The Xiagou Formation is the middle strata of the Xinminbao Group. It is named for its type site in Xiagou, in the Changma Basin of Gansu Province, northwestern China and is considered Early Cretaceous in age. It is known outside the specialized world of Chinese geology as the site of a Lagerstätte in which the fossils were preserved of Gansus yumenensis, the earliest true modern bird.

Description

The laminated yellowish mudstones of the Xiagou Formation are the lithified remnants of varves that were laid down as extremely fine silt settled to the bottom of a tranquil freshwater lake. The result was dense anoxic bottom sediment, where the lack of bacteria slowed the processes of decay, preserving uncompressed fossils in details that include feather impressions and remnants of the webbing between the bird's toes. The age of the formation has not yet been confidently determined. The underlying Chijinpu Formation is likely the same age as the Jehol Group due to the presence of similar fossils, meaning that the Xiagou Formation is probably slightly younger than the Jehol biota, dating to around the late Aptian.

Fossil content

The Xiagou Formation is particularly noted for its high diversity of ancient birds. These include both modern birds close to the ancestors of living species, and related lineages now entirely extinct. Other fossils from the Xiagou Formation are characteristic of an Early Cretaceous lake ecology. There are fossils of abundant fish fauna, Charophyta and ostracods.

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Invertebrates

Arthropods

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
BlattapterixB. gansu
CretacechoristaC. qilianshanensis
EurycoleusE. arcuatus, E. clypeolatus, E. dimorphocellatus, E. parvusA carabid.
GlottocoleusG. lenticulata, G. stellatus
MesoblattinaM. cretacea
MesocoleusM. zhonggouense
MesotricupesM. reticulatus
PetalocupesP. arcus
PhyllocoleusP. striolatus
PlanocoleusP. ensatus
PleurocoleusP. catenatus
SpinusS. yumenense
TetillopsisT. parvula
YumenocoleusY. intermedius, Y. lineatus, Y. longus

Theropods

Birds

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
AvimaiaA. schweitzeraeChangma Basin"Partial skeleton with an egg."An enantiornithean.
BrevidentavisB. zhangiIncomplete skull and cervical vertebrae.An ornithuromorph.
ChangmaornisC. houiChangma Basin"Partial pelvic girdle with hindlimb."An euornithian.
DunhuangiaD. cuiiChangma Basin"Forelimbs."An enantiornithean.
FeitianiusF. paradisiChangma Basin"Hindlimbs with pelvic area and vertebrae."An enantiornithean.
GansusG. yumenensis"Partial hindlimbs."An euornithian.
JiuquanornisJ. niuiChangma Basin"Partial pectoral girdle."An euornithian.
MeemannavisM. ductrixIncomplete skull and cervical and thoracic vertebrae.An ornithuromorph.
NovavisN. pubisculataChangma BasinPartial skeleton including tail vertebrae, a pelvis, and the hindlimbs.An enantiornithean.
QilianiaQ. graffiniChangma Basin"Partial skeleton from two individuals."An enantiornithean.
Unnamed enantiornitheanIndeterminate"Partial forelimb."An enantiornithean.
YumenornisY. huangiChangma Basin"Right forelimb."An euornithian.

Ornithomimosaurs

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
BeishanlongB. grandisYujingzi BasinGray-variegated beds"Fore and hindlimbs with partial vertebrae."An ornithomimosaur.

Therizinosaurs

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
SuzhousaurusS. megatherioidesYujingzi BasinGray-variegated beds"Dorsal vertebrae, ribs and partial shoulder girdle."A therizinosauroid.

Tyrannosauroids

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
XiongguanlongX. baimoensisYujingzi BasinGray-variegated beds"Skull without lower jaws, partial vertebrae, ilium and femur."A tyrannosauroid.

Cerapods

Ceratopsians

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
ArchaeoceratopsA. yujingziensisYujingzi BasinGray-variegated beds"Caudal vertebrae and a partial hindlimb."A neoceratopsian.

Hadrosauroids

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
JintasaurusJ. meniscusYujingzi BasinGray-variegated beds"Partial skull."A hadrosauroid.
XuwulongX. yueluniYujingzi BasinGray-variegated beds"Articulated skeleton lacking limbs."A hadrosauroid.

Sauropods

Macronarians

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
QiaowanlongQ. kangxiiYujingzi BasinGray-variegated beds"Cervical vertebrae and right pelvic girdle."A somphospondylan.

Other vertebrates

Turtles

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
ChangmachelysC. bohliniChangma Basin"Nearly complete skeletons from four individuals."A macrobaenid.

Fish

GenusSpeciesLocationStratigraphic positionMaterialNotesImages
ShuleichthysS. brachypteryx"Complete skeleton from four individuals."An osteoglossomorph.

Plants

GenusSpeciesLocationStratigraphic memberMaterialNotesImages
XenoxylonX. meisteriHongliuxiaWoodA podocarp

See also