Ruixinia is an extinct genus of somphospondylan sauropod dinosaur from the Early Cretaceous (Barremian) Yixian Formation of China. The genus contains a single species, Ruixinia zhangi, known from a partial articulated skeleton. This specimen preserves the most complete series of caudal vertebrae known from any Asian titanosauriform.

Discovery and naming

The Ruixinia holotype specimen, ELDM EL-J009, was found in sediments of the Yixian Formation in Beipiao, Liaoning Province, northeastern China. This locality is dated to the Barremian age of the Early Cretaceous period. The fossil material consists of a partial articulated skeleton including cervical (neck), dorsal (back), and sacral vertebrae, caudal (tail) vertebrae and associated chevrons, dorsal ribs, and a left ilium, pubis, femur, tibia, astragalus, metatarsal V, and possible pedal phalanx. At the time of its description, the fossil material was still partially embedded in matrix, with only the left side of the bones prepared.

In 2022, Ruixinia zhangi was described as a new genus and species of titanosaurian dinosaurs by Jinyou Mo, Feimin Ma, Yilun Yu, and Xing Xu. The binomial honors Ruixin Zhang, a contributor to the Erlianhaote Dinosaur Museum.

Description

Ruixinia was a mid-sized sauropod, with an estimated length of approximately 12 meters.

The neck of Ruixinia was over 4 meters long and consisted of at least 15 cervical vertebrae. Such a high number of cervical vertebrae is similar to Dongbeititan, Euhelopus, and Mamenchisaurus.

The tail of Ruixinia contained at least 52 vertebrae. The last few vertebrae were fused together, an unusual trait in sauropods otherwise only known to occur in some Jurassic Asian sauropods such as Shunosaurus and Mamenchisaurus. However, the structure of the fused vertebrae differs between the three taxa. In Ruixinia, the fused vertebrae form a rodlike structure.

The chevrons are distinctive, particularly that of the 20th caudal vertebra, which has a pentagonal shape.

Classification

Mo et al. (2022) recovered Ruixinia to be a basal titanosaur, placed as the sister taxon to a clade containing Daxiatitan and Xianshanosaurus. However, both Daxiatitan and Xianshanosaurus have also been suggested to have affinities with mamenchisaurids, and Mo et al. noted several similarities between Ruixinia and Mamenchisaurus. Ruixinia was not found to be closely related to either of its contemporaries, Dongbeititan or Liaoningotitan. The cladogram below displays the results of the phylogenetic analyses of Mo et al. (2022).

TitanosauriformesBrachiosauridae Somphospondyli Huanghetitan spp. Ligabuesaurus Dongbeititan Savannasaurus Chubutisaurus Angolatitan Europatitan Tastavinsaurus Liubangosaurus Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Brachiosauridae
SomphospondyliHuanghetitan spp. Ligabuesaurus Dongbeititan Savannasaurus Chubutisaurus Angolatitan Europatitan Tastavinsaurus Liubangosaurus Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Huanghetitan spp. Ligabuesaurus Dongbeititan Savannasaurus Chubutisaurus Angolatitan Europatitan Tastavinsaurus Liubangosaurus Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Huanghetitan spp.
Ligabuesaurus Dongbeititan Savannasaurus Chubutisaurus Angolatitan
Ligabuesaurus Dongbeititan
Ligabuesaurus
Dongbeititan
Savannasaurus Chubutisaurus Angolatitan
Savannasaurus
Chubutisaurus Angolatitan
Chubutisaurus
Angolatitan
Europatitan Tastavinsaurus Liubangosaurus Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Europatitan
Tastavinsaurus
Liubangosaurus Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Liubangosaurus
Euhelopodidae Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Euhelopodidae
Jiangshanosaurus Titanosauria Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Jiangshanosaurus
TitanosauriaTambatitanis Huabeisaurus Dongyangosaurus Andesaurus Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Tambatitanis Huabeisaurus Dongyangosaurus Andesaurus
Tambatitanis
Huabeisaurus Dongyangosaurus Andesaurus
Huabeisaurus
Dongyangosaurus
Andesaurus
Mongolosaurus Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Mongolosaurus
Vahiny Jainosaurus Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Vahiny Jainosaurus
Vahiny
Jainosaurus
Ruyangosaurus Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Ruyangosaurus
Ruixinia Xianshanosaurus Daxiatitan Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Ruixinia Xianshanosaurus Daxiatitan
Ruixinia
Xianshanosaurus Daxiatitan
Xianshanosaurus
Daxiatitan
Rinconsaurus Muyelensaurus Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Rinconsaurus Muyelensaurus
Rinconsaurus
Muyelensaurus
Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus Lithostrotia
Mendozasaurus Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus
Mendozasaurus
Argentinosaurus Futalognkosaurus Puertasaurus Patagotitan Notocolossus
Argentinosaurus Futalognkosaurus
Argentinosaurus
Futalognkosaurus
Puertasaurus Patagotitan Notocolossus
Puertasaurus
Patagotitan Notocolossus
Patagotitan
Notocolossus
Lithostrotia
Lithostrotia

In their 2024 description of the basal titanosaur Gandititan, Han et al. included Ruixinia in their phylogenetic analyses, recovering it as a member of the Euhelopodidae, rather than the Titanosauria, as the sister taxon to Jiangshanosaurus. The results of their phylogenetic analyses are shown in the cladogram below:

TitanosauriformesBrachiosauridae Somphospondyli Astrophocaudia Brontomerus Malarguesaurus Paluxysaurus Europatitan Tastavinsaurus Fukuititan Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform Euhelopodidae Gobititan Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong Huanghetitan spp. Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Brachiosauridae
SomphospondyliAstrophocaudia Brontomerus Malarguesaurus Paluxysaurus Europatitan Tastavinsaurus Fukuititan Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform Euhelopodidae Gobititan Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong Huanghetitan spp. Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Astrophocaudia Brontomerus Malarguesaurus Paluxysaurus
Astrophocaudia
Brontomerus Malarguesaurus Paluxysaurus
Brontomerus
Malarguesaurus
Paluxysaurus
Europatitan Tastavinsaurus Fukuititan Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform Euhelopodidae Gobititan Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong Huanghetitan spp. Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Europatitan Tastavinsaurus Fukuititan Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform
Europatitan Tastavinsaurus
Europatitan
Tastavinsaurus
Fukuititan Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform
Fukuititan
Dongbeititan Ligabuesaurus Angolatitan Chubutisaurus Cloverly titanosauriform
Dongbeititan Ligabuesaurus
Dongbeititan
Ligabuesaurus
Angolatitan Chubutisaurus Cloverly titanosauriform
Angolatitan
Chubutisaurus Cloverly titanosauriform
Chubutisaurus
Cloverly titanosauriform
Euhelopodidae Gobititan Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong Huanghetitan spp. Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
EuhelopodidaeGobititan Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong
Gobititan
Jiangshanosaurus Ruixinia Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong
Jiangshanosaurus Ruixinia
Jiangshanosaurus
Ruixinia
Phuwiangosaurus Tangvayosaurus Qiaowanlong Euhelopus Yongjinglong
Phuwiangosaurus Tangvayosaurus
Phuwiangosaurus
Tangvayosaurus
Qiaowanlong Euhelopus Yongjinglong
Qiaowanlong
Euhelopus Yongjinglong
Euhelopus
Yongjinglong
Huanghetitan spp. Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Huanghetitan spp.
Diamantinasauria Titanosauria Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Diamantinasauria
TitanosauriaAndesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Andesaurus Baotianmansaurus Dongyangosaurus Huabeisaurus Abdarainurus Gandititan
Andesaurus
Baotianmansaurus
Dongyangosaurus
Huabeisaurus
Abdarainurus Gandititan
Abdarainurus
Gandititan
Daxiatitan Xianshanosaurus Ruyangosaurus Lithostrotia
Daxiatitan Xianshanosaurus
Daxiatitan
Xianshanosaurus
Ruyangosaurus Lithostrotia
Ruyangosaurus
Lithostrotia
Lithostrotia

In 2025, Averianov and colleagues reassessed the phylogenetic relationships of Tengrisaurus and other early titanosaurs and allied taxa. They proposed that the initial interpretation of Ruixinia as a non-lithostrotian titanosaur was supported based on the anatomy of the anterior caudal vertebrae, and since this region of the skeleton is not preserved in undisputed euhelopodids. Their phylogenetic analysis including Ruixinia placed it within Titanosauria, diverging immediately after a clade similar to the one including Andesaurus and Gandititan as found by Han et al. (2024).

Paleoecology

Ruixinia is a member of the Jehol Biota. Two other sauropod genera, Dongbeititan and Liaoningotitan, are also known from the Jehol Biota.