Shaximiao Formation
Shaximiao Formation
Stratigraphic range : Middle Jurassic –Late Jurassic PreꞒ Ꞓ O S D C P T J K Pg N
Unit of Chongqing Group
Sub-units Lower, Upper Subunits
Thickness Lower Subunit: Up to 246.5 m (809 ft) Upper Subunit: 450–1,200 m (1,480–3,940 ft)
Lithology
Location
Coordinates 29°12′N 105°54′E/29.2°N 105.9°E/ 29.2; 105.9
Approximate paleocoordinates 28°12′N 110°00′E/28.2°N 110.0°E/ 28.2; 110.0
The Shaximiao Formation (simplified Chinese : 沙溪庙组; traditional Chinese : 沙溪廟組 / 沙溪廟層; pinyin : Shāxīmiào zǔ ) is a Middle to Late Jurassic aged geological formation in Sichuan , China , most notable for the wealth of dinosaurs fossils that have been excavated from its strata. The Shaximiao Formation is exposed in and around the small township of Dashanpu (simplified Chinese : 大山铺镇; traditional Chinese : 大山鋪鎮; pinyin : Dàshānpū zhèn ), situated seven kilometres north-east from Sichuan 's third largest city, Zigong , in the Da'an District .
Geology The Shaximiao Formation includes two distinct subunits: The upper and lower Shaximiao Formations (simplified Chinese : 上·下沙溪庙地层; traditional Chinese : 上·下沙溪廟地層; pinyin : shàng / xià Shāxīmiào dìcéng ), although they are commonly referred to as one, simply being called the "Shaximiao Formation". The upper Shaximiao Formation is also known as the Shangshaximiao Formation , and the lower Shaximiao Formation is also known as the Xiashaximiao Formation , which are direct transliterations of the Chinese names. Both subunits primarily consist of purple-red mudstones, with variable sand inclusion. and siltstones with interbedded sandstones.
Dinosaur finds Dinosaurs of the Shaximiao Formation The Shaximiao Formation has produced mainly sauropods , but has also held numerous other dinosaur types, such as theropods and stegosaurians amongst others. In total, over 8,000 pieces of bone have been unearthed from the area – amounting to nearly 40 tonnes . The site was unknown until the early 1970s, when a Chinese gas company unearthed Gasosaurus in 1972. It would be the first of the many dinosaurs to be uncovered from the area. Most specimens found are held at the Zigong Dinosaur Museum which has been placed on the area during the mid-1980s.
The site is currently a frequented "dinosaur-quarry", but the Shaximiao Formation once hosted a lush forest , evidenced by fossilised wood found alongside the dinosaur remains. Paleontologists speculate that the area also had a lake that was fed by a large river.[clarification needed ] Dinosaur remains would have been swept toward the lake over millions of years, thus accounting for the hundreds of specimens found. Based on biostratigraphy , the Lower Shaximiao Formation has been usually seen to date to 168 to 161 million years old, between the Bathonian to Callovian stages of the Mid Jurassic, while the Upper Shaximiao was thought to be Oxfordian in age. A paper by Wang et al. (2018), (recalibrated dates reported in Moore et al. (2020)), reported a zircon U -Pb age of 160.4 ± 0.4 mya for the lower part of the Shaximiao Formation, suggesting that the Shaximiao Formation is younger than previously thought. Contradicting this, U-Pb dates from a tuff bed supports traditional Middle Jurassic age for the lower part of the formation, with an average age of around 166.0 ± 1.5 Ma. The top of the Shaximiao Formation has been believed to be the end of the Tithonian age.
Dong Zhiming's research The paleontologist who has made the largest contribution to the formation and its excavation is Dong Zhiming . He first examined the formation in 1975, after bone fragments were found embedded in rock from the area. The site was being demolished to make way for both a natural gas field facility and a vehicle park when Dong first saw the area. Amongst the extensive clearings, Dong found numerous bone fragments which were exposed.
However, the specimens were being damaged due to bulldozers in the area and there would be little chance of closing the area as the state had invested millions of yuan in the site already. It was not until 1985 that the government finally agreed to close the construction on the site, and by then Dong and his team had already excavated over 100 dinosaurs from the area, including several rare sauropod skulls. A dinosaur found in the Shaximiao Formation, Dashanpusaurus dongi , was named in tribute of both Dashanpu and Dong Zhiming.
Paleobiota In addition to dinosaur finds, many other prehistoric finds have been uncovered from the Shaximiao Formation. Amongst these finds are fishes , amphibians , turtles , marine reptiles such as crocodiles and also pterosaurs . Bienotheroides , a Tritylodont Synapsid has been found there, as well as Sinobrachyops , a Labyrinthodont .
Amphibians Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Sinobrachyops S. placenticephalus Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA brachyopid stereospondyl
Fish Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Ceratodus C. guanganensis Mamenchisaurus assemblage, Upper Shaximiao FormationA ceratodontid lungfish C. minor Mamenchisaurus assemblage, Upper Shaximiao FormationC. youngi Mamenchisaurus assemblage, Upper Shaximiao FormationC. zigongensis Mamenchisaurus assemblage, Upper Shaximiao Formation, Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationChungkingichthys C. tachuensis Mamenchisaurus assemblage, Upper Shaximiao FormationA ptycholepiform fish Hybodus H. sp.Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA hybodont chondrichthyan Isanichthys I. luchowensis Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA ginglymodian fish, originally identified as Lepidotes Yuchoulepis Y. szechuanensis Shunosaurus-Omeisaurus assemblage, Lower Shaximiao FormationA ptycholepiform fish
Sauropterygians Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Bishanopliosaurus B. zigongensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA pliosauroid plesiosaur
Crocodyliformes Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Sunosuchus S. shunanensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA goniopholidid Hsisosuchus H. dashanpuensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA goniopholid
Pterosaurs Genus Species Stratigraphic position Material Notes Images Angustinaripterus A. longicephalus Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA rhamphorhynchine rhamphorhynchid with a very elongated and flat snout and an estimated wingspan at 1.6 metres (5.2 ft)
Dinosaurs Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Ornithischians Neornithischians Genus Species Stratigraphic position Material Notes Images Agilisaurus A. louderbacki Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationGongbusaurus G. shiyii Mamenchisaurus assemblage, Upper Shaximiao FormationTeeth Hexinlusaurus H. multidens Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationXiaosaurus X. dashanpuensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationYandusaurus Y. hongheensis Mamenchisaurus assemblage, Upper Shaximiao Formation
Stegosaurs Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Bashanosaurus B. primitivus Lower Shaximiao Formation The earliest-diverging stegosaur currently known; it shares features with basal thyreophorans and derived stegosaurs. Chialingosaurus C. kuani Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian. Chungkingosaurus C. jiangbeiensis Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian. Huayangosaurus H. taibaii Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA huayangosaurid with more spike-like plates than Stegosaurus . It was also one of the smallest known stegosaurs , at just 4.5 meters (15 feet) in length. Gigantspinosaurus G. sichuanensis Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian. G. sp.Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian. Tuojiangosaurus T. multispinus Mamenchisaurus assemblage, Upper Shaximiao FormationA stegosaurian. Yingshanosaurus Y. jichuanensis Upper Shaximiao Formation Nearly complete skeleton A stegosaurian.
Sauropods Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Abrosaurus A. dongpoi Lower Shaximiao Formation A macronarian known from a well-preserved skull and other elements. Bashunosaurus B. kaijiangensis Lower Shaximiao Formation An obscure possible macronarian Daanosaurus D. zhangi Mamenchisaurus assemblage, Upper Shaximiao FormationDashanpusaurus D. dongi Lower Shaximiao Formation A basal macronarian Datousaurus D. bashanensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid. Mamenchisaurus M. constructus Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. M. hochuanensis Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. M. youngi Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. M. jingyanensis Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. M. sanjiangensis Upper Shaximiao Formation A single partial skeleton A mamenchisaurid. Omeisaurus O. luoquanensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid . O. tianfuensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid. O. junghsiensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA mamenchisaurid. O. changshouensis Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. O. maoianus Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. O. fuxiensis Mamenchisaurus assemblage, Upper Shaximiao FormationA mamenchisaurid. Protognathosaurus P. oxyodon Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationShunosaurus S. lii Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA eusauropod estimated with a length at 9.5 meters (31 feet), the weight at 3 metric tons (3.3 short tons). The tail ended in a club, equipped on its top with two successive spikes formed by 5 centimetres (2.0 inches)-long cone-shaped osteoderms probably used to fend off enemies. Yuzhoulong Y. qurenensis Lower Shaximiao Formation A basal macronarian known from a subadult specimen Zigongosaurus Z. fuxiensis Mamenchisaurus assemblage, Upper Shaximiao FormationPossibly a species of Mamenchisaurus.
Theropods Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Synonyms Stratigraphic position Material Notes Images Chienkosaurus C. ceratosauroides Mamenchisaurus assemblage, Upper Shaximiao FormationChuandongocoelurus C. primitivus Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationElaphrosaurinae indet.Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationFormerly designated as the paratype of Chuandongocoelurus . Gasosaurus G. constructus Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationAn avetheropod that measured between 3.5 and 4 meters (11 and 13 ft) in length, with a weight of around 150 kilograms (330 pounds). However, some estimates put its weight as high as 400 kilograms (880 pounds), as very little is known about it. Kaijiangosaurus K. lini Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA megalosaurid, early tetanuran, early averostran or basal carnosaur. Leshansaurus L. qianweiensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao formationA megalosaurid . Sinocoelurus S. fragilis Mamenchisaurus assemblage, Upper Shaximiao FormationA theropod of uncertain affinities Sinraptor S. hepingensis Mamenchisaurus assemblage, Upper Shaximiao FormationA metriacanthosaurid, formally named Yangchuanosaurus hepingensis . Szechuanosaurus S. campi Mamenchisaurus assemblage, Upper Shaximiao FormationA metriacanthosaurid. Remains from the Shaximiao Formation might represent an indeterminate theropod. Xuanhanosaurus X. qilixiaensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA metriacanthosaurid. Yangchuanosaurus Y. shangyouensis Y. magnus Mamenchisaurus assemblage, Upper Shaximiao formationA metriacanthosaurid Y. zigongensis Szechuanosaurus zigongensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationZDM 9011 (holotype ), a partial postcranial skeleton; ZDM 9012, a left maxilla ; ZDM 9013, two teeth and ZDM 9014, a right hind limb.
Turtles Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Chengyuchelys C. baenoides Shangshaximiao Formation A xinjiangchelyid . C. dashanpuensis Shangshaximiao Formation A xinjiangchelyid. C. latimarginalis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA xinjiangchelyid. Sichuanchelys S. chowi Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA sichuanchelyid . Tienfuchelys T. zigongensis Shangshaximiao Formation A xinjiangchelyid. Sinaspideretes S. wimani Shangshaximiao Formation A trionychian , possibly the oldest known cryptodire
Therapsids Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Images Bienotheroides B. zigongensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA tritylodontid . Polistodon P. chuannanensis Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationA tritylodontid. Tritylodontidae gen. et sp. indet. Shunosaurus -Omeisaurus assemblage, Lower Shaximiao FormationAn indeterminate tritylodontid.
Plants Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Genus Species Stratigraphic position Material Notes Brachyoxylon B. qijiangense Masangyan village Leaves Family unknown.
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