The Shaximiao Formation (simplified Chinese: 沙溪庙组; traditional Chinese: 沙溪廟組 / 沙溪廟層; pinyin: Shāxīmiào zǔ) is a Middle to Late Jurassic aged geological formation in Sichuan, China, most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the small township of Dashanpu (simplified Chinese: 大山铺镇; traditional Chinese: 大山鋪鎮; pinyin: Dàshānpū zhèn), situated seven kilometres north-east from Sichuan's third largest city, Zigong, in the Da'an District.

Geology

The Shaximiao Formation includes two distinct subunits: The upper and lower Shaximiao Formations (simplified Chinese: 上·下沙溪庙地层; traditional Chinese: 上·下沙溪廟地層; pinyin: shàng / xià Shāxīmiào dìcéng), although they are commonly referred to as one, simply being called the "Shaximiao Formation". The upper Shaximiao Formation is also known as the Shangshaximiao Formation, and the lower Shaximiao Formation is also known as the Xiashaximiao Formation, which are direct transliterations of the Chinese names. Both subunits primarily consist of purple-red mudstones, with variable sand inclusion. and siltstones with interbedded sandstones.

Dinosaur finds

Dinosaurs of the Shaximiao Formation

The Shaximiao Formation has produced mainly sauropods, but has also held numerous other dinosaur types, such as theropods and stegosaurians amongst others. In total, over 8,000 pieces of bone have been unearthed from the area – amounting to nearly 40 tonnes. The site was unknown until the early 1970s, when a Chinese gas company unearthed Gasosaurus in 1972. It would be the first of the many dinosaurs to be uncovered from the area. Most specimens found are held at the Zigong Dinosaur Museum which has been placed on the area during the mid-1980s.

The site is currently a frequented "dinosaur-quarry", but the Shaximiao Formation once hosted a lush forest, evidenced by fossilised wood found alongside the dinosaur remains. Paleontologists speculate that the area also had a lake that was fed by a large river.[clarification needed] Dinosaur remains would have been swept toward the lake over millions of years, thus accounting for the hundreds of specimens found. Based on biostratigraphy, the Lower Shaximiao Formation has been usually seen to date to 168 to 161 million years old, between the Bathonian to Callovian stages of the Mid Jurassic, while the Upper Shaximiao was thought to be Oxfordian in age. A paper by Wang et al. (2018), (recalibrated dates reported in Moore et al. (2020)), reported a zircon U-Pb age of 160.4 ± 0.4 mya for the lower part of the Shaximiao Formation, suggesting that the Shaximiao Formation is younger than previously thought. Contradicting this, U-Pb dates from a tuff bed supports traditional Middle Jurassic age for the lower part of the formation, with an average age of around 166.0 ± 1.5 Ma. The top of the Shaximiao Formation has been believed to be the end of the Tithonian age.

Dong Zhiming's research

The paleontologist who has made the largest contribution to the formation and its excavation is Dong Zhiming. He first examined the formation in 1975, after bone fragments were found embedded in rock from the area. The site was being demolished to make way for both a natural gas field facility and a vehicle park when Dong first saw the area. Amongst the extensive clearings, Dong found numerous bone fragments which were exposed.

However, the specimens were being damaged due to bulldozers in the area and there would be little chance of closing the area as the state had invested millions of yuan in the site already. It was not until 1985 that the government finally agreed to close the construction on the site, and by then Dong and his team had already excavated over 100 dinosaurs from the area, including several rare sauropod skulls. A dinosaur found in the Shaximiao Formation, Dashanpusaurus dongi, was named in tribute of both Dashanpu and Dong Zhiming.

Paleobiota

In addition to dinosaur finds, many other prehistoric finds have been uncovered from the Shaximiao Formation. Amongst these finds are fishes, amphibians, turtles, marine reptiles such as crocodiles and also pterosaurs. Bienotheroides, a Tritylodont Synapsid has been found there, as well as Sinobrachyops, a Labyrinthodont.

Amphibians

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
SinobrachyopsS. placenticephalusShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA brachyopid stereospondyl

Fish

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
CeratodusC. guanganensisMamenchisaurus assemblage, Upper Shaximiao FormationA ceratodontid lungfish
C. minorMamenchisaurus assemblage, Upper Shaximiao Formation
C. youngiMamenchisaurus assemblage, Upper Shaximiao Formation
C. zigongensisMamenchisaurus assemblage, Upper Shaximiao Formation, Shunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
ChungkingichthysC. tachuensisMamenchisaurus assemblage, Upper Shaximiao FormationA ptycholepiform fish
HybodusH.sp.Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA hybodont chondrichthyan
IsanichthysI. luchowensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA ginglymodian fish, originally identified as Lepidotes
YuchoulepisY. szechuanensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA ptycholepiform fish

Sauropterygians

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
BishanopliosaurusB. zigongensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA pliosauroid plesiosaur

Crocodyliformes

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
SunosuchusS. shunanensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA goniopholidid
HsisosuchusH. dashanpuensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA goniopholid

Pterosaurs

GenusSpeciesStratigraphic positionMaterialNotesImages
AngustinaripterusA. longicephalusShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA rhamphorhynchine rhamphorhynchid with a very elongated and flat snout and an estimated wingspan at 1.6 metres (5.2 ft)

Dinosaurs

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Ornithischians

Neornithischians

GenusSpeciesStratigraphic positionMaterialNotesImages
AgilisaurusA. louderbackiShunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
GongbusaurusG. shiyiiMamenchisaurus assemblage, Upper Shaximiao FormationTeeth
HexinlusaurusH. multidensShunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
XiaosaurusX. dashanpuensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
YandusaurusY. hongheensisMamenchisaurus assemblage, Upper Shaximiao Formation

Stegosaurs

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
BashanosaurusB. primitivusLower Shaximiao FormationThe earliest-diverging stegosaur currently known; it shares features with basal thyreophorans and derived stegosaurs.
ChialingosaurusC. kuaniMamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian.
ChungkingosaurusC. jiangbeiensisMamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian.
HuayangosaurusH. taibaiiShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA huayangosaurid with more spike-like plates than Stegosaurus. It was also one of the smallest known stegosaurs, at just 4.5 meters (15 feet) in length.
GigantspinosaurusG. sichuanensisMamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian.
G. sp.Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian.
TuojiangosaurusT. multispinusMamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian.
YingshanosaurusY. jichuanensisUpper Shaximiao FormationNearly complete skeletonA stegosaurian.

Sauropods

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
AbrosaurusA. dongpoiLower Shaximiao FormationA macronarian known from a well-preserved skull and other elements.
BashunosaurusB. kaijiangensisLower Shaximiao FormationAn obscure possible macronarian
DaanosaurusD. zhangiMamenchisaurus assemblage, Upper Shaximiao Formation
DashanpusaurusD. dongiLower Shaximiao FormationA basal macronarian
DatousaurusD. bashanensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid.
MamenchisaurusM. constructusMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
M. hochuanensisMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
M. youngiMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
M. jingyanensisMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
M. sanjiangensisUpper Shaximiao FormationA single partial skeletonA mamenchisaurid.
OmeisaurusO. luoquanensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid.
O. tianfuensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid.
O. junghsiensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid.
O. changshouensisMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
O. maoianusMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
O. fuxiensisMamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid.
ProtognathosaurusP. oxyodonShunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
ShunosaurusS. liiShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA eusauropod estimated with a length at 9.5 meters (31 feet), the weight at 3 metric tons (3.3 short tons). The tail ended in a club, equipped on its top with two successive spikes formed by 5 centimetres (2.0 inches)-long cone-shaped osteoderms probably used to fend off enemies.
YuzhoulongY. qurenensisLower Shaximiao FormationA basal macronarian known from a subadult specimen
ZigongosaurusZ. fuxiensisMamenchisaurus assemblage, Upper Shaximiao FormationPossibly a species of Mamenchisaurus.

Theropods

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesSynonymsStratigraphic positionMaterialNotesImages
ChienkosaurusC. ceratosauroidesMamenchisaurus assemblage, Upper Shaximiao Formation
ChuandongocoelurusC. primitivusShunosaurus-Omeisaurus assemblage, Lower Shaximiao Formation
Elaphrosaurinae indet.Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationFormerly designated as the paratype of Chuandongocoelurus.
GasosaurusG. constructusShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationAn avetheropod that measured between 3.5 and 4 meters (11 and 13 ft) in length, with a weight of around 150 kilograms (330 pounds). However, some estimates put its weight as high as 400 kilograms (880 pounds), as very little is known about it.
KaijiangosaurusK. liniShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA megalosaurid, early tetanuran, early averostran or basal carnosaur.
LeshansaurusL. qianweiensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao formationA megalosaurid.
SinocoelurusS. fragilisMamenchisaurus assemblage, Upper Shaximiao FormationA theropod of uncertain affinities
SinraptorS. hepingensisMamenchisaurus assemblage, Upper Shaximiao FormationA metriacanthosaurid, formally named Yangchuanosaurus hepingensis.
SzechuanosaurusS. campiMamenchisaurus assemblage, Upper Shaximiao FormationA metriacanthosaurid. Remains from the Shaximiao Formation might represent an indeterminate theropod.
XuanhanosaurusX. qilixiaensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA metriacanthosaurid.
YangchuanosaurusY. shangyouensisY. magnusMamenchisaurus assemblage, Upper Shaximiao formationA metriacanthosaurid
Y. zigongensisSzechuanosaurus zigongensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationZDM 9011 (holotype), a partial postcranial skeleton; ZDM 9012, a left maxilla; ZDM 9013, two teeth and ZDM 9014, a right hind limb.

Turtles

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
ChengyuchelysC. baenoidesShangshaximiao FormationA xinjiangchelyid.
C. dashanpuensisShangshaximiao FormationA xinjiangchelyid.
C. latimarginalisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA xinjiangchelyid.
SichuanchelysS. chowiShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA sichuanchelyid.
TienfuchelysT. zigongensisShangshaximiao FormationA xinjiangchelyid.
SinaspideretesS. wimaniShangshaximiao FormationA trionychian, possibly the oldest known cryptodire

Therapsids

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotesImages
BienotheroidesB. zigongensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA tritylodontid.
PolistodonP. chuannanensisShunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA tritylodontid.
Tritylodontidaegen. et sp. indet.Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationAn indeterminate tritylodontid.

Plants

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
GenusSpeciesStratigraphic positionMaterialNotes
BrachyoxylonB. qijiangenseMasangyan villageLeavesFamily unknown.

See also

External links