Philotheca
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Philotheca is a genus of about fifty species of flowering plants in the family Rutaceae. Plants in this genus are shrubs with simple leaves arranged alternately along the stems, flowers that usually have five sepals, five petals and ten stamens that curve inwards over the ovary. All species are endemic to Australia and there are species in every state, but not the Northern Territory.
Description
Plants in the genus Philotheca are shrubs that are either glabrous or have tiny, simple hairs. The leaves are arranged alternately along the stems, narrow oblong to almost cylindrical and sessile or on a very short petiole. From a single to many flowers are arranged in leaf axils or on the ends of the branchlets. The flowers have five sepals and five petals (except in P. virgata which has four). The sepals are free from each other and the petals usually overlap at their bases. There are ten stamens that curve inwards over the ovary with anthers that have an appendage called the "apiculum". The ovary contains five carpels fused near their bases. The seeds are 2–5 mm (0.079–0.197 in) long and are released explosively from their capsule.
Taxonomy and naming
The genus Philotheca was first formally described in 1816 by Edward Rudge from a specimen collected near Port Jackson and the description was published in Transactions of the Linnean Society of London. The first species Rudge described was P. australis but this name is considered a nomen illegitimum and a taxonomic synonym of Philotheca salsolifolia by the Australian Plant Census.
The name Philotheca should have been written Psilotheca after the Ancient Greek words psilos meaning "bare", "smooth", "bald" or "naked" and theke meaning "case", "container", "envelope" or "sheath", referring to "the smooth tube of the stamens".
Many plants formerly in Eriostemon are now in this genus.
Distribution
Plants in the genus Philotheca are found in every state of Australia, but not in the Northern Territory.
Species
The following is a list of Philotheca species accepted by the Australian Plant Census as at April 2019:
- Philotheca acrolophaPaul G.Wilson (Qld.)
- Philotheca angustifolia(Paul G. Wilson) Paul G. Wilson – narrow-leaf wax flower Philotheca angustifolia (Paul G. Wilson) Paul G. Wilson subsp. angustifolia (Vic., S.A.) Philotheca angustifolia subsp. montana(Paul G. Wilson) Paul G. Wilson (Vic.)
- Philotheca apiculata(Paul G. Wilson) Paul G. Wilson (W.A.)
- Philotheca basistylaMollemans (W.A.)
- Philotheca brevifolia(A.Cunn. ex Endl.) Paul G. Wilson (N.S.W.)
- Philotheca brucei(F.Muell.) Paul G.Wilson (W.A.) Philotheca brucei subsp. brevifolia(Paul G. Wilson) Paul G. Wilson Philotheca brucei (F.Muell.) Paul G. Wilson subsp. brucei Philotheca brucei subsp. cinerea(Paul G. Wilson) Paul G. Wilson
- Philotheca buxifolia(Sm.) Paul G.Wilson (N.S.W.) Philotheca buxifolia (Sm.) Paul G. Wilson subsp. buxifolia – box-leaf wax flower Philotheca buxifolia subsp. falcataPaul G. Wilson Philotheca buxifolia subsp. obovata(G.Don) Paul G. Wilson
- Philotheca ciliataHook. (Qld., N.S.W.)
- Philotheca citrinaPaul G.Wilson (W.A.)
- Philotheca coateanaPaul G. Wilson (W.A.)
- Philotheca coccinea(C.A.Gardner) Paul G.Wilson (W.A.)
- Philotheca conduplicata(Paul G. Wilson) P.I.Forst. (Qld., N.S.W.)
- Philotheca cuticularisPaul G. Wilson (Qld.)
- Philotheca cymbiformis(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca deserti(E. Pritzel) Paul G.Wilson (W.A.) Philotheca deserti subsp. brevifolia Paul G.Wilson Philotheca deserti (E.Pritz.) Paul G.Wilson subsp. deserti
- Philotheca difformis(A.Cunn. ex Endl.) Paul G.Wilson Philotheca difformis (A.Cunn. ex Endl.) Paul G. Wilson subsp. difformis – small-leaf wax-flower (Qld., N.S.W., Vic., S.A.) Philotheca difformis subsp. smithiana(Benth.) Paul G. Wilson (Qld., N.S.W.)
- Philotheca epilosa(Paul G. Wilson) P.I.Forst. (Qld., N.S.W.)
- Philotheca eremicolaPaul G.Wilson (W.A.)
- Philotheca ericifolia(A.Cunn. ex Benth.) Paul G.Wilson (N.S.W.)
- Philotheca falcata(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca fitzgeraldii(C.R.P.Andrews) Paul G.Wilson (W.A.)
- Philotheca freycianaRozefelds (Tas.)
- Philotheca gardneri(Paul G.Wilson) Paul G.Wilson (W.A.) Philotheca gardneri (A.Cunn. ex Endl.) Paul G. Wilson subsp. gardneri Philotheca gardneri subsp. globosaPaul G. Wilson
- Philotheca glabra(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca glasshousiensis(Domin) P.I.Forst. (Qld.)
- Philotheca hispidula(Sieber ex Spreng.) Paul G.Wilson (N.S.W.)
- Philotheca kalbarriensisPaul G.Wilson (W.A.)
- Philotheca langeiMollemans (W.A.)
- Philotheca linearis(A.Cunn. ex Endl.) Paul G.Wilson (W.A., S.A., N.S.W.)
- Philotheca myoporoides(DC.) M.J.Bayly Philotheca myoporoides subsp. acuta(Blakely) Bayly (N.S.W.) Philotheca myoporoides subsp. brevipedunculataBayly (N.S.W.) Philotheca myoporoides subsp. euroensisBayly (Vic.) Philotheca myoporoides (DC.) Bayly subsp. myoporoides (N.S.W., Vic.) Philotheca myoporoides subsp. petraea Rozefelds (Vic.)
- Philotheca nodiflora(Lindl.) Paul G.Wilson (W.A.) Philotheca nodiflora subsp. calycina(Turcz.) Paul G.Wilson Philotheca nodiflora subsp. lasiocalyx(Domin) Paul G.Wilson Philotheca nodiflora subsp. latericolaPaul G.Wilson Philotheca nodiflora (Lindl.) Paul G.Wilson subsp. nodiflora
- Philotheca nutans(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca obovalis(A.Cunn.) Paul G.Wilson (N.S.W.)
- Philotheca obovatifolia(Bayly) P.I.Forst. (Qld., N.S.W.)
- Philotheca pachyphylla(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca papillataI.Telford & L.M.Copel. (N.S.W.)
- Philotheca pinoides(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca pungens(Lindl.) Paul G.Wilson — prickly waxflower (Vic., S.A.)
- Philotheca queenslandica(C.T.White) P.I.Forst. (Qld.)
- Philotheca reichenbachiiSieber ex Spreng. (N.S.W.)
- Philotheca rhomboidea(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca salsolifolia(Sm.) Druce (N.S.W., A.C.T.) Philotheca salsolifolia subsp. pedicellataPaul G.Wilson Philotheca salsolifolia (Sm.) Druce subsp. salsolifolia
- Philotheca scabra(Paxton) Paul G.Wilson (N.S.W.) Philotheca scabra subsp. latifolia(Paul G.Wilson) Paul G.Wilson Philotheca scabra (Paxton) Paul G.Wilsonsubsp. scabra
- Philotheca sericea(Paul G.Wilson) Paul G.Wilson (W.A.)
- Philotheca spicata(A.Rich.) Paul G.Wilson (W.A.)
- Philotheca sporadica(M.J.Bayly) Paul G.Wilson (Qld.)
- Philotheca thryptomenoides(S.Moore) Paul G.Wilson (W.A.)
- Philotheca tomentella(Diels) Paul G.Wilson (W.A.)
- Philotheca trachyphylla(F.Muell.) Paul G.Wilson (N.S.W., Vic.)
- Philotheca tubifloraA.S.George (W.A.)
- Philotheca verrucosa(A.Rich.) Paul G.Wilson (Vic., S.A., Tas.)
- Philotheca virgata(Hook.f.) Paul G.Wilson (N.S.W., Vic., Tas.)
- Philotheca wonganensis(Paul G.Wilson) Paul G.Wilson (W.A.)
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