The Campylobacterales are an order of Campylobacterota which make up the epsilon subdivision, together with the small family Nautiliaceae. They are Gram-negative. Most of the species are microaerophilic.

Molecular signatures

Comparative genomic analysis has led to the identification of 49 proteins which are uniquely found in virtually all species of the order Campylobacterales. Additionally, two conserved signature indels have been identified which, along with the proteins, serve as molecular markers for the order. The indels are a three-amino-acid insertion in the B protein of the Uvr ABC system, and a two-amino-acid deletion in phenylalanyl-tRNA synthetase.

Phylogeny

The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI).

16S rRNA based LTP_10_2024120 marker proteins based GTDB 10-RS226
Nitratiruptoraceae Nitratiruptor Nitrosophilus Hydrogenimonadaceae Hydrogenimonas Thioreductoraceae Thioreductor Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"] Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter AliarcobacterThiovulaceae Thiovulum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Nitratiruptoraceae Nitrosophilus alvini Nitrosophilus_A Nitratiruptor Hydrogenimonadaceae Hydrogenimonas Sulfurovaceae Nitratifractor Sulfurovum Helicobacteraceae Wolinella "Pseudohelicobacter" Helicobacter Arcobacteraceae "Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter Sulfurospirillaceae Sulfurospirillum UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
Nitratiruptoraceae Nitratiruptor Nitrosophilus Hydrogenimonadaceae Hydrogenimonas Thioreductoraceae Thioreductor Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"] Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Nitratiruptoraceae Nitratiruptor Nitrosophilus
NitratiruptoraceaeNitratiruptor Nitrosophilus
Nitratiruptor
Nitrosophilus
Hydrogenimonadaceae Hydrogenimonas Thioreductoraceae Thioreductor Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"] Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Hydrogenimonadaceae Hydrogenimonas
HydrogenimonadaceaeHydrogenimonas
Thioreductoraceae Thioreductor Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"] Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Thioreductoraceae Thioreductor
ThioreductoraceaeThioreductor
Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"] Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Helicobacteraceae Wolinella Helicobacter [incl. "Pseudohelicobacter"]
HelicobacteraceaeWolinella Helicobacter [incl. "Pseudohelicobacter"]
Wolinella
Helicobacter [incl. "Pseudohelicobacter"]
Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Sulfurovaceae Nitratifractor Sulfurovum Sulfurimonadaceae Sulfuricurvum Sulfurimonas
SulfurovaceaeNitratifractor Sulfurovum
Nitratifractor
Sulfurovum
SulfurimonadaceaeSulfuricurvum Sulfurimonas
Sulfuricurvum
Sulfurimonas
Sulfurospirillaceae Sulfurospirillum Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Sulfurospirillaceae Sulfurospirillum
SulfurospirillaceaeSulfurospirillum
Campylobacteraceae Campylobacter Arcobacteraceae Poseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
CampylobacteraceaeCampylobacter
ArcobacteraceaePoseidonibacter Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Poseidonibacter
Malaciobacter Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Malaciobacter
Arcobacter Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Arcobacter
Aliarcobacter species-group 2 Halarcobacter Aliarcobacter
Aliarcobacter species-group 2
Halarcobacter Aliarcobacter
Halarcobacter
Aliarcobacter
Thiovulaceae Thiovulum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Nitratiruptoraceae Nitrosophilus alvini Nitrosophilus_A Nitratiruptor Hydrogenimonadaceae Hydrogenimonas Sulfurovaceae Nitratifractor Sulfurovum Helicobacteraceae Wolinella "Pseudohelicobacter" Helicobacter Arcobacteraceae "Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter Sulfurospirillaceae Sulfurospirillum UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
Thiovulaceae Thiovulum Sulfurimonadaceae Sulfuricurvum Sulfurimonas Nitratiruptoraceae Nitrosophilus alvini Nitrosophilus_A Nitratiruptor Hydrogenimonadaceae Hydrogenimonas Sulfurovaceae Nitratifractor Sulfurovum
Thiovulaceae Thiovulum Sulfurimonadaceae Sulfuricurvum Sulfurimonas
ThiovulaceaeThiovulum
SulfurimonadaceaeSulfuricurvum Sulfurimonas
Sulfuricurvum
Sulfurimonas
Nitratiruptoraceae Nitrosophilus alvini Nitrosophilus_A Nitratiruptor Hydrogenimonadaceae Hydrogenimonas Sulfurovaceae Nitratifractor Sulfurovum
Nitratiruptoraceae Nitrosophilus alvini Nitrosophilus_A Nitratiruptor
NitratiruptoraceaeNitrosophilus alvini Nitrosophilus_A Nitratiruptor
Nitrosophilus alvini
Nitrosophilus_A Nitratiruptor
Nitrosophilus_A
Nitratiruptor
Hydrogenimonadaceae Hydrogenimonas Sulfurovaceae Nitratifractor Sulfurovum
HydrogenimonadaceaeHydrogenimonas
SulfurovaceaeNitratifractor Sulfurovum
Nitratifractor
Sulfurovum
Helicobacteraceae Wolinella "Pseudohelicobacter" Helicobacter Arcobacteraceae "Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter Sulfurospirillaceae Sulfurospirillum UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
Helicobacteraceae Wolinella "Pseudohelicobacter" Helicobacter
HelicobacteraceaeWolinella "Pseudohelicobacter" Helicobacter
Wolinella "Pseudohelicobacter"
Wolinella
"Pseudohelicobacter"
Helicobacter
Arcobacteraceae "Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter Sulfurospirillaceae Sulfurospirillum UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
Arcobacteraceae "Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
Arcobacteraceae"Ca. Marinarcus" Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
"Ca. Marinarcus"
Malaciobacter Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
Malaciobacter
Arcobacter Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
Arcobacter
Halarcobacter Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
Halarcobacter
Arcobacter roscoffensis Poseidonibacter Malaciobacter pacificus Aliarcobacter
Arcobacter roscoffensis Poseidonibacter
Arcobacter roscoffensis
Poseidonibacter
Malaciobacter pacificus Aliarcobacter
Malaciobacter pacificus
Aliarcobacter
Sulfurospirillaceae Sulfurospirillum UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
Sulfurospirillaceae Sulfurospirillum
SulfurospirillaceaeSulfurospirillum
UBA1877 Sulfurospirillum_B Campylobacteraceae Campylobacter
UBA1877Sulfurospirillum_B
CampylobacteraceaeCampylobacter

Unassigned genera:

  • "Flexispira" Bryner 1987
  • "Candidatus Thioturbo" Muyzer et al. 2005
  • "Thiofractor" Makita et al. 2012
  • "Thiobarbaceae" Assié et al. 2020 "Candidatus Thiobarba" Assié et al. 2020

See also