The Qiupa Formation (Chinese: 秋扒组; pinyin: Qiūpázǔ) is a Late Cretaceous Maastrichtian geologic formation in Henan Province, central China. It is rich in dinosaur eggs and bones, such as those of carnivorous and herbivorous dinosaurs. The Qiupa Formation is considered to be Late Maastrichtian in age, about 72 million and 66 million years ago.

Geology

The sedimentation is characterized for the presence of calcareous mudstone interbedded with thin fine conglomerates, brownish red thick-bedded siltstone and parallel and cross laminations. This sedimentation indicates habitats composed by large and shallow meanders with braided river deltas.

Fossil content

The Qiupa Formation has yielded numerous dinosaur fossils, particularly eggs. The remains of various theropods such as troodontids, dromaeosaurids and oviraptorosaurs have been found. In addition, indeterminate remains of sauropods, ornithopods, ankylosaurs, lizards, turtles and a possible avian bone are reported. Therizinosaurids Protoceratopsids, and medium sized sauropods are also known from the Qiupa Formation

Dinosaurs

Dinosaurs of the Qiupa Formation
GenusSpeciesMaterialNotesImages
Ankylosauridae indet.Indeterminate"A tooth, well-preserved dorsal vertebra, incomplete dorsal rib and ischium."An ankylosaurid
Elongatoolithidae sp.Indeterminate"145 egg shells probably laid by oviraptorids."Oviraptorid eggs
LuanchuanraptorL. henanensis"Teeth, a frontal and fragmented postcrania."A dromaeosaurid
QiupalongQ. henanensisPartial skeleton preserving the pelvis and the right hindlimb.An ornithomimid
QiupanykusQ. zhangiPartial skeleton with vertebrae and hindlimbs.An alvarezsaurid
"Tyrannosaurus"T. luanchuanensisFive teeth.A tyrannosaurid
YulongY. miniMany juvenile specimens and one subadult skeleton.An oviraptorid
YuornisY. junchangiPartial skeleton.An enantiornithine bird

Mammals

Mammals of the Qiupa Formation
GenusSpeciesMaterialNotesImages
LotheridiumL. mengiNearly complete skull with jaws.A deltatheridiid
YubaatarY. zhongyuanensis"Virtually complete crania and fragmented poscrania."A cimolodont

Squamates

Squamates of the Qiupa Formation
GenusSpeciesMaterialNotesImages
FuniusaurusF. luanchuanensisPartial skull.A polyglyphanodont
TianyusaurusT. zhengiSkull and shoulder girdle.A polyglyphanodont
ZhongyuanxiZ. jiaiPartial skull.A stem-varanid anguimorph
Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon