 2008 "Candidatus Methanodesulfokora" McKay et al. 2019

Phylum Thermoproteota

Phylogeny of Thermoproteota
"BAT" "Bathyarchaeia" "Bifangarchaeales" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" "Nitrososphaeria A" "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales "Sulfobacteria" "Methanomethylicia" "Nezhaarchaeales" "Culexarchaeles" "Methanomethylicales" "Thermoproteia" "Gearchaeales" "Thermofilales" Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales
	"BAT" "Bathyarchaeia" "Bifangarchaeales" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" "Nitrososphaeria A" "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales "Sulfobacteria" "Methanomethylicia" "Nezhaarchaeales" "Culexarchaeles" "Methanomethylicales" "Thermoproteia" "Gearchaeales" "Thermofilales" Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales
"BAT"	"Bathyarchaeia" "Bifangarchaeales" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales" "Nitrososphaeria A" "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales
	"Bathyarchaeia" "Bifangarchaeales" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales"
"Bathyarchaeia"	"Bifangarchaeales" "Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales"
	"Bifangarchaeales"

	"Xuanwuarculales" "Hecatellales" "Houtuarculales" "Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales"
	"Xuanwuarculales" "Hecatellales" "Houtuarculales"
	"Xuanwuarculales"

	"Hecatellales" "Houtuarculales"
	"Hecatellales"

	"Houtuarculales"



	"Wuzhiqiibiales" "Zhuquarculales" "Bathyarchaeales"
	"Wuzhiqiibiales"

	"Zhuquarculales" "Bathyarchaeales"
	"Zhuquarculales"

	"Bathyarchaeales"






	"Nitrososphaeria A" "Caldarchaeales" Nitrososphaeria "Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales
"Nitrososphaeria A"	"Caldarchaeales"

Nitrososphaeria	"Geothermarchaeales" PSMU01 Conexivisphaerales Nitrososphaerales
	"Geothermarchaeales"

	PSMU01 Conexivisphaerales Nitrososphaerales
	PSMU01

	Conexivisphaerales Nitrososphaerales
	Conexivisphaerales

	Nitrososphaerales






"Sulfobacteria"	"Methanomethylicia" "Nezhaarchaeales" "Culexarchaeles" "Methanomethylicales" "Thermoproteia" "Gearchaeales" "Thermofilales" Thermoproteales "Sulfolobia" "Tardisphaerales" Sulfolobales
	"Methanomethylicia" "Nezhaarchaeales" "Culexarchaeles" "Methanomethylicales" "Thermoproteia" "Gearchaeales" "Thermofilales" Thermoproteales
"Methanomethylicia"	"Nezhaarchaeales" "Culexarchaeles" "Methanomethylicales"
	"Nezhaarchaeales"

	"Culexarchaeles" "Methanomethylicales"
	"Culexarchaeles"

	"Methanomethylicales"



"Thermoproteia"	"Gearchaeales" "Thermofilales" Thermoproteales
	"Gearchaeales" "Thermofilales" Thermoproteales
	"Gearchaeales"

	"Thermofilales" Thermoproteales
	"Thermofilales"

	Thermoproteales





	"Sulfolobia" "Tardisphaerales" Sulfolobales
"Sulfolobia"	"Tardisphaerales" Sulfolobales
	"Tardisphaerales"

	Sulfolobales

Thermoproteota Incertae sedis

"Brockiarchaeota"

Class " Bathyarchaeia "

Order "Bifangarchaeales"

Family "Bifangicolaceae" Hou et al. 2023 ?"Candidatus Bifangicola" Hou et al. 2023 [3300025546_26]
Family "Bifangarchaeaceae" Hou et al. 2023 "Candidatus Bifangarchaeum" corrig. Hou et al. 2023 [B24: DTBU01 sp011374155]
Family "Bifangistellaceae" Hou et al. 2023 "Candidatus Bifangistella" Hou et al. 2023 [DRYS01: DRYS01 sp011367275]

Order "Xuanwuarculales"

Family "Mazuisomaceae" Hou et al. 2023 ?"Candidatus Mazuisoma" Hou et al. 2023
Family "Mazuicellaceae" Hou et al. 2023 ?"Candidatus Mazuibellus" Hou et al. 2023 ?"Candidatus Mazuicella" Hou et al. 2023
Family "Mazuousiaecae" Hou et al. 2023 ?"Candidatus Mazuarcula" Hou et al. 2023 ?"Candidatus Mazuousia" Hou et al. 2023
Family "Xuanwuarculaceae" Hou et al. 2023 "Candidatus Xuanwuarcula" Hou et al. 2023 [RBG-16-48-13 sp001775995]

Order "Hecatellales"

Phylogeny of "Hecatellales"
"Hecatellaceae" "Ca. Hecatella" "Ca. Jinwuarcula" "Ca. Jinwuicola" "Ca. Jinwuimorpha"
"Hecatellaceae"	"Ca. Hecatella" "Ca. Jinwuarcula" "Ca. Jinwuicola" "Ca. Jinwuimorpha"
	"Ca. Hecatella" "Ca. Jinwuarcula"
	"Ca. Hecatella"

	"Ca. Jinwuarcula"


	"Ca. Jinwuicola" "Ca. Jinwuimorpha"
	"Ca. Jinwuicola"

	"Ca. Jinwuimorpha"

Family "Hecatellaceae" Rinke et al. 2020 "Candidatus Hecatella" Adam et al. 2022 "Candidatus Jinwuarcula" Hou et al. 2023 [B25] "Candidatus Jinwuicola" Hou et al. 2023 [EX4484-218 sp002254975] ?"Candidatus Jinwuimonas" Hou et al. 2023 "Candidatus Jinwuimorpha" Hou et al. 2023 [EX4484-218 sp003661965] ?"Candidatus Jinwuiprofundus" Hou et al. 2023 ?"Candidatus Jinwuousia" Hou et al. 2023 [JdFR-11 sp002011035]

Order "Houtuarculales"

Phylogeny of "Houtuarculales"
"Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus" "Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Houtuarculaceae" "Ca. Houtuarcula"
	"Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus" "Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Houtuarculaceae" "Ca. Houtuarcula"
	"Crenohoutuithermaceae" "Ca. Crenohoutuithermus" "Fontihoutuithermaceae" "Ca. Fontihoutuithermus"
"Crenohoutuithermaceae"	"Ca. Crenohoutuithermus"

"Fontihoutuithermaceae"	"Ca. Fontihoutuithermus"


	"Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus" "Houtuousiaceae" "Ca. Houtuousia" "Houtuarculaceae" "Ca. Houtuarcula"
	"Aphrohoutuibellaceae" "Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus"
"Aphrohoutuibellaceae"	"Ca. Aphrohoutuibellus" "Ca. Hydrohoutuibellus"
	"Ca. Aphrohoutuibellus"

	"Ca. Hydrohoutuibellus"



	"Houtuousiaceae" "Ca. Houtuousia" "Houtuarculaceae" "Ca. Houtuarcula"
"Houtuousiaceae"	"Ca. Houtuousia"

"Houtuarculaceae"	"Ca. Houtuarcula"

Family "Houtuithermaceae" Hou et al. 2023 ?"Candidatus Houtuithermus" Hou et al. 2023
Family "Marihoutuimonadaceae" Hou et al. 2023 ?"Candidatus Houtuibellus" Hou et al. 2023 ?"Candidatus Marihoutuimonas" Hou et al. 2023
Family "Crenohoutuithermaceae" Hou et al. 2023 "Candidatus Crenohoutuithermus" Hou et al. 2023 [DTDX01]
Family "Fontihoutuithermaceae" Hou et al. 2023 ?"Candidatus Fluxihoutuithermus" Hou et al. 2023 [DTOY01 sp011358085] "Candidatus Fontihoutuithermus" Hou et al. 2023 [DTGE01 sp011389385]
Family "Aphrohoutuibellaceae" Hou et al. 2023 "Candidatus Aphrohoutuibellus" Hou et al. 2023 [RBG-13-38-9] "Candidatus Hydrohoutuibellus" Hou et al. 2023 [DTMT01 sp011355205]
Family "Houtuousiaceae" Hou et al. 2023 "Candidatus Houtuousia" Hou et al. 2023 [40CM-2-53-6]
Family "Houtuarculaceae" Hou et al. 2023 ?"Candidatus Limnohoutuiplasma" Hou et al. 2023 "Candidatus Houtuarcula" Hou et al. 2023 [FEN-987]

Order "Wuzhiqiibiales"

Phylogeny of "Wuzhiqiibiales"
"Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha" "Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium"
	"Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha" "Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium"
	"Wuzhiqiimorphaceae" "Ca. Wuzhiqiimorpha"
"Wuzhiqiimorphaceae"	"Ca. Wuzhiqiimorpha"


	"Wuzhiqiimonadaceae" "Ca. Wuzhiqiimonas" "Wuzhiqiibiaceae" "Ca. Wuzhiqiibium"
"Wuzhiqiimonadaceae"	"Ca. Wuzhiqiimonas"

"Wuzhiqiibiaceae"	"Ca. Wuzhiqiibium"

Family "Wuzhiqiithermaceae" Hou et al. 2023 ?"Candidatus Wuzhiqiithermus" Hou et al. 2023
Family "Wuzhiqiitorraceae" Hou et al. 2023 ?"Candidatus Wuzhiqiitorris" Hou et al. 2023
Family "Wuzhiqiimorphaceae" Hou et al. 2023 "Candidatus Wuzhiqiimorpha" Hou et al. 2023 [PIYN01: PIYN01 sp003601775]
Family "Wuzhiqiimonadaceae" Hou et al. 2023 "Candidatus Wuzhiqiimonas" Hou et al. 2023 [B23: B23]
Family "Wuzhiqiibiaceae" Hou et al. 2023 "Candidatus Wuzhiqiibium" Hou et al. 2023 [TCS64]

Order "Zhuquarculales"

Family "Zhuquarculaceae" Hou et al. 2023 "Candidatus Zhuquarcula" Hou et al. 2023 [EX4484-135]
Family "Zhuquomorphaceae" Hou et al. 2023 "Candidatus Zhuquomorpha" Hou et al. 2023

Order "Bathyarchaeales"

Phylogeny of "Bathyarchaeales"
"Pontizhurongicolaceae" "Ca. Pontizhurongicola" "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Pontizhurongicolaceae" "Ca. Pontizhurongicola" "Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Pontizhurongicolaceae" "Ca. Pontizhurongicola"
"Pontizhurongicolaceae"	"Ca. Pontizhurongicola"


	"Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella" "Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Hanbarchaeaceae" "Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella" "Marizhurongicellaceae" "Ca. Marizhurongibium" "Ca. Marizhurongicella"
"Hanbarchaeaceae"	"Ca. Hanbarchaeum" "Ca. Fufeiimorpha" "Ca. Zhurongicella"
	"Ca. Hanbarchaeum"

	"Ca. Fufeiimorpha" "Ca. Zhurongicella"
	"Ca. Fufeiimorpha"

	"Ca. Zhurongicella"



"Marizhurongicellaceae"	"Ca. Marizhurongibium" "Ca. Marizhurongicella"
	"Ca. Marizhurongibium"

	"Ca. Marizhurongicella"



	"Hanbopyraceae" "Ca. Hanbopyrus" "Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Hanbopyraceae" "Ca. Hanbopyrus"
"Hanbopyraceae"	"Ca. Hanbopyrus"


	"Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum" "Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Jingweiiplasmataceae" "Ca. Jingweiiplasma" "Bathyarchaeaceae" "Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum"
"Jingweiiplasmataceae"	"Ca. Jingweiiplasma"

"Bathyarchaeaceae"	"Ca. Baizosoma" "Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum"
	"Ca. Baizosoma"

	"Ca. Halozhurongicella" "Ca. Baizobellus" "Ca. Bathyarchaeum"
	"Ca. Halozhurongicella"

	"Ca. Baizobellus" "Ca. Bathyarchaeum"
	"Ca. Baizobellus"

	"Ca. Bathyarchaeum"





	"Bathycorpusculaceae" "Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
"Bathycorpusculaceae"	"Ca. Baizomorpha" "Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Baizomorpha"

	"Ca. Buyanhuyuitorris" "Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Buyanhuyuitorris"

	"Ca. Buyanhuyuicella" "Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Buyanhuyuicella"

	"Ca. Hanbocola" "Ca. Hanbothermus" "Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Hanbocola" "Ca. Hanbothermus"
	"Ca. Hanbocola"

	"Ca. Hanbothermus"


	"Ca. Yanziithermus" "Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Yanziithermus"

	"Ca. Yuguotorris" "Ca. Bathycorpusculum"
	"Ca. Yuguotorris"

	"Ca. Bathycorpusculum"

Family "Actozhurongousiaceae" Hou et al. 2023 ?"Candidatus Actozhurongousia" Hou et al. 2023
Family "Yuguousiaceae" Hou et al. 2023 ?"Candidatus Yuguousia" Hou et al. 2023
Family "Pontizhurongicolaceae" Hou et al. 2023 "Candidatus Pontizhurongicola" Hou et al. 2023 [B24-2 sp002490245]
Family "Hanbarchaeaceae" Hou et al. 2023 "Candidatus Fufeiimorpha" Hou et al. 2023 [PIYB01] "Candidatus Hanbarchaeum" Hou et al. 2023 [DTEX01] "Candidatus Zhurongicella" Hou et al. 2023 [B63]
Family "Marizhurongicellaceae" Hou et al. 2023 [B26-1] ?"Candidatus Aoguangisoma" Hou et al. 2023 "Candidatus Marizhurongibium" Hou et al. 2023 [B17-G17 sp003662645] "Candidatus Marizhurongicella" Hou et al. 2023 [B26-1] ?"Candidatus Yanziimonas" Hou et al. 2023 ?"Candidatus Zhurongibium" Hou et al. 2023
Family "Hanbopyraceae" Hou et al. 2023 "Candidatus Hanbopyrus" Hou et al. 2023 [WUQV01 sp009889585]
Family "Jingweiiplasmataceae" Hou et al. 2023 "Candidatus Jingweiiplasma" Hou et al. 2023 [SOJC01]
Family "Bathyarchaeaceae" Khomyakova et al. 2023 "Candidatus Baizobellus" Hou et al. 2023 [SOJZ01] ?"Candidatus Baizosediminiarchaeum" Yu et al. 2023 [BIN-L-1] "Candidatus Baizosoma" Hou et al. 2023 [BA1 sp001399805] ?"Candidatus Baizostella" Hou et al. 2023 "Candidatus Bathyarchaeum" Khomyakova et al. 2023 "Candidatus Halozhurongicella" Hou et al. 2023 [B1-G15]
Family "Bathycorpusculaceae" Loh & Brune 2023 [UBA233] ?"Candidatus Aoshunithermus" Hou et al. 2023 ?"Candidatus Baizomonas" Hou et al. 2023 "Candidatus Baizomorpha" Hou et al. 2023 [UBA233] "Candidatus Bathycorpusculum" Loh & Brune 2023 ["Ca. Hafrusarcha" Pallen, Rodriguez-R & Alikhan 2022] "Candidatus Buyanhuyuicella" Hou et al. 2023 [SOJA01] "Candidatus Buyanhuyuitorris" Hou et al. 2023 [PIYA01] "Candidatus Hanbocola" Hou et al. 2023 [DSZD01] "Candidatus Hanbothermus" Hou et al. 2023 [DRVP01] "Candidatus Yanziithermus" Hou et al. 2023 [JdFR-07] "Candidatus Yuguotorris" Hou et al. 2023 [DSLH01]

Class Nitrososphaeria_A

Order "Caldarchaeales"

Phylogeny of "Caldarchaeales"
"Calditenuaceae" "Ca. Calditenuis" "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum" "Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor"
	"Calditenuaceae" "Ca. Calditenuis" "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum" "Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor"
	"Calditenuaceae" "Ca. Calditenuis" "Caldarchaeaceae" "Ca. Caldarchaeum" "Ca. Pelearchaeum"
"Calditenuaceae"	"Ca. Calditenuis"

"Caldarchaeaceae"	"Ca. Caldarchaeum" "Ca. Pelearchaeum"
	"Ca. Caldarchaeum"

	"Ca. Pelearchaeum"



	"Wolframiiraptoraceae" "Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor"
"Wolframiiraptoraceae"	"Ca. Terraquivivens" "Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor"
	"Ca. Terraquivivens"

	"Ca. Geocrenenecus" "Ca. Benthortus" "Ca. Wolframiiraptor"
	"Ca. Geocrenenecus"

	"Ca. Benthortus" "Ca. Wolframiiraptor"
	"Ca. Benthortus"

	"Ca. Wolframiiraptor"

Family "Calditenuaceae" Balbay et al. 2023 "Candidatus Calditenuis" Beam et al. 2016
Family "Caldarchaeaceae" Rinke et al. 2020 "Candidatus Caldarchaeum" corrig. Nunoura et al. 2011 "Candidatus Pelearchaeum" Balbay et al. 2023 ["Ca. Cadrafarcha" Pallen, Rodriguez-R & Alikhan 2022]
Family "Wolframiiraptoraceae" Buessecker et al. 2022 "Candidatus Benthortus" Buessecker et al. 2022 "Candidatus Geocrenenecus" Buessecker et al. 2022 "Candidatus Terraquivivens" Buessecker et al. 2022 "Candidatus Wolframiiraptor" Buessecker et al. 2022

Class Nitrososphaeria

Nitrososphaeria incertae sedis

?"Candidatus Cenoporarchaeum" corrig. Zhang et al. 2019
?"Candidatus Gigantothauma" corrig. Muller et al. 2010
?"Candidatus Nitrosodeserticola" Hwang et al. 2021

Order "Geothermarchaeales"

Family "Geothermarchaeaceae" Adam et al. 2022 "Candidatus Geothermarchaeum" Adam et al. 2022 "Candidatus Scotarchaeum" Adam et al. 2022

Order PSMU01

Family PSMU01 "Candidatus Australarchaeum" Herbold et al. 2024

Order "Conexivisphaerales"

Family "Conexivisphaeraceae" Kato et al. 2021 (THSC Group) Conexivisphaera Kato et al. 2021

Order Nitrososphaerales

Phylogeny of Nitrososphaerales
"Gagatemarchaeaceae" "Ca. Gagatemarchaeum" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosocosmicus" Nitrososphaera "Ca. Nitrosopolaris" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Gagatemarchaeaceae" "Ca. Gagatemarchaeum" "Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosocosmicus" Nitrososphaera "Ca. Nitrosopolaris" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Gagatemarchaeaceae" "Ca. Gagatemarchaeum"
"Gagatemarchaeaceae"	"Ca. Gagatemarchaeum"


	"Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus" Nitrososphaeraceae "Ca. Nitrosocosmicus" Nitrososphaera "Ca. Nitrosopolaris" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Nitrosocaldaceae" "Ca. Nitrosothermus" "Ca. Nitrosocaldus"
"Nitrosocaldaceae"	"Ca. Nitrosothermus" "Ca. Nitrosocaldus"
	"Ca. Nitrosothermus"

	"Ca. Nitrosocaldus"



	Nitrososphaeraceae "Ca. Nitrosocosmicus" Nitrososphaera "Ca. Nitrosopolaris" Nitrosopumilaceae "Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
Nitrososphaeraceae	"Ca. Nitrosocosmicus" Nitrososphaera "Ca. Nitrosopolaris"
	"Ca. Nitrosocosmicus"

	Nitrososphaera "Ca. Nitrosopolaris"
	Nitrososphaera

	"Ca. Nitrosopolaris"



Nitrosopumilaceae	"Ca. Nitrosotalea" "Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Ca. Nitrosotalea"

	"Ca. Nitrosotenuis" "Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Ca. Nitrosotenuis"

	"Ca. Nitrosopelagicus" "Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Ca. Nitrosopelagicus"

	"Cenarchaeum" "Ca. Nitrosoabyssus" Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	"Cenarchaeum" "Ca. Nitrosoabyssus"
	"Cenarchaeum"

	"Ca. Nitrosoabyssus"


	Nitrosarchaeum "Ca. Nitrosomaritimum" Nitrosopumilus
	Nitrosarchaeum

	"Ca. Nitrosomaritimum" Nitrosopumilus
	"Ca. Nitrosomaritimum"

	Nitrosopumilus

Family "Methylarchaceae" Hua et al. 2019 ?"Candidatus Methylarchaeum" Hua et al. 2019 ?"Candidatus Methanotowutia" Ou et al. 2022
Family "Gagatemarchaeaceae" Sheridan et al. 2023 [UBA183] "Candidatus Gagatemarchaeum" Sheridan et al. 2023 [Bog-1369; GCA_003164815.1] ?"Candidatus Subgagatemarchaeum" Sheridan et al. 2023 [Fn1; IMG2558309099]
Family "Nitrosocaldaceae" Qin et al. 2016 "Candidatus Nitrosocaldus" de la Torre et al. 2008 "Candidatus Nitrosothermus" Luo et al. 2021
Family "Nitrosomiraceae" Zheng et al. 2024 "Candidatus Nitrosomirus" Zheng et al. 2024
Family Nitrososphaeraceae Stieglmeier et al. 2014 "Candidatus Nitrosocosmicus" corrig. Lehtovirta-Morley et al. 2016 "Candidatus Nitrosopolaris" Pessi, Rutanen & Hultman 2022 ["Ca. Flebogarcha" Pallen, Rodriguez-R & Alikhan 2022] Nitrososphaera Stieglmeier et al. 2014
Family Nitrosopumilaceae Qin et al. 2017 "Cenarchaeum" DeLong & Preston 1996 Nitrosarchaeum corrig. Jung et al. 2018 "Candidatus Nitrosoabyssus" Garritano et al. 2024 ?"Candidatus Nitrosokoinonia" Glasl et al. 2023 "Candidatus Nitrosomaritimum" Zhao et al. 2024 "Candidatus Nitrosopelagicus" Santoro et al. 2015 Nitrosopumilus Qin et al. 2017 ["Ca. Nitrosomarinus" Ahlgren et al. 2017] ?"Candidatus Nitrosospongia" Moeller et al. 2019 "Candidatus Nitrosotalea" Lehtovirta 2011 "Candidatus Nitrosotenuis" Lebedeva et al. 2013

Class "Methanomethylicia"

Order "Methanohydrogenicales" Family "Methanohydrogenicaceae" Berghuis et al. 2019 ?"Candidatus Methanohydrogenicus" Berghuis et al. 2019
Order "Methanomethylarchaeales" Family "Methanomethylarchaeceae" corrig. Hua et al. 2019 ?"Candidatus Methanomethylarchaeum" corrig. Hua et al. 2019
Order "Methanomethylovorales" Family "Methanomethylovoraceae" Hua et al. 2019 ?"Candidatus Methanomethylovorus" Hua et al. 2019

Order "Nezhaarchaeales"

Family "Methanohydrogenotrophicaceae" Hua et al. 2019 ?"Candidatus Methanogearchaeum" corrig. Hua et al. 2019 ?"Candidatus Methanohydrogenotrophicum" Hua et al. 2019

Order "Culexarchaeles"

Family "Culexarchaeaceae" Kohtz et al. 2022 ["Methanomediaceae" Berghuis et al. 2019] "Candidatus Culexarchaeum" Kohtz et al. 2022 ["Ca. Methanomedium" Berghuis et al. 2019]
Family "Culexmicrobiaceae" Kohtz et al. 2022 "Candidatus Culexmicrobium" Kohtz et al. 2022

Order "Methanosuratincolales"

Family "Candidatus Methanosuratincolaceae" Wu et al. 2025 ?"Candidatus Methanoluiimicrobium" Wu et al. 2024 "Candidatus Methanomethylicus" Vanwonterghem et al. 2016 ?"Candidatus Methanostamsia" Wu et al. 2024 "Candidatus Methanosuratinicola" corrig. Vanwonterghem et al. 2016

Class Thermoproteia

Order "Gearchaeales"

Family "Gearchaeaceae" corrig. ?Geoarchaeum (sic) Hunt et al. 2016

Order Thermofilales

Family Thermofilaceae Burggraf, Huber & Stetter 1997 Infirmifilum Zayulina et al. 2021 Thermofilum Zillig & Gierl 1983

Order Thermoproteales

Phylogeny of Thermoproteales
Thermocladiaceae Caldivirga Thermocladium Vulcanisaeta Thermoproteaceae Thermoproteus Pyrobaculum
	Thermocladiaceae Caldivirga Thermocladium Vulcanisaeta Thermoproteaceae Thermoproteus Pyrobaculum
Thermocladiaceae	Caldivirga Thermocladium Vulcanisaeta
	Caldivirga Thermocladium
	Caldivirga

	Thermocladium


	Vulcanisaeta


Thermoproteaceae	Thermoproteus Pyrobaculum
	Thermoproteus

	Pyrobaculum

Family Thermocladiaceae Chuvochina et al. 2024 Caldivirga Itoh et al. 1999 Thermocladium Itoh, Suzuki & Nakase 1998 Vulcanisaeta Itoh, Suzuki & Nakase 2002
Family Thermoproteaceae Zillig & Stetter 1982 Pyrobaculum Huber, Kristjansson & Stetter 1988 Thermoproteus Zillig & Stetter 1982

Class "Sulfolobia"

Order "Tardisphaerales"

Family "Marsarchaeaceae" (sic) (NAG2, NCGII)
Family "Tardisphaerace" Prokofeva et al. 2025 ?"Tardisphaera" Prokofeva et al. 2025

Order Sulfolobales

Phylogeny of Sulfolobales
Desulfurococcaceae Staphylothermus Thermogladius Thermosphaera Desulfurococcus Fervidicoccaceae Fervidicoccus Pyrodictiaceae Pyrolobus Hyperthermus Pyrofollis Pyrodictium Acidilobaceae "Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus Ignisphaeraceae Ignisphaera "Zestosphaeraceae" Zestosphaera Ignicoccaceae Ignicoccus Sulfolobaceae Sulfodiicoccus Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Desulfurococcaceae Staphylothermus Thermogladius Thermosphaera Desulfurococcus Fervidicoccaceae Fervidicoccus Pyrodictiaceae Pyrolobus Hyperthermus Pyrofollis Pyrodictium Acidilobaceae "Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus Ignisphaeraceae Ignisphaera "Zestosphaeraceae" Zestosphaera Ignicoccaceae Ignicoccus Sulfolobaceae Sulfodiicoccus Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Desulfurococcaceae Staphylothermus Thermogladius Thermosphaera Desulfurococcus Fervidicoccaceae Fervidicoccus Pyrodictiaceae Pyrolobus Hyperthermus Pyrofollis Pyrodictium Acidilobaceae "Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus
	Desulfurococcaceae Staphylothermus Thermogladius Thermosphaera Desulfurococcus
Desulfurococcaceae	Staphylothermus Thermogladius Thermosphaera Desulfurococcus
	Staphylothermus

	Thermogladius Thermosphaera Desulfurococcus
	Thermogladius

	Thermosphaera Desulfurococcus
	Thermosphaera

	Desulfurococcus





	Fervidicoccaceae Fervidicoccus Pyrodictiaceae Pyrolobus Hyperthermus Pyrofollis Pyrodictium Acidilobaceae "Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus
	Fervidicoccaceae Fervidicoccus
Fervidicoccaceae	Fervidicoccus


	Pyrodictiaceae Pyrolobus Hyperthermus Pyrofollis Pyrodictium Acidilobaceae "Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus
Pyrodictiaceae	Pyrolobus Hyperthermus Pyrofollis Pyrodictium
	Pyrolobus

	Hyperthermus Pyrofollis Pyrodictium
	Hyperthermus

	Pyrofollis Pyrodictium
	Pyrofollis

	Pyrodictium




Acidilobaceae	"Ca. Tiamatella" "Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus
	"Ca. Tiamatella"

	"Ca. Calypsonella" Thermodiscus Aeropyrum Caldisphaera Acidilobus
	"Ca. Calypsonella"

	Thermodiscus Aeropyrum Caldisphaera Acidilobus
	Thermodiscus Aeropyrum
	Thermodiscus

	Aeropyrum


	Caldisphaera Acidilobus
	Caldisphaera

	Acidilobus








	Ignisphaeraceae Ignisphaera "Zestosphaeraceae" Zestosphaera Ignicoccaceae Ignicoccus Sulfolobaceae Sulfodiicoccus Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Ignisphaeraceae Ignisphaera "Zestosphaeraceae" Zestosphaera
Ignisphaeraceae	Ignisphaera

"Zestosphaeraceae"	Zestosphaera


	Ignicoccaceae Ignicoccus Sulfolobaceae Sulfodiicoccus Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
Ignicoccaceae	Ignicoccus

Sulfolobaceae	Sulfodiicoccus Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Sulfodiicoccus

	Sulfurisphaera Stygiolobus Sulfolobus Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Sulfurisphaera Stygiolobus Sulfolobus
	Sulfurisphaera

	Stygiolobus Sulfolobus
	Stygiolobus

	Sulfolobus



	Sulfuracidifex Saccharolobus Metallosphaera "Ca. Aramenus" Acidianus
	Sulfuracidifex Saccharolobus
	Sulfuracidifex

	Saccharolobus


	Metallosphaera "Ca. Aramenus" Acidianus
	Metallosphaera

	"Ca. Aramenus" Acidianus
	"Ca. Aramenus"

	Acidianus

Family Desulfurococcaceae Zillig & Stetter 1983 Desulfurococcus Zillig & Stetter 1983 Staphylothermus Stetter & Fiala 1986 ?Sulfophobococcus Hensel et al. 1997 Thermogladius Osburn & Amend 2011 ex Kochetkova et al. 2016 Thermosphaera Huber et al. 1998 ?"Candidatus Zestomicrobium" Dzhuraeva, Bobodzhanova & Birkeland 2024
Family Fervidicoccaceae Perevalova et al. 2010 Fervidicoccus Perevalova et al. 2010
Family Pyrodictiaceae Burggraf, Huber & Stetter 1997 ?Geogemma Kashefi, Holmes & Lovley 2006 Hyperthermus Zillig, Holz & Wunderl 1991 Pyrodictium Stetter, König & Stackebrandt 1984 "Pyrofollis" Miyazaki et al. 2023 Pyrolobus Blochl et al. 1999
Family Acidilobaceae Prokofeva et al. 2009 Aeropyrum Sako et al. 1996 Acidilobus Prokofeva et al. 2000 Caldisphaera Itoh et al. 2003 ["Ca. Hestiella" St-John and Reysenbach 2024] "Candidatus Calypsonella" St John and Reysenbach 2024 ?Stetteria Jochimsen, Peinemann-Simon & Thomm 1998 Thermodiscus Stetter 2003 "Candidatus Tiamatella" St John and Reysenbach 2024
Family Ignisphaeraceae RChuvochina et al. 2024 Ignisphaera Niederberger et al. 2006
Family "Zestosphaeraceae" Pallen, Rodriguez-R & Alikhan 2022 Zestosphaera St. John et al. 2021
Family Ignicoccaceae Chuvochina et al. 2024 Ignicoccus Huber, Burggraf & Stetter 2000
Family Sulfolobaceae Stetter 1989 Acidianus Segerer et al. 1986 ["Sulfosphaerellus" Zhong et al. 1982; Desulfurolobus Zillig & Böck 1987] "Candidatus Aramenus" Servín-Garcidueñas & Martínez-Romero 2014 ?"Caldariella" de Rosa, Gambacorta & Bu'lock 1975 Metallosphaera Huber et al. 1989 Saccharolobus Sakai & Kurosawa 2018 Stygiolobus Segerer et al. 1991 Sulfodiicoccus Sakai & Kurosawa 2017 Sulfolobus Brock et al. 1972 Sulfuracidifex Itoh et al. 2020 Sulfurisphaera Kurosawa et al. 1998 ?Sulfurococcus Golovacheva, Vale’kho & Troitskii 1995

Phylum " Hadarchaeota "

Class "Persephonarchaeia"

Class "Hadarchaeia"

Order "Hadarchaeales"

Family "Cerberiarchaeaceae" Benito Merino et al. 2024 ?"Candidatus Cerberiarchaeum" Benito Merino et al. 2024 ?"Candidatus Melinoarchaeum" Yu et al. 2024
Family "Hadarchaeaceae" Chuvochina et al. 2019 ex Chuvochina et al. 2023 "Candidatus Hadarchaeum" Chuvochina et al. 2019 ?"Candidatus Hadesarchaeum" Hua et al. 2019 ?"Candidatus Methanourarchaeum" Hua et al. 2019

Phylum Methanobacteriota_B

Class "Theionarchaeia"

Class Thermococci

Phylogeny of Thermococci
Thermococci "Methanofastidiales" "Ca. Methanofastidiosum" "Methanofastidiosaceae" Thermococcales Palaeococcus "Thermococcus_A" "Thermococcus_B" Pyrococcus Thermococcus Thermococcaceae
	Thermococci "Methanofastidiales" "Ca. Methanofastidiosum" "Methanofastidiosaceae" Thermococcales Palaeococcus "Thermococcus_A" "Thermococcus_B" Pyrococcus Thermococcus Thermococcaceae
Thermococci	"Methanofastidiales" "Ca. Methanofastidiosum" "Methanofastidiosaceae" Thermococcales Palaeococcus "Thermococcus_A" "Thermococcus_B" Pyrococcus Thermococcus Thermococcaceae
"Methanofastidiales"	"Ca. Methanofastidiosum"
"Methanofastidiosaceae"
Thermococcales	Palaeococcus "Thermococcus_A" "Thermococcus_B" Pyrococcus Thermococcus
	Palaeococcus

	"Thermococcus_A" "Thermococcus_B" Pyrococcus Thermococcus
	"Thermococcus_A"

	"Thermococcus_B" Pyrococcus Thermococcus
	"Thermococcus_B"

	Pyrococcus Thermococcus
	Pyrococcus

	Thermococcus




Thermococcaceae

Order "Methanofastidiosales"

Family "Methanofastidiosaceae" Rinke et al. 2020 "Candidatus Methanofastidiosum" Nobu et al. 2016

Order Thermococcales

Family Thermococcaceae Zillig et al. 1988 Palaeococcus Takai et al. 2000 Pyrococcus Fiala & Stetter 1986 Thermococcus Zillig 1983 ["Caldococcus" Svetlichny et al. 1987]

Phylum "Hydrothermarchaeota"

Class "Hydrothermarchaeia"

Order "Hydrothermarchaeales"

Family BMS3B "Candidatus Pyrohabitans" Adam et al. 2022
Family "Hydrothermarchaeaceae" Chuvochina et al. 2019 "Candidatus Hydrothermarchaeum" Chuvochina et al. 2019

Phylum Methanobacteriota

Phylogeny of Methanobacteriota
"Methanococcia" Methanocaldococcaceae Methanocaldococcus Methanococcaceae Methanotorris Methanofervidicoccus Methanothermococcus Methanococcus Methanococcales "Methanopyria" Methanopyraceae Methanopyrus Methanopyrales "Methanobacteriia" Methanothermaceae Methanothermus "Methanothermobacteraceae_A" "Methanothermobacter_A" Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium Methanobacteriales
	"Methanococcia" Methanocaldococcaceae Methanocaldococcus Methanococcaceae Methanotorris Methanofervidicoccus Methanothermococcus Methanococcus Methanococcales "Methanopyria" Methanopyraceae Methanopyrus Methanopyrales "Methanobacteriia" Methanothermaceae Methanothermus "Methanothermobacteraceae_A" "Methanothermobacter_A" Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium Methanobacteriales
	"Methanococcia" Methanocaldococcaceae Methanocaldococcus Methanococcaceae Methanotorris Methanofervidicoccus Methanothermococcus Methanococcus Methanococcales
"Methanococcia"	Methanocaldococcaceae Methanocaldococcus Methanococcaceae Methanotorris Methanofervidicoccus Methanothermococcus Methanococcus
Methanocaldococcaceae	Methanocaldococcus

Methanococcaceae	Methanotorris Methanofervidicoccus Methanothermococcus Methanococcus
	Methanotorris

	Methanofervidicoccus Methanothermococcus Methanococcus
	Methanofervidicoccus

	Methanothermococcus Methanococcus
	Methanothermococcus

	Methanococcus




Methanococcales

	"Methanopyria" Methanopyraceae Methanopyrus Methanopyrales "Methanobacteriia" Methanothermaceae Methanothermus "Methanothermobacteraceae_A" "Methanothermobacter_A" Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium Methanobacteriales
"Methanopyria"	Methanopyraceae Methanopyrus
Methanopyraceae	Methanopyrus

Methanopyrales
"Methanobacteriia"	Methanothermaceae Methanothermus "Methanothermobacteraceae_A" "Methanothermobacter_A" Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium
	Methanothermaceae Methanothermus
Methanothermaceae	Methanothermus


	"Methanothermobacteraceae_A" "Methanothermobacter_A" Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium
	"Methanothermobacteraceae_A" "Methanothermobacter_A"
"Methanothermobacteraceae_A"	"Methanothermobacter_A"


	Methanothermobacteraceae Methanothermobacter Methanobacteriaceae "Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium
Methanothermobacteraceae	Methanothermobacter

Methanobacteriaceae	"Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter "Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium
	"Methanacia" "Methanoflexus" "Methanovirga" "Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter
	"Methanacia" "Methanoflexus" "Methanovirga"
	"Methanacia"

	"Methanoflexus" "Methanovirga"
	"Methanoflexus"

	"Methanovirga"



	"Methanorudis" "Methanobaculum" "Methanobinarius" "Methanocatella" "Methanarmilla" Methanobrevibacter
	"Methanorudis" "Methanobaculum" "Methanobinarius"
	"Methanorudis"

	"Methanobaculum" "Methanobinarius"
	"Methanobaculum"

	"Methanobinarius"



	"Methanocatella" "Methanarmilla" Methanobrevibacter
	"Methanocatella"

	"Methanarmilla" Methanobrevibacter
	"Methanarmilla"

	Methanobrevibacter





	"Methanobacterium_E" & "F" "Methanobacterium_D" Methanosphaera Methanobacterium
	"Methanobacterium_E" & "F"

	"Methanobacterium_D" Methanosphaera Methanobacterium
	"Methanobacterium_D"

	Methanosphaera Methanobacterium
	Methanosphaera

	Methanobacterium







Methanobacteriales

Class " Methanococcia "

Order Methanococcales

Family Methanocaldococcaceae Whitman 2002 Methanocaldococcus Whitman 2002
Family Methanococcaceae Balch & Wolfe 1981 Methanococcus Kluyver & van Niel 1936 Methanofervidicoccus Sakai et al. 2019 Methanothermococcus Whitman 2002 Methanotorris Whitman 2002 ["Methanoignis" Boone, Whitman & Rouviére 1993]

Class Methanopyria

Order Methanopyrales

Family Methanopyraceae Huber & Stetter 2002 Methanopyrus Kurr et al. 1992

Class Methanobacteriia

Order Methanobacteriales

Family Methanothermaceae Stetter et al. 1982 Methanothermus Stetter et al. 1982
Family Methanothermobacteraceae Chuvochina et al. 2024 Methanothermobacter Wasserfallen et al. 2000
Family Methanobacteriaceae Barker 1956 "Methanacia" Protasov & Brune 2023 "Methanarmilla" Protasov & Brune 2023 Methanobacterium Kluyver & van Niel 1936 "Methanobaculum" Protasov & Brune 2023 "Methanobinarius" Protasov & Brune 2023 Methanobrevibacter Balch & Wolfe 1981 "Methanocatella" Protasov & Brune 2023 "Methanoflexus" Protasov & Brune 2023 "Methanorudis" Protasov & Brune 2023 Methanosphaera Miller & Wolin 1985 "Methanovirga" Protasov & Brune 2023

Phylum Thermoplasmatota

Phylogeny of Thermoplasmatota
"Izemarchaea" "Thermoprofundales" "Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Izemarchaea" "Thermoprofundales" "Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Izemarchaea" "Thermoprofundales"
"Izemarchaea"	"Thermoprofundales"


	"Poseidoniia" MGIII "Poseidoniales" "Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Poseidoniia" MGIII "Poseidoniales"
"Poseidoniia"	MGIII "Poseidoniales"
	MGIII

	"Poseidoniales"



	"Penumbrarchaeia" "Penumbrarchaeales" "Thermoplasmatia" "Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
"Penumbrarchaeia"	"Penumbrarchaeales"

"Thermoplasmatia"	"Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales "Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Proteinoplasmatales" "Yaplasmatales" "Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales
	"Proteinoplasmatales" "Yaplasmatales"
	"Proteinoplasmatales"

	"Yaplasmatales"


	"Gimiplasmatales" "Sysuiplasmatales" Methanomassiliicoccales
	"Gimiplasmatales"

	"Sysuiplasmatales" Methanomassiliicoccales
	"Sysuiplasmatales"

	Methanomassiliicoccales




	"Halarchaeoplasmatales" "Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Halarchaeoplasmatales"

	"Lutacidiplasmatales" "Aciduliprofundales" "Thermoplasmatales"
	"Lutacidiplasmatales"

	"Aciduliprofundales" "Thermoplasmatales"
	"Aciduliprofundales"

	"Thermoplasmatales"

Class "Izemarchaea"

Order "Thermoprofundales"

(MBG-D, E2)

Class "Poseidoniia"

Order MGIII

Order "Poseidoniales"

Family "Poseidoniaceae" Rinke et al. 2019 "Candidatus Poseidonia" Rinke et al. 2019
Family "Thalassarchaeaceae" Rinke et al. 2019 "Candidatus Thalassarchaeum" corrig. Martin-Cuadrado et al. 2015

Class "Penumbrarchaeia"

Order "Penumbrarchaeales"

Family "Penumbrarchaeaceae" Maeke et al. 2025 "Candidatus Penumbrarchaeum" Maeke et al. 2025 [EX4484-36: EX4484-36: EX4484-6: GCA_965234725.1]

Class Thermoplasmata

Order "Angelarchaeales"

Order "Lunaplasmatales"

Family "Lunaplasmataceae" corrig. Zinke et al. 2021 ?"Candidatus Lunaplasma" (sic) Zinke et al. 2021

Order "Proteinoplasmatales"

(SG8-5)

Order "Yaplasmatales"

(RBG-16-68-12) [incl. "Detorarchales" Pallen & Alikhan 2021; "Limitiplasmatales" corrig. Zheng et al. 2022]

Order "Gimiplasmatales"

Family "Gimiplasmataceae" Hu et al. 2020 "Candidatus Gimiplasma" Hu et al. 2020 (UA10834)

Order "Sysuiplasmatales"

Family "Sysuiplasmataceae" Yuan et al. 2021 "Candidatus Sysuiplasma" Yuan et al. 2021

Order Methanomassiliicoccales

Phylogeny of Methanomassiliicoccales
Methanomassiliicoccaceae Methanomassiliicoccus Methanomethylophilaceae "Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma"
	Methanomassiliicoccaceae Methanomassiliicoccus Methanomethylophilaceae "Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma"
Methanomassiliicoccaceae	Methanomassiliicoccus

Methanomethylophilaceae	"Ca. Methanomicula" "Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma"
	"Ca. Methanomicula"

	"Methanarcanum" "Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma"
	"Methanarcanum"

	"Methanoprimaticola" Methanomethylophilus "Methanogranum" "Methanoplasma"
	"Methanoprimaticola"

	Methanomethylophilus "Methanogranum" "Methanoplasma"
	Methanomethylophilus

	"Methanogranum" "Methanoplasma"
	"Methanogranum"

	"Methanoplasma"

Family "Methanomixtatrophicaceae" Hua et al. 2019 ?"Candidatus Methanomixtatrophicum" Hua et al. 2019
Family Methanomassiliicoccaceae Iino et al. 2013 Methanomassiliicoccus Dridi et al. 2012
Family Methanomethylophilaceae Gaci et al. 2014 ex Borrel et al. 2024 "Methanarcanum" Chibani et al. 2023 "Methanogranum" Iino et al. 2023 Methanomethylophilus Borrel et al. 2024 "Candidatus Methanomicula" Protasov & Brune 2023 ["Ca. Cribefarcha" Pallen, Rodriguez-R & Alikhan 2022] "Methanoplasma" Lang & Brune 2023 "Methanoprimaticola" Chibani et al. 2023 ?"Candidatus Methanospyradousia" Gilroy et al. 2021

Order "Halarchaeoplasmatales"

Phylogeny of "Halarchaeoplasmatales"
"Candidatus Haladaptiplasma" "Candidatus Halarchaeoplasma" "Candidatus Natronoplasma" "Candidatus Saliniplasma"
	"Candidatus Haladaptiplasma" "Candidatus Halarchaeoplasma" "Candidatus Natronoplasma" "Candidatus Saliniplasma"
	"Candidatus Haladaptiplasma" "Candidatus Halarchaeoplasma"
	"Candidatus Haladaptiplasma"

	"Candidatus Halarchaeoplasma"


	"Candidatus Natronoplasma" "Candidatus Saliniplasma"
	"Candidatus Natronoplasma"

	"Candidatus Saliniplasma"

Family "Halarchaeoplasmataceae" Zhou et al. 2023 (PWKY01) "Candidatus Haladaptiplasma" corrig. Zhou et al. 2022 "Candidatus Halarchaeoplasma" Zhou et al. 2023 ["Ca. Haloplasma" Zhou et al. 2022 non Antunes et al. 2008] "Candidatus Natronoplasma" Zhou et al. 2022 "Candidatus Saliniplasma" Zhou et al. 2022

Order "Lutacidiplasmatales"

Family "Lutacidiplasmataceae" Sheridan et al. 2022 "Candidatus Lutacidiplasma" Sheridan et al. 2022 (UBA184)

Order "Aciduliprofundales"

Family "Aciduliprofundaceae" Rinke et al. 2020 "Aciduliprofundum" Reysenbach et al. 2006 (DHVE2 group)

Order Thermoplasmatales

Phylogeny of Thermoplasmatales
Thermoplasma Cuniculiplasma Thermogymnomonas Picrophilus Acidiplasma Ferroplasma
	Thermoplasma Cuniculiplasma Thermogymnomonas Picrophilus Acidiplasma Ferroplasma
	Thermoplasma

	Cuniculiplasma Thermogymnomonas Picrophilus Acidiplasma Ferroplasma
	Cuniculiplasma Thermogymnomonas
	Cuniculiplasma

	Thermogymnomonas


	Picrophilus Acidiplasma Ferroplasma
	Picrophilus

	Acidiplasma Ferroplasma
	Acidiplasma

	Ferroplasma

Family Thermoplasmataceae Reysenbach 2002 Acidiplasma Golyshina et al. 2009 ?"Candidatus Carboxiplasma" Bulaev et al. 2023 Cuniculiplasma corrig. Golyshina et al. 2016 Ferroplasma Golyshina et al. 2000 ?Oxyplasma Golyshina et al. 2024 Picrophilus Schleper et al. 1996 ?"Candidatus Scheffleriplasma" Krause et al. 2022 Thermogymnomonas Itoh, Yoshikawa & Takashina 2007 Thermoplasma Darland et al. 1970

Phylum Halobacteriota

Phylogeny of Halobacteriota
Methanonatronarchaeia Methanonatronarchaeales "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halorutilales Halobacteriales
	Methanonatronarchaeia Methanonatronarchaeales "Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halorutilales Halobacteriales
	Methanonatronarchaeia Methanonatronarchaeales
Methanonatronarchaeia	Methanonatronarchaeales


	"Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales "Methanomicrobia" Methanomicrobiales Halobacteria Halorutilales Halobacteriales
	"Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales "Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales
	"Methanoliparia" "Methanoliparales" "Archaeoglobia" "Mnemosynellales" Archaeoglobales
"Methanoliparia"	"Methanoliparales"

"Archaeoglobia"	"Mnemosynellales" Archaeoglobales
	"Mnemosynellales"

	Archaeoglobales



	"Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales" Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales
	"Syntropharchaeia" "Aerarchaeales" "Syntropharchaeales" "Alkanophagales"
"Syntropharchaeia"	"Aerarchaeales" "Syntropharchaeales" "Alkanophagales"
	"Aerarchaeales"

	"Syntropharchaeales" "Alkanophagales"
	"Syntropharchaeales"

	"Alkanophagales"




	Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales
	Methanocellia Methanocellales Methanosarcinia Methanotrichales "Methanosarcinales_A" Methanosarcinales
Methanocellia	Methanocellales

Methanosarcinia	Methanotrichales "Methanosarcinales_A" Methanosarcinales
	Methanotrichales

	"Methanosarcinales_A" Methanosarcinales
	"Methanosarcinales_A"

	Methanosarcinales







	"Methanomicrobia" Methanomicrobiales Halobacteria Halorutilales Halobacteriales
"Methanomicrobia"	Methanomicrobiales

Halobacteria	Halorutilales Halobacteriales
	Halorutilales

	Halobacteriales

Halobacteriota incertae sedis

?"Candidatus Methanoinsularis" Ou et al. 2022
?"Candidatus Methanoporticola" Ou et al. 2022
Order "Methanoflorentales" Family "Methanoflorentaceae" Mondav et al. 2014 ?"Candidatus Methanoflorens" Mondav et al. 2014

Class "Methanonatronarchaeia"

Order "Methanonatronarchaeales"

Family Methanonatronarchaeaceae Sorokin et al. 2018 "Candidatus Methanohalarchaeum" Sorokin et al. 2018 Methanonatronarchaeum Sorokin et al. 2018

Class "Methanoliparia"

Order "Methanoliparales"

Family "Methanoliparaceae" Borrel et al. 2019 "Candidatus Methanoliparum" Borrel et al. 2019 "Candidatus Methanolliviera" Borrel et al. 2019

Class " Archaeoglobia "

Phylogeny of "Archaeoglobia"
"Mnemosynellales" "Mnemosynellaceae" "Ca. Mnemosynella" Archaeoglobales Archaeoglobaceae "Archaeoglobus_A" & "C" Ferroglobus Geoglobus Archaeoglobus
	"Mnemosynellales" "Mnemosynellaceae" "Ca. Mnemosynella" Archaeoglobales Archaeoglobaceae "Archaeoglobus_A" & "C" Ferroglobus Geoglobus Archaeoglobus
"Mnemosynellales"	"Mnemosynellaceae" "Ca. Mnemosynella"
"Mnemosynellaceae"	"Ca. Mnemosynella"


Archaeoglobales	Archaeoglobaceae "Archaeoglobus_A" & "C" Ferroglobus Geoglobus Archaeoglobus
Archaeoglobaceae	"Archaeoglobus_A" & "C" Ferroglobus Geoglobus Archaeoglobus
	"Archaeoglobus_A" & "C"

	Ferroglobus Geoglobus Archaeoglobus
	Ferroglobus Geoglobus
	Ferroglobus

	Geoglobus


	Archaeoglobus

Order "Mnemosynellales"

Family "Mnemosynellaceae" Adam et al. 2022 "Candidatus Mnemosynella" Adam et al. 2022

Order Archaeoglobales

Family "Allopolytropaceae" corrig. Boyd et al. 2019 ?"Candidatus Allopolytropus" corrig. Boyd et al. 2019 ["Ca. Polytropus" Boyd et al. 2019 non Kirsch 1866]
Family Archaeoglobaceae Huber & Stetter 2002 Archaeoglobus Stetter 1988 Ferroglobus Hafenbradl et al. 1997 Geoglobus Kashefi et al. 2002 ?"Candidatus Methanoglobus" Buessecker et al. 2023 ?"Candidatus Methanomixotrophus" corrig. Liu et al. 2021 ?"Candidatus Methanoproducendum" Hua et al. 2019

Class "Syntropharchaeia"

Order "Aerarchaeales"

Family YL-1 "Candidatus Methanodefluvium" Leu et al. 2025
Family "Aerarchaeaceae" Leu et al. 2025 [CG-2] "Candidatus Aerarchaeum" Leu et al. 2025 (GG-2) "Candidatus Aerovita" Leu et al. 2025 (CM-1, CG-1, MW-1)

Order "Syntropharchaeales"

Family "Syntropharchaeaceae" (ANME-2 cluster) "Candidatus Syntropharchaeum" corrig. Laso-Perez et al. 2016

Order "Alkanophagales"

Family "Methanophagaceae" Chadwick et al. 2022 ?"Candidatus Methanoalium" Chadwick et al. 2022 ?"Candidatus Methanophaga" Chadwick et al. 2022
Family ?"Veteromethanophagaceae" Vulcano et al. 2022
Family "Alkanophagaceae" [B39-G2] "Candidatus Alkanophaga" Zehnle et al. 2023
Family "Methanospiraraceae" Laso-Pérez et al. 2022 "Candidatus Methanospirare" Laso-Pérez et al. 2022 "Candidatus Methanoxibalbensis" Laso-Pérez et al. 2022

Class Methanocellia

Order Methanocellales

Phylogeny of Methanocellales
Methanocellaceae Methanooceanicella "Methanocella_A" Methanocella
Methanocellaceae	Methanooceanicella "Methanocella_A" Methanocella
	Methanooceanicella "Methanocella_A"
	Methanooceanicella

	"Methanocella_A"


	Methanocella

Family Methanocellaceae Sakai et al. 2008 Methanocella Sakai et al. 2008 Methanooceanicella Zhang et al. 2024

Class Methanosarcinia

Order Methanotrichales

Family Methanotrichaceae Akinyemi et al. 2021 "Candidatus Methanocrinis" Khomyakova et al. 2023 Methanothrix Huser, Wuhrmann & Zehnder 1983 [Methanosaeta Patel & Sprott 1990]

Order Methanosarcinales_A

Family Methermicoccaceae Cheng et al. 2007

Methermicoccus Cheng et al. 2007

Order Methanosarcinales

Phylogeny of Methanosarcinales
"Methanogastraceae" "Ca. Methanogaster" "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Ethaniperedens" "Methanoperedentaceae" "Ca. Methanoperedens" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	"Methanogastraceae" "Ca. Methanogaster" "Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Ethaniperedens" "Methanoperedentaceae" "Ca. Methanoperedens" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	"Methanogastraceae" "Ca. Methanogaster"
"Methanogastraceae"	"Ca. Methanogaster"


	"Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Ethaniperedens" "Methanoperedentaceae" "Ca. Methanoperedens" Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	"Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus" "Ethanoperedentaceae" "Ca. Ethaniperedens" "Methanoperedentaceae" "Ca. Methanoperedens"
	"Methanocomedentaceae" "Ca. Methanocomedens" "Ca. Methanomarinus"
"Methanocomedentaceae"	"Ca. Methanocomedens" "Ca. Methanomarinus"
	"Ca. Methanocomedens"

	"Ca. Methanomarinus"



	"Ethanoperedentaceae" "Ca. Ethaniperedens" "Methanoperedentaceae" "Ca. Methanoperedens"
"Ethanoperedentaceae"	"Ca. Ethaniperedens"

"Methanoperedentaceae"	"Ca. Methanoperedens"



	Methanosarcinaceae "Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
Methanosarcinaceae	"Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	"Methanofrustulum" Methanolapillus Methanimicrococcus Methanosarcina
	"Methanofrustulum" Methanolapillus Methanimicrococcus
	"Methanofrustulum"

	Methanolapillus Methanimicrococcus
	Methanolapillus

	Methanimicrococcus



	Methanosarcina


	Methanohalobium Methanosalsum Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	Methanohalobium Methanosalsum
	Methanohalobium

	Methanosalsum


	Methanococcoides Methanohalophilus Methanomethylovorans Methanolobus
	Methanococcoides Methanohalophilus
	Methanococcoides

	Methanohalophilus


	Methanomethylovorans Methanolobus
	Methanomethylovorans

	Methanolobus

Family "Methanogastraceae" corrig. Chadwick et al. 2022 (ANME-2c) "Candidatus Methanogaster" Chadwick et al. 2022
Family "Methanocomedentaceae" corrig. Chadwick et al. 2022 "Candidatus Methanocomedens" Chadwick et al. 2022 (ANME-2a) "Candidatus Methanomarinus" Chadwick et al. 2022 (ANME-2b)
Family "Ethanoperedentaceae" (sic) Pallen, Rodriguez-R & Alikhan 2022 (EX4572-44) "Candidatus Ethaniperedens" corrig. Hahn et al. 2020 ["Ca. Argarchaeum" corrig. Chen et al. 2019]
Family "Methanoperedentaceae" corrig. Haroon et al. 2013 "Candidatus Methanoperedens" Haroon et al. 2013 (ANME-2d, AAA)
Family Methanosarcinaceae Balch & Wolfe 1981 ?Halomethanococcus Yu & Kawamura 1988 Methanimicrococcus corrig. Sprenger et al. 2000 Methanococcoides Sowers & Ferry 1985 "Methanofrustulum" Protasov & Brune 2023 Methanohalobium Zhilina & Zavarzin 1988 Methanohalophilus Paterek & Smith 1988 Methanolapillus Protasov & Brune 2025 Methanolobus König & Stetter 1983 Methanomethylovorans Lomans et al. 2004 Methanosalsum Boone & Baker 2002 Methanosarcina Kluyver & van Niel 1936 ?"Candidatus Methanovorans" Chadwick et al. 2022 (ANME-3)

Class " Methanomicrobia "

Order Methanomicrobiales

Phylogeny of Methanomicrobiales
Methanocorpusculaceae Methanocorpusculum Methanocalculus Methanofollaceae Methanofollis Methanospirillaceae Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum Methanoculleaceae Methanoculleus Methanomicrobiaceae Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
	Methanocorpusculaceae Methanocorpusculum Methanocalculus Methanofollaceae Methanofollis Methanospirillaceae Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum Methanoculleaceae Methanoculleus Methanomicrobiaceae Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
	Methanocorpusculaceae Methanocorpusculum Methanocalculus
Methanocorpusculaceae	Methanocorpusculum Methanocalculus
	Methanocorpusculum

	Methanocalculus



	Methanofollaceae Methanofollis Methanospirillaceae Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum Methanoculleaceae Methanoculleus Methanomicrobiaceae Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
	Methanofollaceae Methanofollis
Methanofollaceae	Methanofollis


	Methanospirillaceae Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum Methanoculleaceae Methanoculleus Methanomicrobiaceae Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
	Methanospirillaceae Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum
Methanospirillaceae	Methanosphaerula Methanolinea Methanoregula "Methanofilum" Methanospirillum
	Methanosphaerula

	Methanolinea Methanoregula "Methanofilum" Methanospirillum
	Methanolinea Methanoregula
	Methanolinea

	Methanoregula


	"Methanofilum" Methanospirillum
	"Methanofilum"

	Methanospirillum





	Methanoculleaceae Methanoculleus Methanomicrobiaceae Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
Methanoculleaceae	Methanoculleus

Methanomicrobiaceae	Methanovulcanius Methanogenium Methanoplanus Methanomicrobium Methanolacinia
	Methanovulcanius Methanogenium
	Methanovulcanius

	Methanogenium


	Methanoplanus Methanomicrobium Methanolacinia
	Methanoplanus

	Methanomicrobium Methanolacinia
	Methanomicrobium

	Methanolacinia

Family Methanocorpusculaceae Zellner et al. 1989 Methanocalculus Ollivier et al. 1998 Methanocorpusculum Zellner et al. 1988 ["Methanorbis" Protasov & Brune 2023]
Family Methanofollaceae Chuvochina et al. 2024 Methanofollis Zellner et al. 1999
Family Methanospirillaceae Boone et al. 2002 "Methanofilum" Protasov & Brune 2023 Methanolinea Imachi et al. 2008 Methanoregula Brauer et al. 2011 Methanosphaerula Cadillo-Quiroz et al. 2009 Methanospirillum Ferry, Smith & Wolfe 1974
Family Methanoculleaceae Chuvochina et al. 2024 Methanoculleus Maestrojun et al. 1990
Family Methanomicrobiaceae Balch & Wolfe 1981 ?Methanochimaera Zhou et al. 2024 Methanogenium Romesser et al. 1981 Methanolacinia Zellner et al. 1990 Methanomicrobium Balch & Wolfe 1981 [Methanoeremita Zhou et al. 2024] Methanoplanus Wildgruber et al. 1984 Methanovulcanius Chien et al. 2023

Class "Hikarchaeia"

Order "Hikarchaeales"

Family "Hikarchaeaceae" Martijn et al. 2020 (MG-IV) ?"Candidatus Hikarchaeum" Martijn et al. 2020

Class "Ordosarchaeia"

Order "Ordosarchaeales"

Family "Ordosarchaeaceae" Zhao et al. 2024 ?"Candidatus Ordosarchaeum" Zhao et al. 2024

Class Halobacteria

Order Halorutilales

Family Halorutilaceae Durán-Viseras et al. 2023 "Candidatus Afararchaeum" Baker et al. 2024 Halorutilus Durán-Viseras et al. 2023

Order Halobacteriales

Phylogeny of Halobacteriales
"Chewarchaeaceae" "Candidatus Chewarchaeum" "Halalkalicoccaceae" Halalkalicoccus "Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	"Chewarchaeaceae" "Candidatus Chewarchaeum" "Halalkalicoccaceae" Halalkalicoccus "Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	"Chewarchaeaceae" "Candidatus Chewarchaeum"
"Chewarchaeaceae"	"Candidatus Chewarchaeum"


	"Halalkalicoccaceae" Halalkalicoccus "Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	"Halalkalicoccaceae" Halalkalicoccus
"Halalkalicoccaceae"	Halalkalicoccus


	"Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	"Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum Natrialbaceae cont'd
	"Salinarchaeaceae" Salinarchaeum "Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum
	"Salinarchaeaceae" Salinarchaeum
"Salinarchaeaceae"	Salinarchaeum


	"Halostellaceae" Halostella Natronoarchaeaceae Natranaeroarchaeum Natronoarchaeum
"Halostellaceae"	Halostella

Natronoarchaeaceae	Natranaeroarchaeum Natronoarchaeum
	Natranaeroarchaeum

	Natronoarchaeum




	Natrialbaceae cont'd


	Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	Haladaptataceae Halomicrococcus Haladaptatus Halorussus Halobacteriaceae Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium
Haladaptataceae	Halomicrococcus Haladaptatus Halorussus
	Halomicrococcus Haladaptatus
	Halomicrococcus

	Haladaptatus


	Halorussus


Halobacteriaceae	Halanaeroarchaeum Halodesulfurarchaeum Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium
	Halanaeroarchaeum Halodesulfurarchaeum
	Halanaeroarchaeum

	Halodesulfurarchaeum


	Halospeciosus Salarchaeum Halocalculus Halarchaeum Halobacterium
	Halospeciosus Salarchaeum Halocalculus Halarchaeum
	Halospeciosus Salarchaeum
	Halospeciosus

	Salarchaeum


	Halocalculus Halarchaeum
	Halocalculus

	Halarchaeum



	Halobacterium




	Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula Haloferacaceae cont'd
	Halococcaceae Halococcus Haloarculaceae Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula
Halococcaceae	Halococcus

Haloarculaceae	Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula
	Halocatena Halomarina Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus
	Halocatena Halomarina
	Halocatena

	Halomarina


	Natronomonas Halorarius Halosegnis Salinirubellus Haloglomus
	Natronomonas

	Halorarius Halosegnis Salinirubellus Haloglomus
	Halorarius Halosegnis
	Halorarius

	Halosegnis


	Salinirubellus Haloglomus
	Salinirubellus

	Haloglomus





	Halorientalis Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula
	Halorientalis

	Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex Halomicrobium Haloarcula
	Halapricum Halococcoides Halorhabdus Salinirussus Salinibaculum Halovenus Halosimplex
	Halapricum Halococcoides Halorhabdus
	Halapricum

	Halococcoides Halorhabdus
	Halococcoides

	Halorhabdus



	Salinirussus Salinibaculum Halovenus Halosimplex
	Salinirussus Salinibaculum Halovenus
	Salinirussus

	Salinibaculum Halovenus
	Salinibaculum

	Halovenus



	Halosimplex



	Halomicrobium Haloarcula
	Halomicrobium

	Haloarcula






	Haloferacaceae cont'd

?"Haloalcalophilium" Lizama et al. 2000a
Family "Chewarchaeaceae" Baker et al. 2024 "Candidatus Chewarchaeum" Baker et al. 2024
Family "Halalkalicoccaceae" Mao et al. 2025 Halalkalicoccus Xue et al. 2005 ?Natronorarus Mao et al. 2025
Family "Salinarchaeaceae" Salinarchaeum Cui et al. 2014
Family "Halostellaceae" Pallen, Rodriguez-R & Alikhan 2022 Halostella Song et al. 2016
Family Natronoarchaeaceae Sorokin et al. 2023 Natranaeroarchaeum Sorokin et al. 2023 Natronoarchaeum Shimane et al. 2010

Phylogeny of Natrialbaceae
Halorubellus Haloarchaeobius Halovivax Salinilacihabitans Natronobeatus Natribaculum Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halorubellus Haloarchaeobius Halovivax Salinilacihabitans Natronobeatus Natribaculum Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halorubellus Haloarchaeobius
	Halorubellus

	Haloarchaeobius


	Halovivax Salinilacihabitans Natronobeatus Natribaculum Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halovivax

	Salinilacihabitans Natronobeatus Natribaculum Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Salinilacihabitans Natronobeatus Natribaculum
	Salinilacihabitans

	Natronobeatus Natribaculum
	Natronobeatus

	Natribaculum



	Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Saliphagus Natronobiforma Halomontanus Natrononativus Natronosalvus
	Saliphagus Natronobiforma Halomontanus
	Saliphagus

	Natronobiforma Halomontanus
	Natronobiforma

	Halomontanus



	Natrononativus Natronosalvus
	Natrononativus

	Natronosalvus



	Halostagnicola Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halostagnicola

	Halosolutus Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halosolutus

	Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halopiger goleimassiliensis Natrarchaeobaculum Salinadaptatus Natrarchaeobius
	Halopiger goleimassiliensis Natrarchaeobaculum
	Halopiger goleimassiliensis

	Natrarchaeobaculum


	Salinadaptatus Natrarchaeobius
	Salinadaptatus

	Natrarchaeobius



	Natronococcus Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Natronococcus

	Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma Natronolimnobius Natrialba
	Halopiger Halopiger djelfimassiliensis Natronobacterium Halobiforma
	Halopiger

	Halopiger djelfimassiliensis Natronobacterium Halobiforma
	Halopiger djelfimassiliensis

	Natronobacterium Halobiforma
	Natronobacterium

	Halobiforma




	Natronolimnobius Natrialba
	Natronolimnobius

	Natrialba



	Natronolimnohabitans Haloterrigena Natronorubrum Natrinema
	Natronolimnohabitans Haloterrigena Natronorubrum
	Natronolimnohabitans Haloterrigena
	Natronolimnohabitans

	Haloterrigena


	Natronorubrum


	Natrinema

Family Natrialbaceae Gupta et al. 2015 Haloarchaeobius Makhdoumi-Kakhki et al. 2012 Halobiforma Hezayen et al. 2002 Halomontanus Mao et al. 2024 [Natronoglomus Sorokin et al. 2025] Halopiger Gutierrez et al. 2007 Halorubellus Cui et al. 2014 Halosolutus Sun et al. 2022 Halostagnicola Castillo et al. 2006 Haloterrigena Ventosa et al. 1999 ?Halovalidus Dong et al. 2025 Halovivax Castillo et al. 2006 Natrarchaeobaculum Sorokin et al. 2020 Natrarchaeobius corrig. Sorokin et al. 2020 Natrialba Kamekura & Dyall-Smith 1996 Natribaculum Liu et al. 2015 [Halovarius Mehrshad et al. 2015] Natronobacterium Tindall et al. 1984 Natronobiforma Sorokin et al. 2019 Natronococcus Tindall et al. 1984 Natronolimnobius Itoh et al. 2005 Natronolimnohabitans Sorokin et al. 2020 Natrinema McGenity et al. 1998 Natronobeatus Li et al. 2023 Natrononativus Li et al. 2023 Natronorubrum Xu et al. 1999 Natronosalvus Tan et al. 2023 Salinadaptatus Xue et al. 2020 Salinilacihabitans Li et al. 2023 Saliphagus Yin et al. 2017
Family Haladaptataceae Rinke et al. 2020 Haladaptatus Savage et al. 2007 Halomicrococcus Chen et al. 2020 Halorussus Cui et al. 2014
Family Halobacteriaceae Halanaeroarchaeum Sorokin et al. 2016 Halarchaeum Minegishi et al. 2010 Halobacterium (Elazari-Volcani 1940) Elazari-Volcani 1957 ["Halobacter" Anderson 1954] Halocalculus Minegishi et al. 2015 Halodesulfurarchaeum Sorokin et al. 2017 Halospeciosus Li et al. 2024 Salarchaeum Shimane et al. 2011
Family Halococcaceae Gupta et al. 2016 Halococcus Schoop 1935
Family Haloarculaceae Halapricum Song et al. 2014 ["Halarchaeoglobus" Sorokin et al. 2021] Haloarcula Torreblanca et al. 1986 [Halomicroarcula Echigo et al. 2013] Halocatena Verma et al. 2020 [Actinarchaeum corrig. Tang et al. 2024] Halococcoides Sorokin et al. 2019 Haloglomus Duran-Viseras, Sanchez-Porro & Ventosa 2020 Halomarina Inoue et al. 2011 Halomicroarcula Echigo et al. 2013 Halomicrobium Oren et al. 2002 [Halosiccatus Mehrshad et al. 2016] Halorientalis Cui et al. 2011 Halorarius Sun et al. 2023 Halorhabdus Waino et al. 2000 Halosegnis Duran-Viseras et al. 2022 [Sala Song et al. 2023 non Ross 1937 non Walker 1867; Salella Deshmukh & Oren 2023] Halosimplex Vreeland et al. 2003 Halovenus Makhdoumi-Kakhki et al. 2012 Natronomonas Kamekura et al. 1997 Salinibaculum Han & Cui 2020 Salinirubellus Hou et al. 2018 Salinirussus Cui et al. 2017

Phylogeny of Haloferacaceae
Salinirubrum Haloplanus Halobium Halegenticoccus Halogranum Haloprofundus Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus Halalkalirubrum Natronocalculus Halohasta Halonotius Halolamina Halorarum Halobaculum Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Salinirubrum Haloplanus Halobium Halegenticoccus Halogranum Haloprofundus Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus Halalkalirubrum Natronocalculus Halohasta Halonotius Halolamina Halorarum Halobaculum Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Salinirubrum

	Haloplanus Halobium Halegenticoccus Halogranum Haloprofundus Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus Halalkalirubrum Natronocalculus Halohasta Halonotius Halolamina Halorarum Halobaculum Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Haloplanus Halobium Halegenticoccus Halogranum Haloprofundus Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus
	Haloplanus

	Halobium Halegenticoccus Halogranum Haloprofundus Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus
	Halobium Halegenticoccus Halogranum Haloprofundus
	Halobium

	Halegenticoccus Halogranum Haloprofundus
	Halegenticoccus

	Halogranum Haloprofundus
	Halogranum

	Haloprofundus




	Salinigranum Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus
	Salinigranum

	Haloferax Halopelagius Halogeometricum Haloquadratum Halobellus
	Haloferax

	Halopelagius Halogeometricum Haloquadratum Halobellus
	Halopelagius

	Halogeometricum Haloquadratum Halobellus
	Halogeometricum

	Haloquadratum Halobellus
	Haloquadratum

	Halobellus








	Halalkalirubrum Natronocalculus Halohasta Halonotius Halolamina Halorarum Halobaculum Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Halalkalirubrum Natronocalculus Halohasta Halonotius
	Halalkalirubrum Natronocalculus
	Halalkalirubrum

	Natronocalculus


	Halohasta Halonotius
	Halohasta

	Halonotius



	Halolamina Halorarum Halobaculum Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Halolamina Halorarum Halobaculum
	Halolamina

	Halorarum Halobaculum
	Halorarum

	Halobaculum



	Halalkaliarchaeum Halopenitus Haloparvum Halorubrum
	Halalkaliarchaeum

	Halopenitus Haloparvum Halorubrum
	Halopenitus

	Haloparvum Halorubrum
	Haloparvum

	Halorubrum

Family Haloferacaceae Gupta et al. 2015 Halalkaliarchaeum Sorokin et al. 2019 Halalkalirubrum Zuo et al. 2021 ?"Candidatus Halectosymbiota" corrig. Filker et al. 2014 Halegenticoccus Liu et al. 2022 Halobaculum Oren et al. 1995 ["Candidatus Halobonum" Ugalde et al. 2013] Halobellus Cui et al. 2011 Halobium Mori et al. 2016 Haloferax Torreblanca et al. 1986 Halogeometricum Montalvo-Rodriguez et al. 1998 [Halosarcina Savage et al. 2008] Halogranum Cui et al. 2010 Halohasta Mou et al. 2013 Halolamina Cui et al. 2011 Halonotius Burns et al. 2010 Haloparvum Chen et al. 2016 Halopelagius Cui et al. 2010 Halopenitus Amoozegar et al. 2012 Haloplanus Elevi Bardavid et al. 2007 Haloprofundus Zhang et al. 2017 Haloquadratum Burns et al. 2007 Halorubrum McGenity & Grant 1996 [Halorubrobacterium Kamekura & Dyall-Smith 1996] Natronocalculus Sorokin et al. 2023 ?Salinirarus Zhang et al. 2025 Salinirubrum corrig. Cui & Qiu 2014 Salinigranum Cui & Zhang 2014