Linyphiidae, spiders commonly known as sheet weavers (from the shape of their webs), or money spiders (in the United Kingdom, Ireland, Australia, New Zealand, and Portugal) is a family of very small spiders comprising around 5,000 described species in around 650 genera worldwide. This makes Linyphiidae the second largest family of spiders after the Salticidae.

The family is poorly understood due to their small body size and wide distribution; new genera and species are still being discovered throughout the world. The newest such genus is Himalafurca from Nepal, formally described in April 2021 by Tanasevitch. Since it is so difficult to identify such tiny spiders, there are regular changes in taxonomy as species are combined or divided.

Money spiders are known for drifting through the air via a technique termed "ballooning". Females of winter-active sheet weavers, such as Tenuiphantes cristatus and Bolephthyphantes index, are known for building sheet webs in depressions on the surface of snow.

Within the agriculture industry, money spiders are regarded as biological control agents against pest species like aphids and springtails.

Description

In Linyphiidae, the clypeus is normally over twice as high as the diameter of the anterior median eyes. The chelicerae have lateral stridulating ridges and lack lateral condyles.

The legs are long and thin, and bear macrosetae. The abdomen is usually oval or elongated.

Distribution

Sheet weaver's web

Spiders of this family occur nearly worldwide. In Norway many species have been found walking on snow at temperatures of down to −7 °C.

While these spiders are light enough to utilize ballooning for travel, they are limited by the physics of an often turbulent atmosphere and microclimate. For this reason ballooning spiders have little control over where they land, leading to a high mortality rate for the practice and its predominant usage by spiderlings and juveniles. The travel of money spiders by ballooning likely contributes to their vast distribution and speciation.

Predators and prey

Taxonomy

The Pimoidae are the sister group to the Linyphiidae.

Sheet weaver's web in morning dew

There are six subfamilies, of which Linyphiinae (the sheetweb spiders), Erigoninae (the dwarf spiders), and Micronetinae, contain the majority of described species.

Many species have been described in monotypic genera, especially in the Erigoninae, which probably reflects the scientific techniques traditionally used in this family. Common genera include Neriene, Lepthyphantes, Erigone, Eperigone, Bathyphantes, Troglohyphantes, Tennesseellum and many others. These are among the most abundant spiders in the temperate regions, although many are also found in the tropics. The generally larger bodied members of the subfamily Linyphiinae are commonly found in classic "bowl and doily" webs or filmy domes. The usually tiny members of the Erigoninae are builders of tiny sheet webs. These tiny spiders (usually 3 mm or less) commonly balloon even as adults and may be very numerous in a given area on one day, only to disappear the next.

Males in the subfamily Erigoninae typically have modified cephalothoraxes. These modifications are diagnostic for a given taxon, being genus or species-specific. These come in an impressive array of forms including, but not limited to, grooves, tubercles, projections, bumps, lobes, and spines. Occasionally, the projections may be decorated with tufts of hair or even bear eyes.

The following are select examples of species in which males possess rather remarkable modifications. Walckenaeria acuminata has its eyes placed on a tall, thin spire whose height exceeds the length of the cephalothorax. Grammonota gigas has a transverse row of four longitudinal lobes behind the eyes. Gnathonargus unicorn has a long, slender, upward-pointing clypeal projection resembling a unicorn horn. Hypselistes florens has a cephalic lobe shaped like an hourglass when viewed from the front. Perregrinus deformis has a short, downcurved clypeal projection resembling a human nose. Praestigia kulczynskii has its anterior median eyes placed ventrally at the end of a long, thick projection issuing from the clypeus. The genera Coreorgonal and Spirembolus have their cephalic regions deeply divided into two pronounced lobes. Eskovia exarmata has a cephalothorax shaped like a trapezoid when viewed laterally. Horcotes quadricristatus has a single, sharp tooth sticking up between the anterior and posterior eyes.

Similarly, the pedipalps of males range from simple to complex in their design, with some possessing striking features and arrangements of palpal sclerites that are unique for a given genus and/or species.

A few spiders in this family include:

Genera

As of October 2025[update], this family includes 641 genera and 4962 species.

Complete genus list
Abacoproeces Simon, 1884 – Alps, Russia Aberdaria Holm, 1962 – Kenya Abiskoa Saaristo & Tanasevitch, 2000 – China, Poland, Russia, Scandinavia Absconditus Irfan, Zhang & Peng, 2022 – China Acanoides Sun, Marusik & Tu, 2014 – China Acanthoneta Eskov & Marusik, 1992 – China, Russia, North America Acartauchenius Simon, 1884 – Algeria, Morocco, Tunisia, Asia, Russia, Italy, Malta Acorigone Wunderlich, 2008 – Azores Acroterius Irfan, Bashir & Peng, 2021 – China Adelonetria Millidge, 1991 – Chile Afribactrus Wunderlich, 1995 – South Africa Afromynoglenes Merrett & Russell-Smith, 1996 – Ethiopia Afroneta Holm, 1968 – Cameroon, DR Congo, Eastern Africa Afrophantes Tanasevitch, 2025 – Cameroon, Ethiopia, Kenya, Tanzania Afrotrichona Tanasevitch, 2020 – Kenya Agnyphantes Hull, 1933 – Kazakhstan, China, Russia, Alaska, Canada Agyneta Hull, 1911 – Worldwide Agyphantes Saaristo & Marusik, 2004 – Russia Ainerigone Eskov, 1993 – Japan, Russia Alioranus Simon, 1926 – Asia, Cyprus, Greece Allomengea Strand, 1912 – Kyrgyzstan, Eastern Asia, Caucasus, Russia, North America Allotiso Tanasevitch, 1990 – Georgia, Turkey Amfractus Irfan, Zhang & Peng, 2022 – China Anacornia Chamberlin & Ivie, 1933 – United States Anguliphantes Saaristo & Tanasevitch, 1996 – Asia, Romania, Russia Anibontes Chamberlin, 1924 – United States Annapolis Millidge, 1984 – North America Anodoration Millidge, 1991 – Argentina, Brazil Anthrobia Tellkampf, 1844 – North America Antrohyphantes Dumitrescu, 1971 – Bulgaria, North Macedonia Aperturina Tanasevitch, 2014 – Malaysia, Thailand Aphileta Hull, 1920 – Kazakhstan, Russia Apobrata Miller, 2004 – Philippines Aprifrontalia Oi, 1960 – Eastern Asia, Russia Arachosinella Denis, 1958 – Afghanistan, Kazakhstan, Kyrgyzstan, Mongolia, Russia Araeoncus Simon, 1884 – Africa, Asia, Europe, North Africa. Introduced to New Zealand Archaraeoncus Tanasevitch, 1987 – Asia, Bulgaria, Ukraine, Russia Arcterigone Eskov & Marusik, 1994 – Russia, Alaska, Canada Arcuphantes Chamberlin & Ivie, 1943 – Eastern Asia, North America Ascetophantes Tanasevitch & Saaristo, 2006 – Nepal Asemostera Simon, 1898 – Central to South America Asiafroneta Tanasevitch, 2020 – Malaysia Asiagone Tanasevitch, 2014 – China, Laos, Thailand Asiceratinops Eskov, 1992 – Russia Asiophantes Eskov, 1993 – Russia Asperthorax Oi, 1960 – China, Japan, Russia Asthenargellus Caporiacco, 1949 – Kenya Asthenargoides Eskov, 1993 – Russia Asthenargus Simon & Fage, 1922 – Africa, Asia, Europe Atypena Simon, 1894 – Asia Auriculaiana Irfan, Zhang & Peng, 2023 – China Australolinyphia Wunderlich, 1976 – Australia Australophantes Tanasevitch, 2012 – Indonesia, Australia Bactrogyna Millidge, 1991 – Chile Baryphyma Simon, 1884 – Kazakhstan, Kyrgyzstan, China, Caucasus, Georgia, Europe, Britain, Western Baryphymula Eskov, 1992 – Japan Bathylinyphia Eskov, 1992 – Kazakhstan, China, Japan, Korea, Russia Bathyphantes Menge, 1866 – Angola, Congo, Asia, Europe, Russia, North America, Oceania, Argentina Batueta Locket, 1982 – Asia Bifurcia Saaristo, Tu & Li, 2006 – China, Russia Birgerius Saaristo, 1973 – Spain, France Bisetifer Tanasevitch, 1987 – Caucasus, Ukraine, Russia Bishopiana Eskov, 1988 – China, Russia Blestia Millidge, 1993 – United States Bolephthyphantes Strand, 1901 – Central Asia, Caucasus, Russia, France, Greenland Bolyphantes C. L. Koch, 1837 – Asia, Europe, Western Mediterranean Bordea Bosmans, 1995 – Portugal, Spain, France Boreomaro Tanasevitch, 2022 – Russia Brachycerasphora Denis, 1962 – Northern Africa, Israel Bureyanus Tanasevitch, 2023 – Russia Bursellia Holm, 1962 – Cameroon, Congo, Eastern Africa Caenonetria Millidge & Russell-Smith, 1992 – Brunei Callitrichia Fage, 1936 – Africa, China, Thailand, India Callosa Zhao & Li, 2017 – China Camafroneta Frick & Scharff, 2018 – Cameroon Cameroneta Bosmans & Jocqué, 1983 – Cameroon Canariellanum Wunderlich, 1987 – Canary Islands Canariphantes Wunderlich, 1992 – Ethiopia, Algeria, Morocco, Tunisia, Israel, Turkey, Europe Capsulia Saaristo, Tu & Li, 2006 – China Caracladus Simon, 1884 – China, Japan, Austria, Germany, Switzerland, Italy, France, Western Carorita Duffey & Merrett, 1963 – China, Ukraine, Russia, Central, Western Cassafroneta Blest, 1979 – New Zealand Catacercus Millidge, 1985 – Chile Catonetria Millidge & Ashmole, 1994 – Ascension Islands Caucasopisthes Tanasevitch, 1990 – Caucasus Cautinella Millidge, 1985 – Chile Caviphantes Oi, 1960 – Asia, Bulgaria, Romania, Ukraine, Russia, North America Centromerita Dahl, 1912 – Caucasus, Turkey. Introduced to North America Centromerus Dahl, 1886 – North Africa, Asia, Europe, North America Centrophantes Miller & Polenec, 1975 – Austria, Slovenia Ceraticelus Simon, 1884 – Germany, Poland, Russia, Finland, Netherlands, North America Ceratinella Emerton, 1882 – China, Japan, Korea, Caucasus, Iran, Turkey, Russia, Italy, North America, Australia Ceratinops Banks, 1905 – North America Ceratinopsidis Bishop & Crosby, 1930 – North America Ceratinopsis Emerton, 1882 – Africa, Asia, Canary Islands, Madeira, Guatemala, North America Ceratocyba Holm, 1962 – Kenya Cheniseo Bishop & Crosby, 1935 – North America Chenisides Denis, 1962 – Congo, Kenya Cherserigone Denis, 1954 – Algeria Chiangmaia Millidge, 1995 – Thailand Chthiononetes Millidge, 1993 – Australia Cinetata Wunderlich, 1995 – Georgia Cirrosus Zhao & Li, 2014 – China Claviphantes Tanasevitch & Saaristo, 2006 – Nepal Cnephalocotes Simon, 1884 – Korea, Ukraine, Russia, Canada, Hawaii Collinsia O. Pickard-Cambridge, 1913 – Kyrgyzstan, China, Japan, Korea, Caucasus, Iran, Austria, Russia, Spain, France, North America, Spitsbergen Coloncus Chamberlin, 1949 – North America Comorella Jocqué, 1985 – Comoros Concavocephalus Eskov, 1989 – Russia Conglin Zhao & Li, 2014 – China Connithorax Eskov, 1993 – Russia Coreorgonal Bishop & Crosby, 1935 – North America Cornicephalus Saaristo & Wunderlich, 1995 – China Cornitibia Lin, Lopardo & Uhl, 2022 – Nepal Cresmatoneta Simon, 1929 – Japan, Korea, India, Caucasus, Turkey, Russia Crispiphantes Tanasevitch, 1992 – China, Korea, Russia Cristatus Irfan, Zhang & Peng, 2022 – China Crosbyarachne Charitonov, 1937 – Turkey, Europe Crosbylonia Eskov, 1988 – Russia Cryptolinyphia Millidge, 1991 – Colombia Ctenophysis Millidge, 1985 – Chile Curtimeticus Zhao & Li, 2014 – China Cyphonetria Millidge, 1995 – Thailand Dactylopisthes Simon, 1884 – Asia, Europe, Alaska, Canada Dactylopisthoides Eskov, 1990 – Russia Decipiphantes Saaristo & Tanasevitch, 1996 – Kazakhstan, Mongolia, Belarus, Russia, Scandinavia Deelemania Jocqué & Bosmans, 1983 – Cameroon, Gabon, Malawi, Ivory Coast Dendronetria Millidge & Russell-Smith, 1992 – Brunei Denisiphantes Tu, Li & Rollard, 2005 – China Diastanillus Simon, 1926 – Austria, Norway, France Dicornua Oi, 1960 – Japan Dicristatus Irfan, Wang & Zhang, 2023 – China Dicymbium Menge, 1868 – Asia, Russia, North America Didectoprocnemis Denis, 1950 – North Africa, Romania, Greece, Portugal, France Diechomma Millidge, 1991 – Colombia Diplocentria Hull, 1911 – China, Mongolia, Russia, Europe, North America Diplocephaloides Oi, 1960 – China, Japan, Korea Diplocephalus Bertkau, 1883 – North Africa, Asia, Europe to Russia, North America, Greenland, Falkland Is Diploplecta Millidge, 1988 – New Zealand Diplostyla Emerton, 1882 – Korea, Caucasus, Iran, Turkey, Russia. Introduced to St. Helena Diplothyron Millidge, 1991 – Central America, Mexico, Colombia, Venezuela Disembolus Chamberlin & Ivie, 1933 – North America Dismodicus Simon, 1884 – Russia, France, North America Doenitzius Oi, 1960 – China, Japan, Korea, Russia Dolabritor Millidge, 1991 – Colombia Donacochara Simon, 1884 – Angola, Europe to Central Asia Drapetisca Menge, 1866 – China, Japan, Caucasus, Russia, North America Drepanotylus Holm, 1945 – China, Mongolia, Bulgaria, Russia, Scandinavia Dresconella Denis, 1950 – France Dubiaranea Mello-Leitão, 1943 – Malaysia, South America Dumoga Millidge & Russell-Smith, 1992 – Indonesia Dunedinia Millidge, 1988 – Australia, New Zealand Eborilaira Eskov, 1989 – Russia Eldonnia Tanasevitch, 2008 – Japan, Korea, Russia Emenista Simon, 1894 – India Emertongone Lin, Lopardo & Uhl, 2022 – North America Enguterothrix Denis, 1962 – Congo, Southeast Asia Entelecara Simon, 1884 – Algeria, Asia, Europe, North America Eordea Simon, 1899 – Indonesia Epibellowia Tanasevitch, 1996 – Japan, Russia Epiceraticelus Crosby & Bishop, 1931 – United States Epigyphantes Saaristo & Tanasevitch, 2004 – Russia Epigytholus Tanasevitch, 1996 – Mongolia, Russia Episolder Tanasevitch, 1996 – Russia Epiwubana Millidge, 1991 – Chile Eridantes Crosby & Bishop, 1933 – North America Erigokhabarum Tanasevitch, 2022 – Russia Erigomicronus Tanasevitch, 2018 – China, Japan, Korea, Russia Erigone Audouin, 1826 – Worldwide Erigonella Dahl, 1901 – Mongolia, Alps, Bulgaria, Russia, France, North Carpathians Erigonoploides Eskov, 1989 – Russia Erigonoplus Simon, 1884 – Morocco, Kazakhstan, Turkmenistan, Western Asia, Europe Erigonops Scharff, 1990 – South Africa Erigophantes Wunderlich, 1995 – Borneo Eskovia Marusik & Saaristo, 1999 – Mongolia, Russia, Canada Eskovina Kocak & Kemal, 2006 – China, Korea, Russia Esophyllas Prentice & Redak, 2012 – United States Estrandia Blauvelt, 1936 – China, Japan, Russia Eulaira Chamberlin & Ivie, 1933 – North America Eurymorion Millidge, 1993 – Bolivia, Brazil Evansia O. Pickard-Cambridge, 1901 – Europe, Japan? Fageiella Kratochvíl, 1934 – Southern Europe Falklandoglenes Usher, 1983 – Falkland Islands Fissiscapus Millidge, 1991 – Colombia, Ecuador Fistulaphantes Tanasevitch & Saaristo, 2006 – Nepal Flagelliphantes Saaristo & Tanasevitch, 1996 – China, Russia Floricomus Crosby & Bishop, 1925 – North America Florinda O. Pickard-Cambridge, 1896 – Mexico, United States Floronia Simon, 1887 – China, Japan, Korea, Russia, Ecuador Formiphantes Saaristo & Tanasevitch, 1996 – Europe Frederickus Paquin, Dupérré, Buckle & Crawford, 2008 – North America Frontella Kulczyński, 1908 – Russia Frontinella F. O. Pickard-Cambridge, 1902 – China, Hispaniola, El Salvador, Mexico, United States Frontinellina van Helsdingen, 1969 – South Africa, North Africa, Asia, Madeira, Russia Frontiphantes Wunderlich, 1987 – Madeira Fusciphantes Oi, 1960 – Japan Gibbafroneta Merrett, 2004 – Congo Gibothorax Eskov, 1989 – Russia Gigapassus Miller, 2007 – Argentina Gladiata Zhao & Li, 2014 – China Glebala Zhao & Li, 2014 – China Glomerosus Zhao & Li, 2014 – China Glyphesis Simon, 1926 – Japan, Europe, North America Gnathonargus Bishop & Crosby, 1935 – United States Gnathonarium Karsch, 1881 – Asia, Russia, Alaska, Canada, North Africa Gnathonaroides Bishop & Crosby, 1938 – North America Gonatium Menge, 1868 – Kenya, Algeria, Morocco, Asia, Europe, North America Gonatoraphis Millidge, 1991 – Colombia Goneatara Bishop & Crosby, 1935 – North America Gongylidiellum Simon, 1884 – Angola, Algeria, Morocco, Asia, Europe, United States, Argentina Gongylidioides Oi, 1960 – Asia, Russia Gongylidium Menge, 1868 – Kazakhstan, China, Turkey, Russia, Italy Gracilentus Irfan, Zhang & Peng, 2022 – China Grammonota Emerton, 1882 – North America, Colombia Gravipalpus Millidge, 1991 – Argentina, Brazil, Peru Habreuresis Millidge, 1991 – Chile Halorates Hull, 1911 – Kazakhstan, China, Nepal, Pakistan, North America, Central, Western Haplinis Simon, 1894 – Australia, New Zealand Haplomaro Miller, 1970 – Angola Helophora Menge, 1866 – Kyrgyzstan, China, Caucasus, Russia, North America Helsdingenia Saaristo & Tanasevitch, 2003 – Cameroon, Comoros, Madagascar, Nigeria, St. Helena, Indonesia, Nepal, Sri Lanka Herbiphantes Tanasevitch, 1992 – China, Japan, Korea, Russia Heterolinyphia Wunderlich, 1973 – Bhutan, India, Nepal Heterotrichoncus Wunderlich, 1970 – Europe Hilaira Simon, 1884 – China, Japan, Mongolia, Nepal, Europe, Russia, North America Himalafurca Tanasevitch, 2021 – Nepal Himalaphantes Tanasevitch, 1992 – China, Japan, India, Nepal, Russia Holma Locket, 1974 – Angola Holmelgonia Jocqué & Scharff, 2007 – Africa Holminaria Eskov, 1991 – China, Mongolia, Russia Horcotes Crosby & Bishop, 1933 – Russia, Scandinavia, North America Houshenzinus Tanasevitch, 2006 – China Hubertella Platnick, 1989 – Nepal Hybauchenidium Holm, 1973 – Mongolia, Russia, Finland, Sweden, North America Hybocoptus Simon, 1884 – Algeria, Morocco, France Hylyphantes Simon, 1884 – Asia, Russia Hyperafroneta Blest, 1979 – New Zealand Hypomma Dahl, 1886 – Equatorial Guinea, Kazakhstan, Kyrgyzstan, China, Japan, Turkey, Russia, North Macedonia, North America, New Caledonia Hypselistes Simon, 1894 – Kazakhstan, Eastern Asia, Russia, North America Hypselocara Millidge, 1991 – Venezuela Ibadana Locket & Russell-Smith, 1980 – Cameroon, Nigeria Iberoneta Deeleman-Reinhold, 1984 – Spain Icariella Brignoli, 1979 – Greece Idionella Banks, 1893 – North America Improphantes Saaristo & Tanasevitch, 1996 – Kenya, Asia, Europe, North Africa Incestophantes Tanasevitch, 1992 – Kazakhstan, China, Mongolia, Georgia, Alps, Russia, Scandinavia, North America Indophantes Saaristo & Tanasevitch, 2003 – China, Indonesia, Malaysia, India, Nepal, Pakistan Intecymbium Miller, 2007 – Argentina, Chile Ipa Saaristo, 2007 – Kazakhstan, Turkmenistan, Mongolia, Caucasus, Turkey, Europe Ipaoides Tanasevitch, 2008 – China Islandiana Brændegaard, 1932 – Russia, Iceland, North America Ivielum Eskov, 1988 – Mongolia, Russia, Canada Jacksonella Millidge, 1951 – Korea, Cyprus, Greece, central Europe, Northern, western Jalapyphantes Gertsch & Davis, 1946 – Dominican Republic, Mexico, Colombia Janetschekia Schenkel, 1939 – Austria, Germany, Switzerland, Albania, Italy, France Javagone Tanasevitch, 2020 – Indonesia Javanaria Tanasevitch, 2020 – Indonesia Javanyphia Tanasevitch, 2020 – Indonesia Jilinus Lin, Lopardo & Uhl, 2022 – China, Korea, Russia Johorea Locket, 1982 – Malaysia, Singapore Juanfernandezia Koçak & Kemal, 2008 – Chile Kaestneria Wiehle, 1956 – China, Japan, Mongolia, Indonesia, Czechia, Poland, Slovakia, Romania, Ukraine, Russia, North America Kagurargus Ono, 2007 – Japan Kalimagone Tanasevitch, 2017 – Malaysia Karita Tanasevitch, 2007 – Germany, Poland, Russia, Ireland, Scandinavia, Belgium, Britain Kenocymbium Millidge & Russell-Smith, 1992 – Indonesia, Malaysia, Thailand Ketambea Millidge & Russell-Smith, 1992 – Asia, Russia Kikimora Eskov, 1988 – Russia, Finland, Norway Knischatiria Wunderlich, 1976 – Indonesia, Malaysia, Australia Koinothrix Jocqué, 1981 – Cape Verde Kolymocyba Eskov, 1989 – Russia Kratochviliella Miller, 1938 – Bulgaria, Greece, North Macedonia, France Labicymbium Millidge, 1991 – South America Labulla Simon, 1884 – Portugal, Spain, France Labullinyphia van Helsdingen, 1985 – China, Sri Lanka Labullula Strand, 1913 – Angola, Cameroon, Comoros Laetesia Simon, 1908 – Thailand, Australia, Vanuatu, New Zealand Lamellasia Tanasevitch, 2014 – Thailand Laminacauda Millidge, 1985 – Panama, South America Laminafroneta Merrett, 2004 – Cameroon, Congo, Eastern Africa Laogone Tanasevitch, 2014 – China, Laos Laperousea Dalmas, 1917 – Australia, New Zealand Lasiargus Kulczyński, 1894 – Kazakhstan, Kyrgyzstan, Mongolia, Russia Lenis Irfan, Zhang & Peng, 2025 – China Lepthyphantes Menge, 1866 – Africa, Asia, Europe, Puerto Rico, Brazil, Chile, North America, New Caledonia Leptorhoptrum Kulczyński, 1894 – Japan, Russia Leptothrix Menge, 1869 – Europe Lessertia Smith, 1908 – Algeria, Morocco, Turkey, Canary Islands, Madeira, Italy, Spain. Introduced to Canada, New Zealand Lessertinella Denis, 1947 – Alps, Slovakia, Romania, North Macedonia Lidia Saaristo & Marusik, 2004 – Kazakhstan, Kyrgyzstan Limoneta Bosmans & Jocqué, 1983 – Cameroon, Kenya, South Africa Linyphantes Chamberlin & Ivie, 1942 – North America Linyphia Latreille, 1804 – Algeria, Morocco, Asia, United States to South America, Australia, Cook Islands, Samoa Locketidium Jocqué, 1981 – Kenya, Malawi, Tanzania Locketiella Millidge & Russell-Smith, 1992 – Brunei, Indonesia Locketina Kocak & Kemal, 2006 – Brunei, Malaysia Lomaita Bryant, 1948 – Dominican Republic Lophomma Menge, 1868 – Russia, North America Lotusiphantes Chen & Yin, 2001 – China Lucrinus O. Pickard-Cambridge, 1904 – South Africa Lutosus Irfan, Zhang & Peng, 2022 – China Lygarina Simon, 1894 – South America Machadocara Miller, 1970 – Congo, Kenya, Zambia Macrargus Dahl, 1886 – China, Mongolia, Europe Maculoncus Wunderlich, 1995 – Taiwan, Caucasus, Israel, Cyprus, Greece Malkinola Miller, 2007 – Chile Mansuphantes Saaristo & Tanasevitch, 1996 – Caucasus, Turkey, Europe Maorineta Millidge, 1988 – Marshall Islands, New Zealand, Cook Islands Maro O. Pickard-Cambridge, 1907 – Japan, Mongolia, Russia, North America Martensinus Wunderlich, 1973 – Nepal Masikia Millidge, 1984 – Russia, North America Maso Simon, 1884 – Asia, Russia, Portugal, Alaska, United States Masoncus Chamberlin, 1949 – North America Masonetta Chamberlin & Ivie, 1939 – United States Mecopisthes Simon, 1926 – Algeria, Morocco, Turkmenistan, China, Japan, Caucasus, Israel, Europe Mecynargoides Eskov, 1988 – Mongolia, Russia Mecynargus Kulczyński, 1894 – Kazakhstan, Kyrgyzstan, China, Mongolia, Europe, Alaska, Canada, Greenland Mecynidis Simon, 1894 – Angola, Kenya, Tanzania, South Africa Megafroneta Blest, 1979 – New Zealand Megalepthyphantes Wunderlich, 1994 – Algeria, Morocco, Asia, Russia, Cyprus, Greece Mermessus O. Pickard-Cambridge, 1899 – China, Japan, Korea, Russia, North America, South America. Introduced worldwide Mesasigone Tanasevitch, 1989 – Kazakhstan, China, Iran, Russia Metafroneta Blest, 1979 – New Zealand Metaleptyphantes Locket, 1968 – Africa, Indonesia Metamynoglenes Blest, 1979 – New Zealand Metapanamomops Millidge, 1977 – Finland Metopobactrus Simon, 1884 – Japan, Korea, Caucasus, Turkey, Europe, Canada Micrargpelecopsis Wunderlich, 2025 – Portugal Micrargus Dahl, 1886 – Uganda, Europe, Asia, North America, North Africa Microbathyphantes van Helsdingen, 1985 – Angola, Cameroon, Seychelles, Nigeria, Asia Microctenonyx Dahl, 1886 – Nigeria, Italy, Macaronesia, North Africa to Kyrgyzstan. Introduced worldwide Microcyba Holm, 1962 – Africa Microlinyphia Gerhardt, 1928 – Africa, Asia, Russia, North America Microneta Menge, 1869 – Asia, Russia, Sweden, United States to Brazil, New Guinea, North Africa Microplanus Millidge, 1991 – Panama, Colombia Midia Saaristo & Wunderlich, 1995 – Europe Miftengris Eskov, 1993 – Russia Millidgea Locket, 1968 – Angola Millidgella Kammerer, 2006 – Argentina, Chile Minicia Thorell, 1875 – Algeria, Kazakhstan, Western Asia, Europe Minyriolus Simon, 1884 – Caucasus, Austria, Russia, Croatia, Italy, France, Argentina Mioxena Simon, 1926 – Angola, Congo, Kenya Mitrager van Helsdingen, 1985 – Indonesia, India, Nepal Moebelia Dahl, 1886 – China, Caucasus, Germany, France Moebelotinus Wunderlich, 1995 – Mongolia, Russia Molestia Tu, Saaristo & Li, 2006 – China, France Monocephalus Smith, 1906 – Europe Monocerellus Tanasevitch, 1983 – Russia Montilaira Chamberlin, 1921 – North America Moreiraxena Miller, 1970 – Angola Moyosi Miller, 2007 – Argentina, Brazil, Guyana Mughiphantes Saaristo & Tanasevitch, 1999 – Asia, Europe Murphydium Jocqué, 1996 – Kenya, Somalia Mycula Schikora, 1994 – Austria, Germany, Italy Myrmecomelix Millidge, 1993 – Ecuador, Peru Mythoplastoides Crosby & Bishop, 1933 – North America Napometa Benoit, 1977 – St. Helena Nasoona Locket, 1982 – Asia, Venezuela Nasoonaria Wunderlich & Song, 1995 – China, Southeast Asia Nematogmus Simon, 1884 – Asia, Russia, North Africa Nenilinium Eskov, 1988 – Mongolia, Russia Nentwigia Millidge, 1995 – Indonesia, Thailand Neocautinella Baert, 1990 – South America Neodietrichia Özdikmen, 2008 – North America Neoeburnella Koçak, 1986 – Ivory Coast Neomaso Forster, 1970 – South America Neonesiotes Millidge, 1991 – Oceania Neriene Blackwall, 1833 – Africa, Asia, Europe, North America, Brazil, Uruguay Neserigone Eskov, 1992 – Japan, Russia Nesioneta Millidge, 1991 – Asia, Oceania, United Arab Emirates Nihonella Ballarin & Yamasaki, 2021 – Japan Nippononeta Eskov, 1992 – Eastern Asia, Russia Nipponotusukuru Saito & Ono, 2001 – Japan Nispa Eskov, 1993 – Japan, Russia Notholepthyphantes Millidge, 1985 – Chile Nothophantes Merrett & Stevens, 1995 – Britain Notiogyne Tanasevitch, 2007 – Russia Notiohyphantes Millidge, 1985 – Brazil, Chile, Galapagos Notiomaso Banks, 1914 – Argentina, Chile Notioscopus Simon, 1884 – South Africa, China, Mongolia, Russia Notolinga Lavery & Dupérré, 2019 – Argentina Novafroneta Blest, 1979 – New Zealand Novafrontina Millidge, 1991 – Colombia, Ecuador, Peru Novalaetesia Millidge, 1988 – New Zealand Nusoncus Wunderlich, 2008 – Europe Oaphantes Chamberlin & Ivie, 1943 – North America Obrimona Strand, 1934 – Sri Lanka Obscuriphantes Saaristo & Tanasevitch, 2000 – Kazakhstan, China, Caucasus, Turkey, Russia, Southern Europe Oculocornia Oliger, 1985 – Russia Oedothorax Bertkau, 1883 – Ethiopia, Asia, Europe, North America, North Africa Oia Wunderlich, 1973 – Asia, Russia Oilinyphia Ono & Saito, 1989 – China, Japan, Thailand Oilinyphioides Irfan, Zhang & Peng, 2025 – China Okhotigone Eskov, 1993 – China, Japan, Russia Onychembolus Millidge, 1985 – Argentina, Chile Oreocyba Holm, 1962 – Kenya, Uganda Oreoneta Kulczyński, 1894 – Kazakhstan, China, Mongolia, Europe, North America Oreonetides Strand, 1901 – China, Japan, Taiwan, Europe, North America Oreophantes Eskov, 1984 – North America Orfeo Miller, 2007 – Brazil Origanates Crosby & Bishop, 1933 – United States Orsonwelles Hormiga, 2002 – Hawaii Oryphantes Hull, 1933 – Kazakhstan, Russia, North America Ostearius Hull, 1911 – China. Introduced to Africa, Indonesia, Malaysia, Canada, New Zealand Ouedia Bosmans & Abrous, 1992 – Algeria, Tunisia, Southern Europe Pachydelphus Jocqué & Bosmans, 1983 – Gabon, Ivory Coast, Sierra Leone Pacifiphantes Eskov & Marusik, 1994 – China, Korea, Russia, North America Pahangone Tanasevitch, 2018 – Malaysia Paikiniana Eskov, 1992 – China, Japan, Korea Palaeohyphantes Millidge, 1984 – Australia Palliduphantes Saaristo & Tanasevitch, 2001 – North Africa, Kyrgyzstan, Nepal, Caucasus, Iran, Turkey, Europe Panamomops Simon, 1884 – Kazakhstan, Kyrgyzstan, Pakistan, Caucasus, Europe Paracornicularia Crosby & Bishop, 1931 – United States Paracymboides Tanasevitch, 2011 – India Paraeboria Eskov, 1990 – Russia Parafroneta Blest, 1979 – New Zealand Paraglyphesis Eskov, 1991 – Russia Paragongylidiellum Wunderlich, 1973 – India, Nepal Paraletes Millidge, 1991 – Brazil, Peru Parameioneta Locket, 1982 – China, Southeast Asia Parapelecopsis Wunderlich, 1992 – Georgia, Madeira, Portugal, France, Britain Parasisis Eskov, 1984 – China, Japan, Korea, Russia Paratapinocyba Saito, 1986 – Japan Paratmeticus Marusik & Koponen, 2010 – Japan, Russia Parawubanoides Eskov & Marusik, 1992 – Mongolia, Russia Parbatthorax Tanasevitch, 2019 – China, Nepal Parhypomma Eskov, 1992 – Japan Paro Berland, 1942 – French Polynesia Parvunaria Tanasevitch, 2018 – Myanmar Patagoneta Millidge, 1985 – Chile Pecado Hormiga & Scharff, 2005 – Algeria, Morocco, Spain Pelecopsidis Bishop & Crosby, 1935 – United States Pelecopsis Simon, 1864 – Africa, Asia, Europe, North America Peponocranium Simon, 1884 – Mongolia, Georgia, Europe Perlongipalpus Eskov & Marusik, 1991 – Mongolia, Russia Perregrinus Tanasevitch, 1992 – China, Mongolia, Russia, Canada Perro Tanasevitch, 1992 – Russia, Canada Phanetta Keyserling, 1886 – United States Phyllarachne Millidge & Russell-Smith, 1992 – Brunei Piesocalus Simon, 1894 – Indonesia Piniphantes Saaristo & Tanasevitch, 1996 – Asia, Alps, France Pityohyphantes Simon, 1929 – Kazakhstan, Japan, Russia, North America Plaesianillus Simon, 1926 – France Platyspira Song & Li, 2009 – China Plectembolus Millidge & Russell-Smith, 1992 – Indonesia, Malaysia, Philippines Plesiophantes Heimer, 1981 – Caucasus, Georgia, Turkey, Russia Plicatiductus Millidge & Russell-Smith, 1992 – Indonesia Pocadicnemis Simon, 1884 – Asia, Russia, Southern Europe, North America Pocobletus Simon, 1894 – North to South America Poecilafroneta Blest, 1979 – New Zealand Poeciloneta Kulczyński, 1894 – Kazakhstan, Kyrgyzstan, China, Georgia, Iran, Russia, North America Porrhomma Simon, 1884 – Asia, Europe, North America Praestigia Millidge, 1954 – Japan, Mongolia, Europe, Alaska, Canada, Greenland Primerigonina Wunderlich, 1995 – Panama Prinerigone Millidge, 1988 – Africa, Asia Procerocymbium Eskov, 1989 – Russia, Canada Proelauna Jocqué, 1981 – Angola, Malawi, Tanzania Proislandiana Tanasevitch, 1985 – Armenia, Turkey, Russia Promynoglenes Blest, 1979 – New Zealand Pronasoona Millidge, 1995 – Malaysia, Thailand Prosoponoides Millidge & Russell-Smith, 1992 – China, Southeast Asia, India Protoerigone Blest, 1979 – New Zealand Protopalpus Tanasevitch, 2021 – Thailand Protosintula Tanasevitch, 2021 – Israel Pseudafroneta Blest, 1979 – New Zealand Pseudocarorita Wunderlich, 1980 – Central Europe Pseudocyba Tanasevitch, 1984 – Kazakhstan, Russia Pseudohilaira Eskov, 1990 – Russia Pseudomaro Denis, 1966 – Europe Pseudomaso Locket & Russell-Smith, 1980 – Nigeria Pseudomicrargus Eskov, 1992 – Japan Pseudomicrocentria Miller, 1970 – Africa, Malaysia, Singapore Pseudoporrhomma Eskov, 1993 – Russia Pseudotapinopa Wunderlich, 2024 – Ecuador Pseudotyphistes Brignoli, 1972 – South America Pseudowubana Eskov & Marusik, 1992 – Mongolia, Russia Psilocymbium Millidge, 1991 – South America Putaoa Hormiga & Tu, 2008 – China, Taiwan Racata Millidge, 1995 – Indonesia, Thailand Rhabdogyna Millidge, 1985 – Chile Ringina Tambs-Lyche, 1954 – Crozet Islands Russocampus Tanasevitch, 2004 – Caucasus, Ukraine, Russia Ryojius Saito & Ono, 2001 – China, Japan, Korea Saaristoa Millidge, 1978 – Japan, North America Sachaliphantes Saaristo & Tanasevitch, 2004 – China, Japan, Korea, Russia Saitonia Eskov, 1992 – China, Japan, Korea Saloca Simon, 1926 – Nepal, Turkey, Russia, Central Satilatlas Keyserling, 1886 – Russia, North America, Central, Western Sauron Eskov, 1995 – Kazakhstan, Turkey, Russia Savignia Blackwall, 1833 – Comoros, Ethiopia, Algeria, Kazakhstan, East Asia, Caucasus, Russia, Europe, Canada, Australia Savigniorrhipis Wunderlich, 1992 – Azores Scandichrestus Wunderlich, 1995 – Russia, Estonia, Finland, Sweden Sciastes Bishop & Crosby, 1938 – Austria, Switzerland, Russia, Italy, France, North America Scirites Bishop & Crosby, 1938 – North America Scironis Bishop & Crosby, 1938 – North America Scolecura Millidge, 1991 – Argentina, Brazil, Colombia Scolopembolus Bishop & Crosby, 1938 – North America Scotargus Simon, 1913 – Algeria, Morocco, Kazakhstan, Nepal, Caucasus, Iran, Canary Islands, Russia Scotinotylus Simon, 1884 – Kazakhstan, China, Japan?, Mongolia, Europe, North America Scutpelecopsis Marusik & Gnelitsa, 2009 – Caucasus, Iran, Turkey, Bulgaria, Romania, Southern Europe Scylaceus Bishop & Crosby, 1938 – North America Scyletria Bishop & Crosby, 1938 – North America Selenyphantes Gertsch & Davis, 1946 – Costa Rica, Guatemala, Mexico Semljicola Strand, 1906 – Kazakhstan, China, Mongolia, Russia, Norway, Scandinavia, North America, Britain Sengletus Tanasevitch, 2008 – Egypt, Iran, Israel Shaanxinus Tanasevitch, 2006 – China, Taiwan, Vietnam, Russia Shanus Tanasevitch, 2006 – China Sibirocyba Eskov & Marusik, 1994 – Russia Silometopoides Eskov, 1990 – Kazakhstan, Eastern Asia, Russia, North America Silometopus Simon, 1926 – Egypt?, Kazakhstan, China, Japan, Iran, Israel, Europe Simplicistilus Locket, 1968 – Angola, Democratic Republic of the Congo, Nigeria Singatrichona Tanasevitch, 2019 – Singapore Sinisterigone Irfan, Zhang & Peng, 2022 – China Sinogone Irfan, Wang & Zhang, 2023 – China Sinolinyphia Wunderlich & Li, 1995 – China, Russia Sinopimoa Li & Wunderlich, 2008 – China Sintula Simon, 1884 – Algeria, Morocco, Tunisia, Western Asia, Europe Sisicottus Bishop & Crosby, 1938 – Russia, North America Sisicus Bishop & Crosby, 1938 – Europe, Russia, North America Sisis Bishop & Crosby, 1938 – North America Sisyrbe Bishop & Crosby, 1938 – United States Sitalcas Bishop & Crosby, 1938 – United States Smerasia Zhao & Li, 2014 – China Smermisia Simon, 1894 – Costa Rica, South America Smodix Bishop & Crosby, 1938 – North America Solenysa Simon, 1894 – Eastern Asia Soucron Crosby & Bishop, 1936 – North America Souessa Crosby & Bishop, 1936 – North America Souessoula Crosby & Bishop, 1936 – North America Sougambus Crosby & Bishop, 1936 – North America Souidas Crosby & Bishop, 1936 – North America Soulgas Crosby & Bishop, 1936 – North America Spanioplanus Millidge, 1991 – South America Spelaeus Irfan, Zhang & Peng, 2025 – China Sphecozone O. Pickard-Cambridge, 1871 – Trinidad, United States, South America Spiralophantes Tanasevitch & Saaristo, 2006 – Nepal Spirembolus Chamberlin, 1920 – North America Staveleya Sherwood, 2021 – Switzerland, Ukraine, Italy, France Stemonyphantes Menge, 1866 – Asia, Ukraine, Russia, Italy, Slovenia, France, North America Sthelota Simon, 1894 – Guatemala, Panama Stictonanus Millidge, 1991 – Chile, Falkland Islands Strandella Oi, 1960 – China, Japan, Korea, Russia Strongyliceps Fage, 1936 – Kenya, Uganda Styloctetor Simon, 1884 – Asia, Alps, Russia, North America, North Africa Subbekasha Millidge, 1984 – Canada Syedra Simon, 1884 – DR Congo, China, Japan, Korea, Europe Symmigma Crosby & Bishop, 1933 – North America Tachygyna Chamberlin & Ivie, 1939 – North America Taibainus Tanasevitch, 2006 – China Taibaishanus Tanasevitch, 2006 – China Tallusia Lehtinen & Saaristo, 1972 – Kazakhstan, China, Japan, Caucasus, Turkey, Russia, Greece, Italy Tanasevitchia Marusik & Saaristo, 1999 – Russia Tapinocyba Simon, 1884 – Algeria, Morocco, Asia, Europe, North America Tapinocyboides Wiehle, 1960 – India, Russia Tapinopa Westring, 1851 – Algeria, Asia, Russia, Southern Europe, North America Taranucnus Simon, 1884 – Japan, Turkey, Poland, Slovakia, Romania, Ukraine, Russia, North America Tarsiphantes Strand, 1905 – Russia, Alaska, Canada, Greenland Tchatkalophantes Tanasevitch, 2001 – Kazakhstan, Kyrgyzstan, China, Mongolia, Pakistan Tegulinus Tanasevitch, 2017 – Indonesia Tennesseellum Petrunkevitch, 1925 – Bonaire, North America. Introduced to Marshall Islands Tenuiphantes Saaristo & Tanasevitch, 1996 – Tanzania, Algeria, Asia, Europe, North America, Balkans?. Introduced to New Zealand, Argentina, Chile, Falkland Is Ternatus Sun, Li & Tu, 2012 – China Tessamoro Eskov, 1993 – Russia Thainetes Millidge, 1995 – Thailand Thaiphantes Millidge, 1995 – Thailand Thaleria Tanasevitch, 1984 – Russia, Alaska Thapsagus Simon, 1894 – Madagascar Thaumatoncus Simon, 1884 – Algeria, Tunisia, Israel, Italy, Spain, France Theoa Saaristo, 1995 – Seychelles, China, Southeast Asia Theoneta Eskov & Marusik, 1991 – Russia Theonina Simon, 1929 – Caucasus, Georgia?, Israel, Ukraine, Russia, Central Europe to Greece, North Africa Thyreobaeus Simon, 1889 – Madagascar Thyreosthenius Simon, 1884 – Caucasus, Russia Tibiaster Tanasevitch, 1987 – Kazakhstan, Russia Tibioploides Eskov & Marusik, 1991 – China, Japan, Russia, Estonia, Scandinavia, Alaska Tibioplus Chamberlin & Ivie, 1947 – Kyrgyzstan, Mongolia, Russia, Scandinavia, Alaska, Canada Tiso Simon, 1884 – Asia, Madeira, Russia, Canada, Greenland Tmeticodes Ono, 2010 – Japan Tmeticus Menge, 1868 – Eastern Asia, Russia, North America Tojinium Saito & Ono, 2001 – Japan Toltecaria Miller, 2007 – Mexico Tomohyphantes Millidge, 1995 – Indonesia Totua Keyserling, 1891 – Brazil Trachyneta Holm, 1968 – Congo, Malawi Traematosisis Bishop & Crosby, 1938 – North America Trematocephalus Dahl, 1886 – China, Sri Lanka, Caucasus, Iran, Turkey, Russia, France Trichobactrus Wunderlich, 1995 – Mongolia, Russia Trichoncoides Denis, 1950 – Kazakhstan, Caucasus, Iran, Turkey, Russia, France, North Africa Trichoncus Simon, 1884 – Kenya, North Africa, Kazakhstan, China, Western Asia, Europe Trichoncyboides Wunderlich, 2008 – Bulgaria, Romania, Italy, France Trichopterna Kulczyński, 1894 – Tanzania, Algeria, Morocco, Caucasus, Turkey, Russia, Southern Europe Trichopternoides Wunderlich, 2008 – Russia Triplogyna Millidge, 1991 – Argentina, Brazil, Colombia Troglohyphantes Joseph, 1882 – Algeria, Tunisia, Western Asia, Europe Troxochrota Kulczyński, 1894 – Germany, Switzerland, Romania, Russia, Estonia, Scandinavia Troxochrus Simon, 1884 – Angola, Caucasus, Israel, Turkey, Russia, Sweden, Greece Tubercithorax Eskov, 1988 – Russia Tunagyna Chamberlin & Ivie, 1933 – Russia, North America Turbinellina Millidge, 1993 – Argentina, Chile Turinyphia van Helsdingen, 1982 – China, Japan, Korea, Azores, Madeira, Alps Tusukuru Eskov, 1993 – Russia, North America Tutaibo Chamberlin, 1916 – Guatemala, Mexico, United States, South America Tybaertiella Jocqué, 1979 – Ethiopia, Ivory Coast, Nigeria, East Africa, Central, West Typhistes Simon, 1894 – Ethiopia, Sri Lanka Typhlonyphia Kratochvíl, 1936 – Croatia Typhochrestinus Eskov, 1990 – Russia Typhochrestoides Eskov, 1990 – Russia Typhochrestus Simon, 1884 – Northern Africa, Caucasus, Israel, Turkey, Europe, North America Uahuka Berland, 1935 – Marquesas Islands Uapou Berland, 1935 – Marquesas Islands Ulugurella Jocqué & Scharff, 1986 – Tanzania Ummeliata Strand, 1942 – Asia, Russia Uralophantes Esyunin, 1992 – Ukraine, Russia Ussurigone Eskov, 1993 – Russia Vagiphantes Saaristo & Tanasevitch, 2004 – Central Asia, Pakistan Venia Seyfulina & Jocqué, 2009 – Kenya Vermontia Millidge, 1984 – Russia, North America Vesicapalpus Millidge, 1991 – Argentina, Brazil Vietnagone Tanasevitch, 2019 – China, Vietnam Viktorium Eskov, 1988 – Russia Vittatus Zhao & Li, 2014 – China Wabasso Millidge, 1984 – Russia, Iceland, Scandinavia, North America, Britain Walckenaeria Blackwall, 1833 – Africa, Asia, Europe, North America, Colombia Walckenaerianus Wunderlich, 1995 – Kazakhstan, Mongolia, Bulgaria, Russia Weintrauboa Hormiga, 2003 – China, Japan, Russia Wiehlea Braun, 1959 – Germany, Spain, Belgium, France, Britain Wiehlenarius Eskov, 1990 – Austria, Switzerland, Russia, Greece, Italy Wubana Chamberlin, 1919 – North America Wubanoides Eskov, 1986 – Japan, Mongolia, Czechia, Germany, Russia Wulinggone Irfan, Zhang & Peng, 2025 – China Wulingnus Irfan, Zhang & Peng, 2025 – China Wuliphantes Irfan, Wang & Zhang, 2023 – China Xim Ibarra-Núñez, Chamé-Vázquez & Maya-Morales, 2021 – Mexico Yakutopus Eskov, 1990 – Russia Yuelushannus Irfan, Zhou, Bashir, Mukhtar & Peng, 2020 – China Zerogone Eskov & Marusik, 1994 – Russia Zhezhoulinyphia Irfan, Zhou & Peng, 2019 – China Zilephus Simon, 1902 – Argentina Zornella Jackson, 1932 – Kazakhstan, Mongolia, Poland, Belarus, Russia, Estonia, Lithuania, Scandinavia, North America Zygottus Chamberlin, 1949 – North America
  • Bosselaers, J & Henderickx, H. (2002) A new Savignia from Cretan caves (Araneae: Linyphiidae). Zootaxa 109:1-8 [permanent dead link]
  • Hågvar, S. & Aakra, K. 2006. Spiders active on snow in Southern Norway. Norw. J. Entomol. 53, 71-82.

External links

  • Spiders rebuild webs en masse after floods in Wagga Wagga