Salamanca Formation
Salamanca Formation
Stratigraphic range : latest Maastrichtian -Danian (Tiupampan -Riochican ) ~66.0–57 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Unit of Río Chico Group
Underlies Peñas Coloradas Formation (Río Chico Group)
Lithology
Location
Country Argentina
Type section
The Salamanca Formation is a geologic formation in the Golfo San Jorge Basin of central Patagonia that yields well-preserved, well-dated fossils from the early Paleocene. Studies of these fossils are providing new data on plant and animal diversity following the end-Cretaceous extinction event .
The Salamanca Formation crops out in the San Jorge Basin in southern Chubut and northern Santa Cruz provinces, Argentina, overlying the Cretaceous Chubut Group and is part of the Paleocene and Eocene Río Chico Group. The formation yields abundant plant remains as well as fossils of invertebrates , marine macrofaunas, reptiles , and mammals . The formation consists primarily of estuarine to shallow marine deposits.
Palynological analysis of the Salamanca Formation shows low floral diversity after the end-Cretaceous mass extinction, followed by a rapid recovery. 50% of all pollen types are angiosperms, whereas gymnosperms accounted for only ~13% of total richness; however, Classopollis pollen , representing the extinct conifer family Cheirolepidiaceae , is the most abundant palynomorph. Wood assemblages from the Salamanca Formation are dominated by conifers, but the presence of fossil angiosperm woods indicate that they were also part of the canopy. The co-occurrence of palms, dicot woods with indistinct growth rings, and alligatorids , indicates temperature remained above freezing year-round. The results of leaf physiognomic analyses indicate that the climate in the San Jorge Basin during the early Paleocene was warm subtropical.
Paleoflora Angiosperm Name Species Locality Material Notes Image Akania A. sp.Palacio de los Loros A flowering plant of the family Akaniaceae Banaraphyllum B. ovatum Palacio de los Loros A flowering plant Cissites C. patagonicum Palacio de los Loros A flowering plant belonging to Vitaceae , related to the modern genus Cissus Cunoniantha C. bicarpellata A flowering plant belonging to the family Cunoniaceae Dryophyllum D. australis Palacio de los Loros A flowering plant Fagophyllum F. duseni Palacio de los Loros Leaves, related to Fagaceae Lacinipetalum L. spectabilum Palacio de los Loros A flowering plant belonging to the family Cunoniaceae Laurophyllum L. chubutensis L. piatnitzkyi Palacio de los Loros A flowering plant belonging to the family Lauraceae Myrica M. premira Palacio de los Loros A flowering plant belonging to the family Myricaceae Nothofagus sp. Leaves Notiantha N. grandensis Rancho Grande Flowers A flowering plant belonging to the family Rhamnaceae Palaeophytocrene P. ga Estancia Las Violetas A fruit A flowering plant belonging to the family Icacinaceae Paracacioxylon P frenguellii Palacio de los Loros Wood A flowering plant belonging to the family Mimosoideae Paranymphaea P aristolochiaformis Palacio de los Loros A flowering plant belonging to the family Nymphaeaceae Sterculia S. acuminataloba Palacio de los Loros A flowering plant belonging to the family Malvaceae Stephania S. psittaca Palacio de los Loros A flowering plant belonging to the family Menispermaceae Suessenia S. grandensis Rancho Grande Leaves A flowering plant belonging to the family Rhamnaceae Wilkinsoniphyllum W. menispermoides Palacio de los Loros Leaves A flowering plant belonging to the family Menispermaceae Tripylocarpa T. aestuaria Seed A flowering plant belonging to the palm tribe Cocoseae Volkheimerites V. labyrinthus Flowering plant pollen of uncertain placement Fabaceae Sapindaceae Urticaceae Rosaceae
Gymnosperms Name Species Locality Material Notes Image Agathis A. immortalis Leaves, twigs, pollen cones, ovuliferous complexes, pollen, seeds A conifer belonging to Araucariaceae . Oldest known species of Agathis Classopollis C. sp.Palacio de los Loros Pollen belonging to the extinct conifer family Cheirolepidiaceae Podocarpaceae Leaves and cones
Ferns Name Species Locality Material Notes Image Azolla A. keuja Palacio de los Loros A fern belonging to the family Salviniaceae Kirketapel K. salamanquensis A scale-leaved conifer belonging to the family Podocarpaceae . Lygodium Sp. A fern belonging to the Schizaeales , another indeterminate distinct fern is also present
Paleofauna Fish Name Species Locality Material Notes Image Xiphactinus X. sp.A member of the Ichthyodectiformes .
Amphibians Name Species Locality Material Notes Image Gigantobatrachus G. casamiquelai Punta Peligro A frog Xerocephalella X. sabrosa Punta Peligro A frog belonging to the family Calyptocephalellidae .
Reptiles Testudines Name Species Locality Material Notes Image Hydromedusa H. casamayorensis Punta Peligro A member of the family Chelidae . Najadochelys N. patagonica Punta Peligro Peligrochelys P. walshae Punta Peligro A member of Meiolaniformes Salamanchelys S. palaeocenica Punta Peligro A member of the family Chelidae. Trionyx T. argentina Punta Peligro A member of the family Trionychidae Yaminuechelys Y. maior Punta Peligro A member of the family Chelidae.
Crocodylomorphs Name Species Locality Material Notes Image Eocaiman E. palaeocenicus Punta Peligro A caimanine alligatorid. Necrosuchus Tewkensuchus Necrosuchus N. ionensis Estancia Las Violetas A caimanine alligatorid. Protocaiman P. peligrensis Punta Peligro A partial skull A caimanine alligatorid. Tewkensuchus T. salamanquensis Punta Peligro A holotype specimen consists of a partial skull A sebecosuchian
Lepidosaurs Name Species Locality Material Notes Image Kawasphenodon K. peligrensis Punta Peligro A sphenodontian
Mammals Monotremes Name Species Locality Material Notes Image Monotrematum M. sudamericanum Punta Peligro Teeth from the upper and lower jaws A monotreme
Allotheres Name Species Locality Material Notes Image Sudamerica S. ameghinoi Punta Peligro A gondwanathere
Meridiolestidans Name Species Locality Material Notes Image Peligrotherium P. tropicalis Punta Peligro A member of the clade Mesungulatoidea .
Meridiungulates Name Species Locality Material Notes Image Escribania E. chubutensis E. talonicuspsis Punta Peligro A member of the family Didolodontidae . Raulvaccia R. peligrensis Punta Peligro A member of the family Didolodontidae. Requisia R. vidmari Punta Peligro A member of the family Notonychopidae.
Metatherians Name Species Locality Material Notes Image Derorhynchus D. aff. minutus Punta Peligro A metatherian of uncertain affinities Didelphopsis Indeterminate ?Sparassodonta Indeterminate Astragalus Polydolopimorphia Indeterminate Two taxa, one of which is a member of Bonapartheriidae Metatheria Indeterminate Right petrosal (MPEF-PV 2235)
Invertebrates After unless otherwise noted
Name Species Material Notes Image Acesta A bivalve belonging to Limidae Arca A bivalve belonging to Arcidae ?Astarte A bivalve belonging to Astartidae Aporrhaidae Indeterminate Gastropod Austrophaera A bivalve Barbatia A bivalve belonging to Arcidae Bathytormus A bivalve belonging to Crassatellidae Buccinidae Four distinct genera Gastropod Cancellariidae Indeterminate A gastropod Calyptraeidae Indeterminate Gastropod Cardites A bivalve belonging to Carditidae Cavoscala A gastropod belonging to Epitoniidae Claibornicardia A bivalve belonging to Carditidae Costacopluma C. salamanca A crab belonging to the family Retroplumidae Crenilabium A gastropod belonging to Acteonidae Cubitostrea A bivalve belonging to Ostreidae Cucullaea Bivalve Darwinices A gastropod belonging to Naticidae ?Dosinia A bivalve belonging to Veneridae Fusinus A gastropod belonging to Fasciolariidae Fyfea A gastropod Globisininae Indeterminate Gastropods belonging to the family Naticidae Glycymerita A bivalve belonging to Glycymerididae Gryphaostrea A bivalve Gyroscala A gastropod belonging to Epitoniidae Harpidae Nov. Gen. A gastropod Heteroterma A gastropod belonging to Tudiclidae ?Larma A bivalve belonging to Veneridae Ledina A bivalve belonging to Nuculanidae Leionucula A bivalve belonging to Nuculidae Lucinidae Indeterminate Bivalve Marwickia A bivalve belonging to Veneridae Microfulgur A gastropod belonging to Fasciolariidae Neilo A bivalve belonging to Malletiidae Nucula A bivalve belonging to Nuculidae Panopea A bivalve belonging to Hiatellidae Patellogastropoda Indeterminate Gastropod Perissodonta A gastropod belonging to Struthiolariidae Phacoides A bivalve belonging to Lucinidae Pinna A bivalve belonging to Pinnidae Priscaphander A gastropod belonging to Scaphandridae Priscophicus A gastropod belonging to Ficidae Pseudomaura A gastropod Pteromyrtea A bivalve belonging to Lucinidae Pseudotylostoma A gastropod Pycnodonte A bivalve belonging to Gryphaeidae Ringiculidae Indeterminate A gastropod Rocalaria A gastropod belonging to Fasciolariidae Rotundicardia A bivalve belonging to Carditidae Sacella A bivalve belonging to Nuculanidae Spineilo A bivalve belonging to Malletiidae Struthioptera A gastropod belonging to Aporrhaidae Sulcobuccinum A gastropod belonging to Pseudolividae Taioma A gastropod belonging to Fasciolariidae Talochlamys A bivalve belonging to Pectinidae Tejonia A gastropod belonging to Naticidae Titomaya T. longobucca & T. chalcedonica A bivalve belonging to Venerida Tornatellaea A gastropod belonging to Acteonidae Turritellidae Nov. gen. A gastropod Venericardia A bivalve belonging to Carditidae Volutidae Indeterminate Gastropod Yoldia A bivalve belonging to Yoldiidae ?Zemacies A gastropod belonging to Borsoniidae