The Salamanca Formation is a geologic formation in the Golfo San Jorge Basin of central Patagonia that yields well-preserved, well-dated fossils from the early Paleocene. Studies of these fossils are providing new data on plant and animal diversity following the end-Cretaceous extinction event.

The Salamanca Formation crops out in the San Jorge Basin in southern Chubut and northern Santa Cruz provinces, Argentina, overlying the Cretaceous Chubut Group and is part of the Paleocene and Eocene Río Chico Group. The formation yields abundant plant remains as well as fossils of invertebrates, marine macrofaunas, reptiles, and mammals. The formation consists primarily of estuarine to shallow marine deposits.

Palynological analysis of the Salamanca Formation shows low floral diversity after the end-Cretaceous mass extinction, followed by a rapid recovery. 50% of all pollen types are angiosperms, whereas gymnosperms accounted for only ~13% of total richness; however, Classopollis pollen, representing the extinct conifer family Cheirolepidiaceae, is the most abundant palynomorph. Wood assemblages from the Salamanca Formation are dominated by conifers, but the presence of fossil angiosperm woods indicate that they were also part of the canopy. The co-occurrence of palms, dicot woods with indistinct growth rings, and alligatorids, indicates temperature remained above freezing year-round. The results of leaf physiognomic analyses indicate that the climate in the San Jorge Basin during the early Paleocene was warm subtropical.

Paleoflora

Angiosperm

NameSpeciesLocalityMaterialNotesImage
AkaniaA. sp.Palacio de los LorosA flowering plant of the family Akaniaceae
BanaraphyllumB. ovatumPalacio de los LorosA flowering plant
CissitesC. patagonicumPalacio de los LorosA flowering plant belonging to Vitaceae, related to the modern genus Cissus
CunonianthaC. bicarpellataA flowering plant belonging to the family Cunoniaceae
DryophyllumD. australisPalacio de los LorosA flowering plant
FagophyllumF. duseniPalacio de los LorosLeaves, related to Fagaceae
LacinipetalumL. spectabilumPalacio de los LorosA flowering plant belonging to the family Cunoniaceae
LaurophyllumL. chubutensis L. piatnitzkyiPalacio de los LorosA flowering plant belonging to the family Lauraceae
MyricaM. premiraPalacio de los LorosA flowering plant belonging to the family Myricaceae
Nothofagussp.Leaves
NotianthaN. grandensisRancho GrandeFlowersA flowering plant belonging to the family Rhamnaceae
PalaeophytocreneP. gaEstancia Las VioletasA fruitA flowering plant belonging to the family Icacinaceae
ParacacioxylonP frenguelliiPalacio de los LorosWoodA flowering plant belonging to the family Mimosoideae
ParanymphaeaP aristolochiaformisPalacio de los LorosA flowering plant belonging to the family Nymphaeaceae
SterculiaS. acuminatalobaPalacio de los LorosA flowering plant belonging to the family Malvaceae
StephaniaS. psittacaPalacio de los LorosA flowering plant belonging to the family Menispermaceae
SuesseniaS. grandensisRancho GrandeLeavesA flowering plant belonging to the family Rhamnaceae
WilkinsoniphyllumW. menispermoidesPalacio de los LorosLeavesA flowering plant belonging to the family Menispermaceae
TripylocarpaT. aestuariaSeedA flowering plant belonging to the palm tribe Cocoseae
VolkheimeritesV. labyrinthusFlowering plant pollen of uncertain placement
Fabaceae
Sapindaceae
Urticaceae
Rosaceae

Gymnosperms

NameSpeciesLocalityMaterialNotesImage
AgathisA. immortalisLeaves, twigs, pollen cones, ovuliferous complexes, pollen, seedsA conifer belonging to Araucariaceae. Oldest known species of Agathis
ClassopollisC. sp.Palacio de los LorosPollen belonging to the extinct conifer family Cheirolepidiaceae
PodocarpaceaeLeaves and cones

Ferns

NameSpeciesLocalityMaterialNotesImage
AzollaA. keujaPalacio de los LorosA fern belonging to the family Salviniaceae
KirketapelK. salamanquensisA scale-leaved conifer belonging to the family Podocarpaceae.
LygodiumSp.A fern belonging to the Schizaeales, another indeterminate distinct fern is also present

Paleofauna

Fish

NameSpeciesLocalityMaterialNotesImage
XiphactinusX. sp.A member of the Ichthyodectiformes.

Amphibians

NameSpeciesLocalityMaterialNotesImage
GigantobatrachusG. casamiquelaiPunta PeligroA frog
XerocephalellaX. sabrosaPunta PeligroA frog belonging to the family Calyptocephalellidae.

Reptiles

Testudines

NameSpeciesLocalityMaterialNotesImage
HydromedusaH. casamayorensisPunta PeligroA member of the family Chelidae.
NajadochelysN. patagonicaPunta Peligro
PeligrochelysP. walshaePunta PeligroA member of Meiolaniformes
SalamanchelysS. palaeocenicaPunta PeligroA member of the family Chelidae.
TrionyxT. argentinaPunta PeligroA member of the family Trionychidae
YaminuechelysY. maiorPunta PeligroA member of the family Chelidae.

Crocodylomorphs

NameSpeciesLocalityMaterialNotesImage
EocaimanE. palaeocenicusPunta PeligroA caimanine alligatorid.Necrosuchus Tewkensuchus
NecrosuchusN. ionensisEstancia Las VioletasA caimanine alligatorid.
ProtocaimanP. peligrensisPunta PeligroA partial skullA caimanine alligatorid.
TewkensuchusT. salamanquensisPunta PeligroA holotype specimen consists of a partial skullA sebecosuchian

Lepidosaurs

NameSpeciesLocalityMaterialNotesImage
KawasphenodonK. peligrensisPunta PeligroA sphenodontian

Mammals

Monotremes

NameSpeciesLocalityMaterialNotesImage
MonotrematumM. sudamericanumPunta PeligroTeeth from the upper and lower jawsA monotreme

Allotheres

NameSpeciesLocalityMaterialNotesImage
SudamericaS. ameghinoiPunta PeligroA gondwanathere

Meridiolestidans

NameSpeciesLocalityMaterialNotesImage
PeligrotheriumP. tropicalisPunta PeligroA member of the clade Mesungulatoidea.

Meridiungulates

NameSpeciesLocalityMaterialNotesImage
EscribaniaE. chubutensis E. talonicuspsisPunta PeligroA member of the family Didolodontidae.
RaulvacciaR. peligrensisPunta PeligroA member of the family Didolodontidae.
RequisiaR. vidmariPunta PeligroA member of the family Notonychopidae.

Metatherians

NameSpeciesLocalityMaterialNotesImage
DerorhynchusD. aff. minutusPunta PeligroA metatherian of uncertain affinities
DidelphopsisIndeterminate
?SparassodontaIndeterminateAstragalus
PolydolopimorphiaIndeterminateTwo taxa, one of which is a member of Bonapartheriidae
MetatheriaIndeterminateRight petrosal (MPEF-PV 2235)

Invertebrates

After unless otherwise noted

NameSpeciesMaterialNotesImage
AcestaA bivalve belonging to Limidae
ArcaA bivalve belonging to Arcidae
?AstarteA bivalve belonging to Astartidae
AporrhaidaeIndeterminateGastropod
AustrophaeraA bivalve
BarbatiaA bivalve belonging to Arcidae
BathytormusA bivalve belonging to Crassatellidae
BuccinidaeFour distinct generaGastropod
CancellariidaeIndeterminateA gastropod
CalyptraeidaeIndeterminateGastropod
CarditesA bivalve belonging to Carditidae
CavoscalaA gastropod belonging to Epitoniidae
ClaibornicardiaA bivalve belonging to Carditidae
CostacoplumaC. salamancaA crab belonging to the family Retroplumidae
CrenilabiumA gastropod belonging to Acteonidae
CubitostreaA bivalve belonging to Ostreidae
CucullaeaBivalve
DarwinicesA gastropod belonging to Naticidae
?DosiniaA bivalve belonging to Veneridae
FusinusA gastropod belonging to Fasciolariidae
FyfeaA gastropod
GlobisininaeIndeterminateGastropods belonging to the family Naticidae
GlycymeritaA bivalve belonging to Glycymerididae
GryphaostreaA bivalve
GyroscalaA gastropod belonging to Epitoniidae
HarpidaeNov. Gen.A gastropod
HeterotermaA gastropod belonging to Tudiclidae
?LarmaA bivalve belonging to Veneridae
LedinaA bivalve belonging to Nuculanidae
LeionuculaA bivalve belonging to Nuculidae
LucinidaeIndeterminateBivalve
MarwickiaA bivalve belonging to Veneridae
MicrofulgurA gastropod belonging to Fasciolariidae
NeiloA bivalve belonging to Malletiidae
NuculaA bivalve belonging to Nuculidae
PanopeaA bivalve belonging to Hiatellidae
PatellogastropodaIndeterminateGastropod
PerissodontaA gastropod belonging to Struthiolariidae
PhacoidesA bivalve belonging to Lucinidae
PinnaA bivalve belonging to Pinnidae
PriscaphanderA gastropod belonging to Scaphandridae
PriscophicusA gastropod belonging to Ficidae
PseudomauraA gastropod
PteromyrteaA bivalve belonging to Lucinidae
PseudotylostomaA gastropod
PycnodonteA bivalve belonging to Gryphaeidae
RingiculidaeIndeterminateA gastropod
RocalariaA gastropod belonging to Fasciolariidae
RotundicardiaA bivalve belonging to Carditidae
SacellaA bivalve belonging to Nuculanidae
SpineiloA bivalve belonging to Malletiidae
StruthiopteraA gastropod belonging to Aporrhaidae
SulcobuccinumA gastropod belonging to Pseudolividae
TaiomaA gastropod belonging to Fasciolariidae
TalochlamysA bivalve belonging to Pectinidae
TejoniaA gastropod belonging to Naticidae
TitomayaT. longobucca & T. chalcedonicaA bivalve belonging to Venerida
TornatellaeaA gastropod belonging to Acteonidae
TurritellidaeNov. gen.A gastropod
VenericardiaA bivalve belonging to Carditidae
VolutidaeIndeterminateGastropod
YoldiaA bivalve belonging to Yoldiidae
?ZemaciesA gastropod belonging to Borsoniidae