The Scotokaryotes is a proposed basal eukaryote clade sister to Diaphoretickes. Likely Scotokaryote groupings include the Metamonads, the Malawimonas and the Podiata. The name comes from the Greek word scotos (dark) due to the complete absence of photosynthetic organisms in this clade, contrasting with Diaphoretickes and Excavata.

An alternative to the UnikontBikont division was suggested by Derelle et al. in 2015, where they proposed the acronyms Opimoda–Diphoda respectively, as substitutes to the older terms (Opimoda includes the old 'unikonts', plus some former bikonts; Diphoda includes most of the old 'bikonts', but not all). The name Opimoda is formed from the letters (shown in capitals) of OPIsthokonta and aMOebozoa.

Sarcomastigota Cavalier-Smith, 1983 is a proposed subkingdom (currently shown to be paraphyletic) that includes all members of Opimoda that are not animals or fungi. Sulcozoa Cavalier-Smith, 2012 is a proposed phylum (currently shown to be paraphyletic) within Sarcomastigota that does not include the phyla Amoebozoa (clade) and Choanozoa (paraphyletic), i.e. it includes the proposed subphyla Apusozoa and Varisulca.

Phylogeny

A proposed cladogram is

EukaryotaMetamonada Diphoda Discoba Diaphoretickes Opimoda Ancyromonadida Malawimonadida Podiata CRuMs Amorphea Amoebozoa Obazoa Breviatea Opisthokonta Apusomonadida
Metamonada
Diphoda Discoba Diaphoretickes Opimoda Ancyromonadida Malawimonadida Podiata CRuMs Amorphea Amoebozoa Obazoa Breviatea Opisthokonta Apusomonadida
DiphodaDiscoba Diaphoretickes
Discoba
Diaphoretickes
OpimodaAncyromonadida Malawimonadida Podiata CRuMs Amorphea Amoebozoa Obazoa Breviatea Opisthokonta Apusomonadida
Ancyromonadida Malawimonadida
Ancyromonadida
Malawimonadida
PodiataCRuMs Amorphea Amoebozoa Obazoa Breviatea Opisthokonta Apusomonadida
CRuMs
AmorpheaAmoebozoa Obazoa Breviatea Opisthokonta Apusomonadida
Amoebozoa
ObazoaBreviatea Opisthokonta Apusomonadida
Breviatea
Opisthokonta Apusomonadida
Opisthokonta
Apusomonadida

See also