The Cucaracha Formation (Tca) is a geologic formation in Panama. It preserves vertebrate and plant fossils dating back to the Neogene period; Early to Middle Miocene epochs (Hemingfordian). Fossils of the crocodylian Centenariosuchus, the turtle Rhinoclemmys panamaensis and the artiodactyl Paratoceras have been found in the formation.

The Cucaracha Formation is approximately 125 metres (410 ft) thick, and dated to 19 to 14 Ma.

Fossil content

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Mammals

Bats

Bats reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
AmericanycterisA. cyrtodonCentenario Bridge & Hodges HillPartial jaw & right premolarA leaf-nosed bat, first reported as a new "large insectivorous phyllostomine"

Carnivorans

Carnivorans reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
BassaricyonoidesB. sp.Gaillard CutA procyonid
Daphoenodoncf. D. sp.Gaillard CutA bear dog
Hemicyonidae indet.IndeterminateGaillard CutRadius & ulna fragmentA bear
TomarctusT. brevirostrisGaillard CutJaw & teethA borophagine dog

Rodents

Rodents reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
NototamiasN. sp.Gaillard CutA ground squirrel
PetauristodonP. sp.Gaillard CutA flying squirrel
ProheteromysP. sp.Gaillard CutA heteromyid
TexomysT. stewartiPedro Miguel Lock & Gaillard CutTeethA jimomyid

Ungulates

Ungulates reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
AnchitheriumA. clarenceiGaillard Cut & Centenario BridgeMandible fragment, upper cheek tooth & maxillaAn equid
ArchaeohippusA. sp.Gaillard CutCalcaneus & toothAn equid
CynorcaC. occidentaleGaillard Cut, Centenario Bridge & Hodges HillJaw elements with teethSpecies reassigned to the genus Tedfordhyus
FloridacerasF. whiteiGaillard CutJaws, teeth & femur fragmentA rhinoceros
Floridatraguluscf. F. nanusGaillard CutA camelid
Machaeromeryxcf. M. sp.Gaillard CutA moschid
MenocerasM. barbouriGaillard CutTeeth & limb elementsA rhinoceros
MerycochoerusM. matthewiGaillard CutSkull elementsAn oreodont
Parablastomeryxcf. P. sp.Gaillard CutA moschid
ParatocerasP. coatesiGaillard Cut, Centenario Bridge, Hodges Hill & Cartagena HillSkull elements, teeth & limb fragmentsA protoceratid, first reported as P. wardi
P. wardiGaillard CutSkull elements, teeth & limb fragmentsRemains reassigned to P. coatesi
TedfordhyusT. occidentalisGaillard Cut, Centenario Bridge & Hodges HillJaw elements with teethA peccary, originally named Cynorca occidentale

Reptiles

Birds

Birds reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
Accipitridae indet.IndeterminateWest side of the Panama CanalTalonA large eagle, talon resembles those of the crowned eagle and martial eagle

Crocodilians

Crocodilians reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
CentenariosuchusC. gilmoreiHodges HillSkull & jawA caiman
cf. C. gilmoreiCentenario BridgeRight angularA caiman
DadagavialisD. gunaiEast margin of the Panama CanalPartial snoutA gavialid
cf. D. gunaiCentenario BridgePartial mandibleA gavialid
Eusuchia indet.IndeterminatePanama Canal ZoneHeadless skeletonOsteoderms comparable to those of gavialids such as Gavialosuchus, but crocodilian osteoderms are non-diagnostic

Squamates

Squamates reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
BoaB. cf. B. constrictorCartagena Hill & Hodges HillIncomplete vertebraeA boid snake

Turtles

Turtles reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
Podocnemididae indet.Incertae sedisCentenario BridgeRight epiplastronA side-necked turtle
RhinoclemmysR. panamaensisCentenario BridgeShell elementsA geoemydid turtle
R. sp.Centenario Bridge & Culebra ReachIsolated bonesA geoemydid turtle
StaurotypusS. moschusCentenario BridgePeripheral part of shellA kinosternid turtle
Testudinidae indet.Incertae sedisCulebra ReachShell & limb elementsA tortoise
Trionychidae indet.Incertae sedisCentenario BridgeRight costalA softshell turtle

Plants

Plants reported from the Cucaracha Formation
GenusSpeciesPresenceMaterialNotesImages
AmpelorhizaA. heteroxylonCulebra CutRootsA member of Paullinieae
AntrocaryonA. panamaensisLirio EastEndocarpsA member of Anacardiaceae
DracontomelonD. montesiiLirio EastEndocarpsA member of Anacardiaceae
GuazumaoxylonG. miocenicaHodges HillWoodA member of Malvaceae, similar to Guazuma
MammeaM. paramericanaNorthern Lirio EastStemA member of Calophyllaceae
OreomunneaO. grahamiiLirio EastNutsA member of Juglandaceae
PanascleroticoxylonP. crystallosaHodges HillWoodA member of Malpighiales
ParinariP. panamensisLirio East, Gold Hill & Empire sitesEndocarpsA member of Chrysobalanaceae
ParinarioxylonP. panamensisLirio EastWoodA member of Chrysobalanaceae
PeriplanetoxylonP. panamenseHodges HillWoodA member of Malvaceae
PrioriaP. canalensisHodges HillWoodA legume
P. hodgesiiHodges HillWoodA legume
RoureaR. blattaNorthern Lirio East siteWoodA liana belonging to the Connaraceae
SpondiasS. rothwelliiLirio EastEndocarpsA member of Anacardiaceae

See also

Bibliography

  • Kirby, Michael Xavier; Jones, Douglas S.; MacFadden, Bruce J. (2008), "Lower Miocene Stratigraphy along the Panama Canal and Its Bearing on the Central American Peninsula", PLoS ONE, 3 (7): 1–14, Bibcode:, doi:, PMC , PMID
  • Stewart, R.H.; Stewart, J.L.; Woodring, W.P. (1980), (PDF), USGS, p. 1
  • Woodring, W.P (1957), (PDF), USGS, pp. 1–186