Closeup of hand, showing claws
Closeup of skull

Megatheriidae is a family of extinct ground sloths that lived from approximately 23 mya—11,000 years ago.

Megatheriids appeared during the Late Oligocene (Deseadan in the SALMA classification), some 29 million years ago, in South America. The group includes the largest known ground sloths, the elephant sized Megatherium (given its name 'great beast' by Georges Cuvier) and Eremotherium. An early genus that was originally considered a megatheriid, the more slightly built Hapalops, reached a length of about 1.2 metres (3.9 ft). The nothrotheres have recently been placed in their own family, Nothrotheriidae.

The skeletal structure of these ground sloths indicates that the animals were massive. Their thick bones and even thicker joints (especially those on the hind legs) gave their appendages tremendous power that, combined with their size and fearsome claws, provided a formidable defense against predators.

The earliest megatheriid in North America was Sibotherium which arrived 5.3 million years ago, after crossing the recently formed Panamanian land bridge. At more than five tons in weight, 6 metres (20 ft) in length, and able to reach as high as 5.2 metres (17 ft), Eremotherium when fully erect was taller than an African bush elephant bull. Unlike relatives, this species retained a plesiomorphic extra claw. While other species of Eremotherium had four fingers with only two or three claws, E. eomigrans had five fingers, four of them with claws up to nearly 30 centimetres (1 ft) long.

Phylogeny

The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data (see Fig. 4 of Presslee et al., 2019).

FolivoraMegalocnidae (Caribbean sloths) Mylodontoidea † ScelidotheriidaeMylodontidae Choloepodidae (two-toed sloths) Megatherioidea † MegalonychidaeMegalonyx jeffersoni Bradypodidae Bradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus (three-toed sloths) † NothrotheriidaeNothrotheriops shastensis † Megatheriidae † Megatherium americanum
Megalocnidae (Caribbean sloths)
Mylodontoidea † ScelidotheriidaeMylodontidae Choloepodidae (two-toed sloths) Megatherioidea † MegalonychidaeMegalonyx jeffersoni Bradypodidae Bradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus (three-toed sloths) † NothrotheriidaeNothrotheriops shastensis † Megatheriidae † Megatherium americanum
Mylodontoidea † ScelidotheriidaeMylodontidae Choloepodidae (two-toed sloths) Megatherioidea † MegalonychidaeMegalonyx jeffersoni Bradypodidae Bradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus (three-toed sloths) † NothrotheriidaeNothrotheriops shastensis † Megatheriidae † Megatherium americanum
MylodontoideaScelidotheriidaeMylodontidae Choloepodidae (two-toed sloths)
Scelidotheriidae
Mylodontidae Choloepodidae (two-toed sloths)
Mylodontidae
Choloepodidae (two-toed sloths)
MegatherioideaMegalonychidaeMegalonyx jeffersoni Bradypodidae Bradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus (three-toed sloths) † NothrotheriidaeNothrotheriops shastensis † Megatheriidae † Megatherium americanum
MegalonychidaeMegalonyx jeffersoni Bradypodidae Bradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus (three-toed sloths)
MegalonychidaeMegalonyx jeffersoni
BradypodidaeBradypus torquatus Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus
Bradypus torquatus
Bradypus pygmaeus Bradypus tridactylus Bradypus variegatus
Bradypus pygmaeus
Bradypus tridactylus Bradypus variegatus
Bradypus tridactylus
Bradypus variegatus
(three-toed sloths)
NothrotheriidaeNothrotheriops shastensis † Megatheriidae † Megatherium americanum
NothrotheriidaeNothrotheriops shastensis
† MegatheriidaeMegatherium americanum

The following phylogeny is based on Varela et al. 2019.

FolivoraPseudoglyptodon Bradypodidae Bradypus Eufolivora Mylodontidae Nematherium Scelidotheriinae Analcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon Megatherioidea Schismotherium Pelecyodon Megalonychidae Hapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus Megatheria Analcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
Pseudoglyptodon
Bradypodidae Bradypus Eufolivora Mylodontidae Nematherium Scelidotheriinae Analcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon Megatherioidea Schismotherium Pelecyodon Megalonychidae Hapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus Megatheria Analcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
BradypodidaeBradypus
EufolivoraMylodontidae Nematherium Scelidotheriinae Analcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon Megatherioidea Schismotherium Pelecyodon Megalonychidae Hapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus Megatheria Analcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
MylodontidaeNematherium Scelidotheriinae Analcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
Nematherium
Scelidotheriinae Analcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
ScelidotheriinaeAnalcitherium Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes
Analcitherium
Neonematherium Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes
Neonematherium
Sibyllotherium Proscelidodon Catonyx Scelidotherium Valgipes
Sibyllotherium
Proscelidodon Catonyx Scelidotherium Valgipes
Proscelidodon
Catonyx Scelidotherium Valgipes
Catonyx
Scelidotherium Valgipes
Scelidotherium
Valgipes
Pseudoprepotherium Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
Pseudoprepotherium
Orophodontinae? Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys Mylodontinae Mylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
Orophodontinae?Brievabradys Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys
Brievabradys
Octodontotherium Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys
Octodontotherium
Paroctodontotherium Octomylodon Urumacotherium Baraguatherium Octodontobradys
Paroctodontotherium
Octomylodon Urumacotherium Baraguatherium Octodontobradys
Octomylodon
Urumacotherium Baraguatherium Octodontobradys
Urumacotherium
Baraguatherium Octodontobradys
Baraguatherium
Octodontobradys
MylodontinaeMylodon Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
Mylodon
Thinobadistes Lestodon Lestobradys Bolivartherium Simomylodon Pleurolestodon Glossotherium Paramylodon
Thinobadistes Lestodon Lestobradys Bolivartherium
Thinobadistes
Lestodon Lestobradys Bolivartherium
Lestodon
Lestobradys Bolivartherium
Lestobradys
Bolivartherium
Simomylodon Pleurolestodon Glossotherium Paramylodon
Simomylodon Pleurolestodon
Simomylodon
Pleurolestodon
Glossotherium Paramylodon
Glossotherium
Paramylodon
MegatherioideaSchismotherium Pelecyodon Megalonychidae Hapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus Megatheria Analcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
Schismotherium Pelecyodon
Schismotherium
Pelecyodon
Megalonychidae Hapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus Megatheria Analcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
MegalonychidaeHapalops Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Hapalops
Hyperleptus Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Hyperleptus
Eucholoeops Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Eucholoeops
Pseudortotherium Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Pseudortotherium
Megalonychotherium Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Megalonychotherium
Megalonychinae Pliometanastes Megalonyx Australonyx Megistonyx Ahytherium Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
MegalonychinaePliometanastes Megalonyx Australonyx Megistonyx Ahytherium
Pliometanastes
Megalonyx Australonyx Megistonyx Ahytherium
Megalonyx
Australonyx Megistonyx Ahytherium
Australonyx
Megistonyx Ahytherium
Megistonyx
Ahytherium
Pliomorphus Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
Pliomorphus
Megalocninae Megalocnus Parocnus Choloepodinae Acratocnus Neocnus Choloepus
MegalocninaeMegalocnus Parocnus
Megalocnus
Parocnus
CholoepodinaeAcratocnus Neocnus Choloepus
Acratocnus
Neocnus Choloepus
Neocnus
Choloepus
MegatheriaAnalcimorphus Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
Analcimorphus
Nothrotheriidae Thalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus Megatheriidae Planopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
NothrotheriidaeThalassocninae Thalassocnus Nothrotheriinae Lakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus
ThalassocninaeThalassocnus
NothrotheriinaeLakukullus Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus
Lakukullus
Mionothropus Nothrotherium Nothrotheriops Pronothrotherium Aymaratherium Xyophorus
Mionothropus Nothrotherium Nothrotheriops
Mionothropus
Nothrotherium Nothrotheriops
Nothrotherium
Nothrotheriops
Pronothrotherium Aymaratherium Xyophorus
Pronothrotherium
Aymaratherium Xyophorus
Aymaratherium
Xyophorus
MegatheriidaePlanopsinae Prepoplanops Planops Prepotherium Megatheriinae Diabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
PlanopsinaePrepoplanops Planops Prepotherium
Prepoplanops
Planops Prepotherium
Planops
Prepotherium
MegatheriinaeDiabolotherium Megathericulus Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
Diabolotherium Megathericulus
Diabolotherium
Megathericulus
Anisodontherium Pyramiodontherium Proeremotherium Eremotherium Megatherium
Anisodontherium
Pyramiodontherium Proeremotherium Eremotherium Megatherium
Pyramiodontherium
Proeremotherium Eremotherium Megatherium
Proeremotherium
Eremotherium Megatherium
Eremotherium
Megatherium

External links

  • Data related to Megatheriidae at Wikispecies