Tanysauria (meaning "long lizards") is an extinct clade of long-necked archosauromorph reptiles from the Triassic period of Europe, Asia, North America, and South America. It comprises the families Tanystropheidae and Trachelosauridae (found by the authors to be coextensive with Dinocephalosauridae, and a senior synonym of the latter). Within the Archosauromorpha, Tanysauria represents the sister group to the Crocopoda. It contains many of the species formerly placed in "Protorosauria", which is now thought to be a paraphyletic group.

Definition

The clade Tanysauria was described by Stephan Spiekman and co-authors in 2024. Its name comes from the Greek τᾰνυ [tany] meaning "long" and σαῦρος [sauros] meaning "lizard," which refers to the long necks and, in some cases, bodies of tanysaurians.

According to the phylogenetic definition given by Spiekman et al., the clade Tanysauria includes all taxa that are more closely related to Tanystropheus longobardicus, Dinocephalosaurus orientalis, and Trachelosaurus fischeri than to Protorosaurus speneri, Prolacerta broomi, Mesosuchus browni, Azendohsaurus madagaskarensis, or Proterosuchus fergusi.

Classification

In their 2024 redescription of Trachelosaurus, Spiekman and colleagues named Tanysauria as a new clade containing the Tanystropheidae and Trachelosauridae (a senior synonym of Dinocephalosauridae). This clade is the sister taxon to the Crocopoda. The results of their phylogenetic analyses are shown in the cladogram below:

ArchosauromorphaProtorosaurusAenigmastropheus CrocopodaTanysauria Jesairosaurus Trachelosauridae Austronaga Pectodens Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus Tanystropheidae Augustaburiania Luxisaurus Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
ProtorosaurusAenigmastropheus
Protorosaurus
Aenigmastropheus
CrocopodaTanysauria Jesairosaurus Trachelosauridae Austronaga Pectodens Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus Tanystropheidae Augustaburiania Luxisaurus Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Crocopoda
TanysauriaJesairosaurus Trachelosauridae Austronaga Pectodens Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus Tanystropheidae Augustaburiania Luxisaurus Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Jesairosaurus
Trachelosauridae Austronaga Pectodens Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus Tanystropheidae Augustaburiania Luxisaurus Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
TrachelosauridaeAustronaga Pectodens Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus
Austronaga Pectodens
Austronaga
Pectodens
Fuyuansaurus Trachelosaurus Gracilicollum Dinocephalosaurus
Fuyuansaurus
Trachelosaurus Gracilicollum Dinocephalosaurus
Trachelosaurus
Gracilicollum Dinocephalosaurus
Gracilicollum
Dinocephalosaurus
TanystropheidaeAugustaburiania Luxisaurus Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Augustaburiania
Luxisaurus
Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Macrocnemus bassanii Macrocnemus fuyuanensis Macrocnemus obristi
Macrocnemus bassanii
Macrocnemus fuyuanensis Macrocnemus obristi
Macrocnemus fuyuanensis
Macrocnemus obristi
UFRGS-PV-492-T Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
UFRGS-PV-492-T
Amotosaurus Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Amotosaurus
Langobardisaurus Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Langobardisaurus
Raibliania Tanystropheus spp. Sclerostropheus Ozimek Tanytrachelos
Raibliania Tanystropheus spp.
Raibliania
Tanystropheus spp.
Sclerostropheus Ozimek Tanytrachelos
Sclerostropheus
Ozimek Tanytrachelos
Ozimek
Tanytrachelos