Ichthyosauridae is an extinct family of thunnosaur ichthyosaurs from the latest Triassic and Early Jurassic (Rhaetian to Pliensbachian stages) of Europe, and possibly also from the middle Early Cretaceous (Hauterivian or Barremian stage) of Iraq. Named by Charles Lucien Bonaparte, in 1841, it is usually thought to contain a single genus, Ichthyosaurus, which is known from several species from the Early Jurassic. In 2013, Fischer et al. named and described Malawania anachronus from the middle Early Cretaceous of Iraq. It was found to share several synapomorphies with the type species of this family, Ichthyosaurus communis, and a large phylogenetic analysis recovered these species as sister taxa. Despite its geologically younger age, M. anachronus was also assigned to Ichthyosauridae.

Phylogeny

Fischer et al. (2013) applied the name Ichthyosauridae Bonaparte, 1841 for the clade that contains Malawania anachronus and Ichthyosaurus communis. This clade was recovered in the phylogenetic analysis that was based on the largest currently available data matrix of parvipelvian species. Most other phylogenetic analyses, that were based on smaller matrices, also recovered this clade or found M. anachronus to be in a more basal position. Ichthyosauridae was recovered as the sister taxon of Baracromia, a clade first defined by Fischer et al. (2013) that includes all other thunnosaurs. The cladogram below follows the large phylogenetic analysis of Fischer et al. (2013).

Life reconstruction of Ichthyosaurus anningae

Cladogram

Mikadocephalus gracilirostris Parvipelvia Hudsonelpidia brevirostris Macgowania janiceps Neoichthyosauria Temnodontosaurus Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini Suevoleviathan disinteger Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Mikadocephalus gracilirostris
ParvipelviaHudsonelpidia brevirostris Macgowania janiceps Neoichthyosauria Temnodontosaurus Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini Suevoleviathan disinteger Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Hudsonelpidia brevirostris
Macgowania janiceps Neoichthyosauria Temnodontosaurus Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini Suevoleviathan disinteger Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Macgowania janiceps
NeoichthyosauriaTemnodontosaurus Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini Suevoleviathan disinteger Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Temnodontosaurus Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini
Temnodontosaurus
Leptonectes tenuirostris Eurhinosaurus longirostris Excalibosaurus costini
Leptonectes tenuirostris
Eurhinosaurus longirostris Excalibosaurus costini
Eurhinosaurus longirostris
Excalibosaurus costini
Suevoleviathan disinteger Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Suevoleviathan disinteger
Hauffiopteryx typicus Thunnosauria Ichthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Hauffiopteryx typicus
ThunnosauriaIchthyosauridae Ichthyosaurus communis Malawania anachronus Baracromia Stenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
IchthyosauridaeIchthyosaurus communis Malawania anachronus
Ichthyosaurus communis
Malawania anachronus
BaracromiaStenopterygius quadriscissus Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Stenopterygius quadriscissus
Chacaicosaurus cayi Ophthalmosauridae Arthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Chacaicosaurus cayi
OphthalmosauridaeArthropterygius chrisorum Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Arthropterygius chrisorum
Ophthalmosaurinae Mollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus Platypterygiinae Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
OphthalmosaurinaeMollesaurus periallus Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus
Mollesaurus periallus
Ophthalmosaurus icenicus (type species) Baptanodon natans ("O." natans) Acamptonectes densus
Ophthalmosaurus icenicus (type species)
Baptanodon natans ("O." natans) Acamptonectes densus
Baptanodon natans ("O." natans)
Acamptonectes densus
PlatypterygiinaeBrachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus "Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Brachypterygius extremus Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus
Brachypterygius extremus
Maiaspondylus lindoei Aegirosaurus leptospondylus Sveltonectes insolitus
Maiaspondylus lindoei
Aegirosaurus leptospondylus Sveltonectes insolitus
Aegirosaurus leptospondylus
Sveltonectes insolitus
"Platypterygius" hercynicus Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
"Platypterygius" hercynicus
Caypullisaurus bonapartei Athabascasaurus bitumineus "Platypterygius" australis
Caypullisaurus bonapartei
Athabascasaurus bitumineus "Platypterygius" australis
Athabascasaurus bitumineus
"Platypterygius" australis