Anningasaura is an extinct genus of basal plesiosaur. It is known from a single type species, A. lymense, discovered in Early Jurassic rocks of Lyme Regis in the United Kingdom.

Discovery

1896 lithograph of the skull of specimen NHMUK OR 49202

Anningasaura is known only from the holotype specimen NHMUK OR 49202, which consists of a complete skull, palate, and mandible, with eight associated cervical vertebrae, including the atlas-axis complex. The partial skeleton came from a juvenile plesiosaur. It was originally referred to "Plesiosaurus" macrocephalus by Charles William Andrews in 1896, which is otherwise known only from the very young type specimen NHMUK PV R1336. NHMUK OR 49202 was collected at Lyme Regis, of Dorset, from the Hettangian to early Sinemurian-aged beds of the Lower Lias Group.

Description

NHMUK OR 49202 possesses plesiomorphic characters, including premaxillae that do not separate the frontals on the midline, narrow cranioquadrate passages and the lack of a constricting groove around the occipital condyle. It also shows several autapomorphies not observed in other plesiosaurians. Its posteromedial processes of the premaxillae (or possible anterior portion of the frontal) forming a dorsoventrally thick, mediolaterally expanded platform and its cultriform process of the parasphenoid is wider mediolaterally than the combined posterior interpterygoid vacuities. It also has two closely spaced foramina in the lateral surface of the exoccipital. Additional autapomorphies are the presence of supplementary foramen penetrating the parietal sagittal crest, the absence of a pterygoid-vomerine contact and the absence of a contact between the pterygoids in palatal aspect.

Phylogeny

A phylogenetic analysis performed by Benson et al. (2012) found it to be a basal, non-neoplesiosaurian, plesiosaur. The cladogram below shows Anningasaura phylogenetic position among other plesiosaurs following Benson et al. (2012).

Pistosauria"Pistosaurus postcranium" Pistosaurus Yunguisaurus liae Augustasaurus hagdorni Plesiosauria Bobosaurus forojuliensis Anningasaura lymense Rhomaleosauridae Stratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus Neoplesiosauria Pliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
"Pistosaurus postcranium"
Pistosaurus
Yunguisaurus liae
Augustasaurus hagdorni Plesiosauria Bobosaurus forojuliensis Anningasaura lymense Rhomaleosauridae Stratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus Neoplesiosauria Pliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Augustasaurus hagdorni
PlesiosauriaBobosaurus forojuliensis Anningasaura lymense Rhomaleosauridae Stratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus Neoplesiosauria Pliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Bobosaurus forojuliensis
Anningasaura lymense Rhomaleosauridae Stratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus Neoplesiosauria Pliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Anningasaura lymense
Rhomaleosauridae Stratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus Neoplesiosauria Pliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
RhomaleosauridaeStratesaurus taylori Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Stratesaurus taylori
Macroplata tenuiceps Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Macroplata tenuiceps
Avalonnectes arturi Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Avalonnectes arturi
Eurycleidus arcuatus Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Eurycleidus arcuatus
Meyerasaurus victor Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Meyerasaurus victor
Maresaurus coccai Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Maresaurus coccai
Atychodracon megacephalus Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Atychodracon megacephalus
Archaeonectrus rostratus Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Archaeonectrus rostratus
Rhomaleosaurus cramptoni Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Rhomaleosaurus cramptoni
Rhomaleosaurus thorntoni Rhomaleosaurus zetlandicus
Rhomaleosaurus thorntoni
Rhomaleosaurus zetlandicus
NeoplesiosauriaPliosauridae Thalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes) Plesiosauroidea Eoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
PliosauridaeThalassiodracon hawkinsii Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes)
Thalassiodracon hawkinsii
Hauffiosaurus spp. Attenborosaurus conybeari advanced pliosaurids (Peloneustes)
Hauffiosaurus spp.
Attenborosaurus conybeari advanced pliosaurids (Peloneustes)
Attenborosaurus conybeari
advanced pliosaurids (Peloneustes)
PlesiosauroideaEoplesiosaurus antiquior Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Eoplesiosaurus antiquior
Plesiosaurus dolichodeirus Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Plesiosaurus dolichodeirus
Plesiopterys wildi Cryptoclidus eurymerus Microcleididae Eretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Plesiopterys wildi Cryptoclidus eurymerus
Plesiopterys wildi
Cryptoclidus eurymerus
MicrocleididaeEretmosaurus rugosus Westphaliasaurus simonsensii Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Eretmosaurus rugosus
Westphaliasaurus simonsensii
Seeleyosaurus guilelmiimperatoris Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Seeleyosaurus guilelmiimperatoris
Microcleidus tournemiensis Microcleidus brachypterygius Microcleidus homalospondylus
Microcleidus tournemiensis
Microcleidus brachypterygius Microcleidus homalospondylus
Microcleidus brachypterygius
Microcleidus homalospondylus

Etymology

Side view of a skull cast

Anningasaura was first described and named by Peggy Vincent and Roger B. J. Benson in 2012 and the type species is Anningasaura lymense. The generic name honors Mary Anning, a British fossil collector who became known around the world for finds she made in the Jurassic marine fossil beds at Lyme Regis in Dorset. The specific name is derived from the name of the Lyme Regis locality where the only known specimen was collected.

See also