Novavis (meaning "strange bird") is an extinct genus of enantiornithean birds from the Early Cretaceous Xiagou Formation of China. The genus contains a single species, N. pubisculata, known from a partial skeleton. The pubis bone of Novavis is unusually short compared to other related birds.

Discovery and naming

The Novavis holotype specimen, IVPP V31957, was discovered in 2005 in sediments of the Xiagou Formation (Changma Basin locality) near Changma Town in Yumen City of Gansu Province, China. The specimen is incomplete, comprising six or seven caudal vertebrae, a partial pelvic girdle (the right ischium and both pubes), and most of both hindlimbs, preserved on a single slab. Like many other bird fossils initially found in this locality, the holotype features a thick reflective layer of consolidant to secure the specimen. However, this obscures some details and prevents the precise identification of some bones. The specimen was initially housed at the Chinese Academy of Geological Sciences under the catalogue number CAGS-IG-05-CM-012 before being moved to the Institute of Vertebrate Paleontology and Paleoanthropology.

In 2025, O'Connor et al. described Novavis pubisculata as a new genus and species of enantiornithean birds based on these fossil remains. The generic name, Novavis, combines the Latin words novus, meaning "strange" and avis, meaning "bird". It is also intended to honor Nova R. J. Sumita. The specific name, pubisculata, is derived from roots that mean "short" or "diminutive" pubis. The full binomial name translates to "strange bird with short pubis", in reference to the unusual proportions of the pelvic girdle.

Four other enantiornithean birds—all known from incomplete skeletons—were named from the Xiagou Formation prior to Novavis: Qiliania, Dunhuangia, Feitianius, and Avimaia. The formation's bird fauna is otherwise dominated by more than 70 specimens of the ornithuromorph Gansus yumenensis.

Classification

In their phylogenetic analyses, O'Connor et al. (2025) recovered Novavis as a member of the extinct Cretaceous bird clade Enantiornithes. They noted that their analyses found Novavis to be more basal than the other Xiagou Formation enantiornitheans that it coexisted with (highlighted below) except for Dunhuangia, which it was recovered in a polytomy with alongside Eocathayornis. Their results are displayed in the cladogram below:

OrnithothoracesEuornithesEnantiornithes Protopteryx Iberomesornis Pengornis Eopengornis Rapaxavis Longirostravis Shanweiniao Longipteryx Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Euornithes
EnantiornithesProtopteryx Iberomesornis Pengornis Eopengornis Rapaxavis Longirostravis Shanweiniao Longipteryx Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Protopteryx Iberomesornis
Protopteryx
Iberomesornis
Pengornis Eopengornis Rapaxavis Longirostravis Shanweiniao Longipteryx Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Pengornis Eopengornis
Pengornis
Eopengornis
Rapaxavis Longirostravis Shanweiniao Longipteryx Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Rapaxavis Longirostravis
Rapaxavis
Longirostravis
Shanweiniao Longipteryx Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Shanweiniao
Longipteryx
Novavis Dunhuangia Eocathayornis Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Novavis
Dunhuangia
Eocathayornis
Eoalulavis Concornis Cathayornis yandica Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Eoalulavis Concornis Cathayornis yandica
Eoalulavis
Concornis
Cathayornis yandica
Neuquenornis Halimornis Enantiophoenix Feitianius GSGM-FV-00516 (indeterminate unnamed enantiornithean) Qiliania Avimaia Shenqiornis Eoenantiornis Elsornis
Neuquenornis
Halimornis
Enantiophoenix
Feitianius
GSGM-FV-00516 (indeterminate unnamed enantiornithean)
Qiliania
Avimaia
Shenqiornis
Eoenantiornis
Elsornis

See also