A black-and-white drawing of an Oncopus truncatus Thorrell, facing right, with 8 legs
Drawing of female Oncopus truncatus Thorrell

Sandokanidae is a family of harvestmen in the suborder Laniatores, formerly referred to as Oncopodidae (the name was replaced because of the secondary homonymy of the type genus Oncopus, replaced by Sandokan; this made the change of family name mandatory)

Name

The name of the type genus is based on the name of a fictional Bornean prince-pirate.

Description

Sandokanidae range in body size from about 2–11 mm. Their legs are relatively short and stout. Most species are amber colored with some dark brown patterns. A few undescribed Gnomulus species are orange.

Distribution

This family is known from Southeast Asia from Indonesia, almost reaching New Guinea on Waigeo, up north into the Himalayan region.

Relationships

Sandokanidae are probably the sister group to all other Grassatores.

Species

  • Biantoncopus Martens & Schwendinger, 1998

Biantoncopus fuscus Martens & Schwendinger, 1998 — Leyte (Philippines)

  • Caenoncopus Martens & Schwendinger, 1998 — Sumatra

Caenoncopus affinis Martens & Schwendinger, 1998 Caenoncopus cuspidatus (Schwendinger, 1992) Caenoncopus tenuis Martens & Schwendinger, 1998

  • Gnomulus Thorell, 1890

Gnomulus aborensis (Roewer, 1913) — Arunachal Pradesh (northeast India) Gnomulus annulipes (Pocock, 1897) — Sarawak (Borneo) Gnomulus armillatus (Thorell, 1891) — Sumatra Gnomulus asli Martens & Schwendinger, 1998 — West Malaysia Gnomulus baharu Schwendinger, in Martens & Schwendinger 1998 Gnomulus carinatus Schwendinger & Martens, 2002 — Kalimantan (Indonesia) Gnomulus claviger Schwendinger & Martens, 2002 — Luzon (Philippines) Gnomulus coniceps Martens & Schwendinger, 1998 — Luzon Gnomulus conigerus (Schwendinger, 1992) — Sabah (Malaysia) Gnomulus crassipes Schwendinger & Martens, 2002 — Luzon Gnomulus crucifer Martens & Schwendinger, 1998 — Luzon Gnomulus drescoi (Silhavy, 1962) — Sumatra Gnomulus exsudans Schwendinger & Martens, 2002 — Sabah Gnomulus goodnighti (Suzuki, 1977) — Mindanao (Philippines) Gnomulus hamatus Schwendinger & Martens, 2002 — Luzon Gnomulus hirsutus Martens & Schwendinger, 1998 — West Malaysia Gnomulus hutan Schwendinger & Martens, 2002 — Sarawak Gnomulus hyatti (Martens, 1977) — Nepal Gnomulus imadatei (Suzuki, 1970) — Brunei Gnomulus insularis (Roewer, 1927) — Penang? (Malaysia) Gnomulus javanicus Schwendinger & Martens, 2002 — Java Gnomulus laevis (Roewer, 1915) — Borneo Gnomulus lannaianus (Schwendinger, 1992) — Thailand Gnomulus laruticus Martens & Schwendinger, 1998 — West Malaysia Gnomulus latoperculum Schwendinger & Martens, 2002 — Sulawesi Gnomulus leofeae Schwendinger & Martens, 2002 — Burma Gnomulus leyteensis Martens & Schwendinger, 1998 — Leyte Gnomulus lomani Schwendinger & Martens, 2002 — Borneo, Sumatra? Gnomulus maculatus Martens & Schwendinger, 1998 — Luzon Gnomulus marginatus Schwendinger & Martens, 2002 — Thailand Gnomulus matabesar Schwendinger & Martens, 2002 — Halmahera (Indonesia) Gnomulus minor N. Tsurusaki, 1990 — Philippines Gnomulus monticola Schwendinger & Martens, 2002 — West Malaysia Gnomulus obscurus Schwendinger & Martens, 2002 — Sarawak Gnomulus palawanensis (S. Suzuki, 1982) — Palawan (Philippines) Gnomulus piliger (Pocock, 1903) — Thailand Gnomulus pilosus Schwendinger & Martens, 2002 — West Malaysia Gnomulus pulvillatus (Pocock, 1903) — West Malaysia Gnomulus rostratoideus Schwendinger & Martens, 2002 — West Malaysia, Singapore Gnomulus rostratus Thorell, 1890 — Penang Gnomulus ryssie Schwendinger & Martens, 2002 — Thailand Gnomulus sinensis Schwendinger & Martens, 2002 — Sichuan (China) Gnomulus spiniceps Schwendinger & Martens, 2002 — Vietnam Gnomulus sumatranus Thorell, 1891 — Sumatra Gnomulus sundaicus (Schwendinger, 1992) — Sarawak Gnomulus thorelli (Sørensen, 1932) — Java Gnomulus tuberculatus Schwendinger & Martens, 2002 — Sumatra Gnomulus tumidifrons Schwendinger & Martens, 2002 — Halmahera

  • Sandokan Özdikmen & Kury, 2007

Sandokan doriae Thorell, 1876 — Sarawak Sandokan expatriatus (Schwendinger & Martens, 2004) — Thailand? Sandokan feae Thorell, 1890 — Penang Sandokan hosei Pocock, 1897 — Sarawak Sandokan lingga (Schwendinger & Martens, 2004) — Lingga Islands (Indonesia) Sandokan malayanus (Schwendinger & Martens, 2004) — West Malaysia Sandokan megachelis (Schwendinger, 1992) — Sabah Sandokan tiomanensis (Schwendinger & Martens, 2004) — Tioman Island (Malaysia) Sandokan truncatus Thorell, 1891 — Singapore

  • Palaeoncopus Martens & Schwendinger, 1998 — Sumatra

Palaeoncopus gunung Martens & Schwendinger, 1998 Palaeoncopus katik Martens & Schwendinger, 1998 Palaeoncopus kerdil Martens & Schwendinger, 1998

  • Joel Hallan's Biology Catalog: