Grimmen Formation
Grimmen Formation
Stratigraphic range : Lower Toarcian ~182–179 Ma PreꞒ Ꞓ O S D C P T J K Pg N Tenuicostatum-Exaratum
Unit of Kamienna Group
Underlies Glashütte Formation, Dörnten Member (Posidonia Shale Formation ) Overlies Posidonia Shale Formation , Wolgast FormationThickness Up to 100 m (330 ft)
Lithology
Other Mudstone , fine-grained Sandstone , Siderite and carbonate concretionsLocation
Region Mecklenburg-Vorpommern
Country Germany
Extent North German Basin
Type section
Named for Grimmen , GermanyThe Grimmen Formation is a Lower Jurassic (Toarcian ) geological formation in northeastern Germany, primarily exposed in the Grimmen and Klein Lehmhagen clay pits and documented in wells such as Reinberg 1E. Formally established in 2025, it was previously part of the informal "Green Series" of the Ciechocinek Formation . It represents a prodelta to brackish -marine depositional system in the eastern North German Basin (NGB), shaped by sea-level changes and the Toarcian anoxic event .
History Studies of Toarcian strata near Grimmen began in 1874 with the discovery of fossiliferous clays in a railway cutting near Schönenwalde, 5 km north of Grimmen, initially misidentified as Middle Jurassic due to ammonite finds. In 1909, the succession was reclassified as Lower Toarcian (Lias epsilon), distinct from the Posidonia Shale Formation . From 1959 to 1995, the Grimmen clay pit was excavated, revealing a glacially dislocated raft of Liassic clays and sands deformed by Pleistocene ice advances. Exploration wells (e.g., Reinberg 1E, Kb Barth 10, Kb Grambow 5) since the 1950s provided extensive core data, with Reinberg 1E designated as the reference section in 2025. The clay pit , abandoned and water-filled since 1995, was established as the type section in 2025, with 2016 and 2020 excavations refining an 18 m composite log spanning the Pliensbachian -Toarcian boundary to the elegans Subzone.
Sedimentology/Lithology Profile at Grimmen The Grimmen Formation comprises greenish to bluish claystones , siltstones , and fine-grained sandstones , with siderite and carbonate concretions , reaching up to 100 m thick in the eastern North German Basin . The type section in the Grimmen clay pit forms an 18 m composite log, transitioning from Pliensbachian sandy deposits of the Wolgast Formation to Toarcian organo-detrital clays and heteroliths of the Lehmhagen Member (Posidonia Shale Formation ), followed by the Reinberg Member of the Grimmen Formation. The Reinberg Member (~10 m thick) starts with bluish clays , grading to greenish, pellet-laminated clays with exaratum and lower elegans marker beds (marly limestones with fecal pellets). Pellet laminae, up to 5 mm thick, decrease upsection, while silty/sandy intercalations and siderite concretions increase, forming a coarsening-upward trend with pyrite -filled burrows. The upper Grimmen Formation features greenish clays with thin silt /sand laminae in 8–15 m symmetric cycles, capped by the Glashütte Formation's deltaic sandstones . Heavy minerals (zircon , rutile , tourmaline ) and smectite -group clay minerals indicate volcanogenic input, while kaolinite and chlorite are common. Glacially dislocated Eocene greensand and clay are locally intercalated. Organic-rich layers contain charcoal fragments, suggesting wildfires .
Toarcian material found in glacial Erratics in Ahrensburg and the Hagen Forest have been in controversy due to its dubious origin, being linked with the Rya Formation and Sorthat Formation , as well this unit. They were originally considered or local or Baltic in derivation, but that changued with the recovery of erratic concretions in the Baltic sea cliffs near Lübeck , being found as part of the Weichselian Glacial Maximum. Liassic–Cretaceous sediments in the assemblage are most probably associated with the tectonic Sorgenfrei-Tornquist Zone . The origin of this erratics from southwestern Baltic, Poland or Danish archipelago is unlikely, as those zones are dominated by Late Cretaceous–Paleocene strata, suggesting that this Toarcian assamblages should come from south/SW between STZ–TTZ and the German Baltic coast. The most clear hint link this deposits with the Grimmen Fm, as they're identical in fauna and facies composition of Grimmen and Dobbertin, also affected by subglacial erosion and thrusting, suggesting a close stratigraphic and palaeogeographical origin.
Paleoenvironment Pliensbachian–Toarcian plankton communities in the NGB Simplified Paleogeography of the NGB in the Toarcian, with the extent of the Grimmen Formation and adjacent units The Grimmen Formation was deposited in a prodelta to brackish -marine environment in the eastern North German Basin , influenced by sea-level changes and the Toarcian anoxic event . The Reinberg Member reflects a transition from organo-detrital Posidonia Shale Formation to brackish -marine clays , with high total organic carbon (TOC, ~3 wt%) at the base decreasing to <1 wt% upsection, indicating a shift to dysoxic conditions. Basal bluish clays (exaratum Subzone) represent a restricted marine basin, followed by greenish clays with pellet laminae, suggesting increased fluvial input and suspension-load plumes during wetter phases. The coarsening-upward trend and onset of bioturbation (pyrite -filled burrows) reflect prodelta progradation from Fennoscandia , culminating in the Glashütte Formation's deltaic systems. High kaolinite and smectite content indicate a warm, humid climate with biochemical weathering and volcanogenic influence, akin to modern tropical settings. The absence of major ripples suggests minimal wave action, with sedimentation driven by fluvial input and storm events. The formation extends from western Poland to the Ringkøbing-Fyn-Møn-Arkona High, interfingering with the Ciechocinek Formation in the east. The Grimmen Formation's biostratigraphy is primarily based on ammonite zones, with the falciferum Zone (elegantulum, exaratum, elegans, falciferum subzones) for the lower Toarcian . The type section spans the Pliensbachian -Toarcian boundary (tenuicostatum Subzone) to the elegans Subzone, with key fossils preserved in concretions and clays .
Reinberg Member biota Ammonites define subzones, with Dactylioceras semicelatum and possible D. tenuicostatum at the boundary, followed by Lobolytoceras siemensi and Tiltoniceras antiquum in the Lehmhagen Member (semicelatum Subzone). Elegantulum concretions yield abundant Eleganticeras elegantulum and rare Hildaites murleyi , while exaratum concretions in the Reinberg Member contain Cleviceras exaratum , Phylloceras heterophyllum , and Lytoceras crenatum . Belemnites (Passaloteuthis bisulcata ) are common in the semicelatum Subzone but absent from the elegantulum Subzone due to anoxia . Teuthoids and coleoids occur in elegantulum and exaratum concretions .
Holoplanktonic gastropod Coelodiscus minutus and larval 'Inoceramus ' dubius are abundant, with adults in elegantulum and exaratum concretions , often in regurgitalites . Ostracods and foraminifera are absent in black shales but occur above the Dörnten Member. Calcareous nannoplankton (e.g., Rhombolithion ) are diverse in the elegantulum Subzone, correlating with NJ5b–NJ6 zones. Palynofacies show a dinoflagellate cyst blackout in the elegantulum-exaratum Subzones, with amorphous organic matter (AOM) dominance and micro-charcoal. Megaspores (e.g., Paxillitriletes phyllicus ) from Fennoscandia indicate a humid climate .
Reinberg Member Fossil Record The Reinberg Member, notably at Grimmen and Dobbertin, is a Konservatlagerstätte with exceptional marine and terrestrial fossils in exaratum concretions . Ammonites (e.g., Cleviceras exaratum ), Coelodiscus minutus , and 'Inoceramus' dubius are preserved in calcite , with predation traces. Decapod crustaceans (e.g., Palaeoastacus -like), coleoid gladii, and fish (leptolepids, Grimmenodon aureum ) occur in calcium phosphate . Coprolites , likely from fish , sharks , and marine reptiles (e.g., ichthyosaurs , plesiosaurs ), are common. Terrestrial insects , especially in exaratum concretions types 1 and 2, include well-preserved wings and rare 3D specimens, comparable to Dobbertin's fauna. These fossils, formed in anoxic , fine-grained limestone below the storm wave base , reflect a nutrient-rich environment fertilized by insect carcasses.
Biota Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon Notes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon Morphotaxon
Foraminifera Genus Species Location Material Notes Images Berthelinella B. paradoxa Dobbertin Tests Foraminiferan of the family PlectofrondiculariidaeBolivina B. liasica Dobbertin Tests Foraminiferan of the family Bolivinidae Cornuspira C. orbicula Dobbertin Tests Foraminiferan of the subfamily Cornuspirinae Dentalina D. deslongchampsi D. primaeva D. pseudocommunis D. submucronata D. subtenuicollis D. tenuistriata D. torta D. vetusta Dobbertin Tests Foraminiferan of the family Nodosariidae Eoguttulina E. bilocularis E. liassica E. cf.simplex Dobbertin Tests Foraminiferan of the subfamily Polymorphininae Falsopalmula F. insignis Dobbertin Tests Foraminiferan of the family Robuloididae Frondicularia F. bicostata F. sulcata F. terquemi F . sp.Dobbertin Tests Foraminiferan of the family Nodosariidae Glomospira G. gordialis G . sp.Dobbertin Tests Foraminiferan of the subfamily Usbekistaniinae Lagena L. globosa L. oxystoma L. globosa L. tenuicostata L . sp.Dobbertin Tests Foraminiferan of the family Nodosariidae Lenticulina L. acutiangulata L. müensteri L. varians L. sublaevis Dobbertin Tests Foraminiferan of the family Vaginulinidae Lingulina L. tenera Dobbertin Tests Foraminiferan of the subfamily Lingulininae Marginulina M. prima M. simplex M. oolithica Dobbertin Tests Foraminiferan of the subfamily Marginulininae Nodosaria N. dispar N. agglutinans N. elongata N. germanica N. hirsuta N. dispar N. mitis N. regularis N. simplex Dobbertin Tests Foraminiferan of the family Nodosariidae Ophtalmidium O. orbiculare O. carinatum O . sp.Dobbertin Tests Foraminiferan of the family Ophthalmidiidae Rectoglandulina R. oviformis R. sexcostata Dobbertin Tests Foraminiferan of the family Nodosariidae Stilostomella S .? sp.Dobbertin Tests Foraminiferan of the family Stilostomellidae Trochamminoides T . sp.Dobbertin Tests Foraminiferan of the family Trochamminoidae Turrispirillina T. conoidea Dobbertin Tests Foraminiferan of the family Spirillinidae Vaginulina V. anomala Dobbertin Tests Foraminiferan of the subfamily Vaginulininae
Ichnofossils Genus Species Type Location Origin Images Chondrites C. bollensis Fodinichnia Dobbertin Clay Pit Sipunculans Polychaetes Enteropneustans EchiuransIllustration of Chondrites bollensis Megagrapton M . isp.Pascichnia Dobbertin Clay Pit Polychaetes Sipunculans Enteropneustans Echiurans Planolites P. montanus P. beverleyensis Pascichnia Kozłowice Clay Pit "Boroszów" Brickyard Polychaetes Insects Example of Planolites fossil Skolithos S . isp.Domichnia Müssentin gravel pit Polychaetes Phoronidans Taenidium T. serpentinum Fodinichnia Dobbertin Clay Pit Deposit-feeding Sipuncula Polychaetes Phoronidans
Brachiopoda Genus Species Stratigraphic position Material Notes Images Discina D. cornucopiae D. papyracea D . sp.Klein Lehmhagen pit, Grimmen Dobbertin Shells A saltwater brachiopod , member of Discinidae inside Discinida .
Bivalvia Genus Species Stratigraphic position Material Notes Images Bositra B. buchii B. radiata Klein Lehmhagen pit, Grimmen Dobbertin Ahrensburg Shells A saltwater oyster , member of Posidoniidae. Specimens on a matrix Goniomya G. rhombifera Klein Lehmhagen pit, Grimmen Dobbertin Shells A saltwater clam , member of Pholadomyidae . Gresslya G . sp.Ahrensburg Shells A saltwater clam, member of Ceratomyidae. Lucina L. plana Klein Lehmhagen pit, Grimmen Dobbertin Shells A saltwater clam, member of Lucinidae . Lucina Meleagrinella M. substriata Klein Lehmhagen pit, Grimmen Dobbertin clay pit Shells A saltwater scallop , member of Oxytomidae. Meleagrinella Nucula N. caecilia Dobbertin Shells A saltwater clam, member of Nuculidae . Nucula Parainoceramya P. dubia P. amygdaloides P . sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Shells A saltwater/brackish clam, member of Inoceramidae . Parainoceramya Parallelodon P. cypriniformis Ahrensburg Shells A saltwater clam, member of Parallelodontidae . Stalagmina S. koeneni Klein Lehmhagen pit, Grimmen Dobbertin Shells A saltwater clam, member of Inoceramidae. Steinmannia S. bronni S. radiata Klein Lehmhagen pit, Grimmen Dobbertin Shells A saltwater oyster, member of Posidoniidae. Specimens on a matrix Tancredia T. elegans Ahrensburg Shells A saltwater clam, member of Tancrediidae.
Gastropoda Genus Species Stratigraphic position Material Notes Images Cerithium C. armatum C. roeceri Klein Lehmhagen pit, Grimmen Shells A sea snail , member of Cerithiinae. Cerithium Coelodiscus C. minutus Clay pit of Dobbertin Klein Lehmhagen pit, Grimmen Müssentin gravel pit Ahrensburg Shells A holoplanktonic snail, type member of the family Coelodiscidae Turritella T . sp.Dobbertin Shells A sea snail, member of Turritellidae .
Cephalopoda Genus Species Stratigraphic position Material Notes Images Acrocoelites A. pyramidalis Klein Lehmhagen pit, Grimmen Müssentin gravel pit Multiple specimens. A Belemnoidea member of the family Megateuthididae Acrocoelites Beloteuthis B. schubleri Klein Lehmhagen pit, Grimmen Multiple specimens A Mesoteuthoidea, member of the family Beloteuthidae. Beloteuthis Dactylioceras D. athleticum D. temperatum D. attenuatum D. tenuicostatum D. semicelatum D. cf. aequistriatum D. eickenbergi D. ernsti Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Shells An ammonite , type member of the subfamily Dactylioceratinae . Dactylioceras Eleganticeras E. elegantulum E . sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Shells An ammonite, member of the family Hildoceratidae . Eleganticeras Hildaites H. subserpentinus Ahrensburg Shells An ammonite, member of the family Hildoceratidae. Jeletzkyteuthis J. coriaceus Klein Lehmhagen pit, Grimmen Two specimens A Loligosepiidae Loligosepiida (Vampyromorpha ). Lobolytoceras L. siemensi L . sp.Klein Lehmhagen pit, Grimmen Dobbertin Thüringen Brandenburg Braunschweig Boulder from Ahrensburg Shells An ammonite, member of the family Lytoceratidae . Lobolytoceras Nodicoeloceras N. crassoides Ahrensburg Shells An ammonite, type member of the subfamily Dactylioceratinae. Peronoceras P. fibulatum Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Shells An ammonite, member of the subfamily Dactylioceratinae. Pseudolioceras P. lythense Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Shells An ammonite, member of the subfamily Harpoceratinae . Pseudolioceras Phylloceras P. heterophyllum Ahrensburg Shells An ammonite, type member of the subfamily Phylloceratinae. Dactylioceras Teudopsis T. bollensis Klein Lehmhagen pit, Grimmen Few specimens A vampire squid, member of Teudopsidae. Teudopsis Tiltoniceras T. antiquum T. costatum T. acutum T. schroederi T. costatum T. capillatum T . sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Shells An ammonite, member of the family Hildoceratidae. Tiltoniceras Whitbyiceras W. pingue Ahrensburg Shells An ammonite, member of the family Phymatoceratidae.
Crustacea Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.
Genus Species Stratigraphic position Material Notes Images Bairdia B. hilda Dobbertin Valves A marine/brackish ostracod of the family Bairdiidae . Bythocypris B. elongata Dobbertin Valves A marine/brackish ostracod of the family Bythocyprididae . Cytherelloidea Indeterminate Dobbertin Valves A marine/brackish ostracod of the superfamily Cytherelloidea. Euestheria E. opalina E . sp.Klein Lehmhagen pit, Grimmen Dobbertin Valves A freshwater clam shrimp (Phyllopodan ) of the family Lioestheriidae. Originally identified as a bivalve of the genus Posidonia and latter a member of the genus Lioestheria . Glyphea G. liasina Dobbertin Isolated molts Isolated chelae A marine decapod, type member of the family Glypheidae . Glyphea Hungarella H. adenticulata H . sp.Dobbertin Valves A marine/brackish ostracod of the family Healdiidae Paracypris P. semidisca Dobbertin Valves A marine/brackish ostracod of the subfamily Paracypridinae. Polycope P. cincinnata P . sp.Dobbertin Valves A marine/brackish ostracod of the family Polycopidae . Procytheridea P . sp.Dobbertin Valves A marine/brackish ostracod of the family Protocytheridae . Proeryon P. hartmanni cf.P . sp.Klein Lehmhagen pit, Grimmen Dobbertin Isolated molts Single specimen with preserved upper thorax and chelae A marine decapod, member of the subfamily Proeryoninae. Proeryon Trachycythere T. tubulosa Dobbertin Valves A marine/brackish ostracod of incertae sedis affinity
Arachnida Genus Species Stratigraphic position Material Notes Images Seppo S. koponeni Klein Lehmhagen pit, Grimmen Single incomplete specimen A spider , possible member of the superfamily Palpimanoidea . It is the first confirmed spider from the lower Jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action. Probably a ground-dwelling predator that hunted the abundant insect fauna present on the layers.
Insecta Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation. In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of Eleganticeras elegantulum macrochonchs and in fish coprolites which are the most frequent fossils at all. In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex. On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.
Notoptera Genus Species Stratigraphic position Material Notes Images Geinitzia G. schlieffeni G. debilis G. minor Dobbertin, Mecklenburg (Greifswald collection) Specimens A notopteran, member of the family Geinitziidae . Nele N. jurassica Klein Lehmhagen pit, Grimmen Specimens A notopteran, member of the family Bajanzhargalanidae . Grylloblatta , extant ice crawler, Nele was probably similarProsepididontus P. calopteryx Dobbertin, Mecklenburg (Greifswald collection) Specimens A notopteran, member of Geinitziidae.
Odonatoptera Genus Species Stratigraphic position Material Notes Images Obotritagrion O. tenuiformum O. petersi Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Specimens An odonatopteran, member of the family Protomyrmeleontidae. Protomyrmeleon P. brunonis P. quadriordinum Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens
Odonata Genus Species Stratigraphic position Material Notes Images Anisozygopteron A. geinitzianum Dobbertin, Mecklenburg (Greifswald collection) Specimens A dragonfly, member of the family Myopophlebiidae. Anomothemis A. brevistigma Dobbertin, Mecklenburg (Greifswald collection) Specimens incertae sedis Archithemis A. brodiei Dobbertin, Mecklenburg (Greifswald collection) Specimens Dialothemis D. dubia Dobbertin, Mecklenburg (Greifswald collection) Specimens A dragonfly, member of the family Selenothemistidae. Eosagrion E. risi Dobbertin, Mecklenburg (Greifswald collection) Specimens A dragonfly, member of the family Eosagrionidae. Grimmenopteron G. elegantulum Klein Lehmhagen pit, Grimmen Specimens A dragonfly, member of the family Liassophlebiidae. Heterophlebia H. buckmani H. megapolitana Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Specimens A dragonfly, member of the family Heterophlebiidae. Heterothemis H. brodiei H. germanica Dobbertin, Mecklenburg (Greifswald collection) Ahrensburg erratics Specimens A dragonfly, member of the family Liassogomphidae. Liadothemis L. hydrodictyon L. major L. geinitzi L. insignis Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens Incertae sedis . Magnasupplephlebia represents a very large dragonfly, with a wingspan of 13 cm. Other odonatan remains are unable to being referred to a concrete group due to their incomplete status.Magnasupplephlebia M. kallweita M. intercalaria Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Specimens Parelthothemis P. dobbertinensis Dobbertin, Mecklenburg (Greifswald collection) Specimens Petrothemis P. singularis Dobbertin, Mecklenburg (Greifswald collection) Specimens Pycnothemis P. densa Dobbertin, Mecklenburg (Greifswald collection) Specimens Rhabdothemis R. strigivena Dobbertin, Mecklenburg (Greifswald collection) Specimens Selenothemis S. liadis Dobbertin, Mecklenburg (NHMW collection) Specimens A dragonfly, type member of the family Selenothemistidae. Temnostigma T. singulare Dobbertin, Mecklenburg (Greifswald collection) Specimens Incertae sedis .Trigonophlebia T. zessini Klein Lehmhagen pit, Grimmen Specimens A dragonfly, member of the family Heterophlebiidae. Turanopteron T. pommerana Klein Lehmhagen pit, Grimmen Specimens A dragonfly, member of the family Asiopteridae. Sphenophlebia pommerana is a junior synonym.
Paraneoptera Genus Species Stratigraphic position Material Notes Images Archipsylla A. primitiva Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A paraneopteran, member of the family Archipsyllidae . Liassopsocus L. lanceolatus Klein Lehmhagen pit, Grimmen Specimens A paraneopteran, member of the family Psocidiidae.
Eoblattida Genus Species Stratigraphic position Material Notes Images Griphopteron G. molle Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens An Eoblattidan , member of the family Blattogryllidae.
Thysanoptera Genus Species Stratigraphic position Material Notes Images Liadoptilia L. misera Klein Lehmhagen pit, Grimmen Specimens Thysanopterans, members of the family Lophioneurida . Aeroplankton is extraordinarily well preserved in Grimmen, with the most abundant representatives of the aeroplankton (around 3 mm) being Lophioneurids, specially Undacypha europaea . Extant Thysanopteran, extinct genera were probably similar Trichorthophlebia T. pilifera Dobbertin, Mecklenburg (Greifswald collection) Undacypha U. europaea Dobbertin, Mecklenburg (Greifswald collection)
Plecoptera Genus Species Stratigraphic position Material Notes Images Dicronemoura D. furcata Klein Lehmhagen pit, Grimmen Specimens A stonefly, member of the family Perlariopseidae. Dobbertiniopteryx D. capniomimus Dobbertin, Mecklenburg (Ansorge coll) Specimens A winter stonefly, member of the family Capniidae . extant Capniidae, Dobbertiniopteryx was probably similar
Orthoptera Genus Species Stratigraphic position Material Notes Images Locustopsis L. gyra L. bernstorffi L. sippeli L. dubia L. bernstorffi L. nana L. reducta L. elongata L. elegans L. pulchella L. procera L. mecklenburgica L. maculosa L . sp 1 L . sp 2 L . sp 3 L . sp 4Grimmen (Krempien collection) Lehmhagen, near Grimmen (Zessin collection) Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens Grasshoppers of the family Locustopsidae .Locustophanes L. rhipidophorus Dobbertin, Mecklenburg (Greifswald collection) Specimens A locust , type member of the family Locustidae . Acanthacris , extant Locust, Locustophanes was probably similarPanorpidium P. geinitzi P. magna P. minima P. media P. reticulata P. mesostena Grimmen (Krempien collection) Lehmhagen, near Grimmen (Zessin collection) Klein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (Zessin collection) Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A grasshopper, member of the family Elcanidae . Parelcana P. tenuis Klein Lehmhagen pit, Grimmen Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection) Specimens Plesioschwinzia P. thalassophila P. lacera Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens Grasshoppers of the family Locustopsidae. Protochaeta P. masculina P. lanceolata Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A grasshopper, member of the family Regiatidae . Protogryllus P. dobbertinensis P. germanicus P . sp. P. vicinus P. liadis P. major P. stenobasis P. femina P. acutipennis P. minor P. irregularis Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (BMNH collection) Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection) Ahrensburg Erratics Specimens A cricket -like grasshopper, type member of the family Protogryllidae . Schwinzia S. sola Dobbertin, Mecklenburg (Zessin collection) Specimens Grasshoppers of the family Locustopsidae. Synelcana S. muelleri Dobbertin, Mecklenburg (Zessin collection) Specimens A grasshopper, member of the family Elcanidae. Zalmonites Z. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens A grasshopper, incertae sedis.
Phasmatodea Genus Species Stratigraphic position Material Notes Images Durnovaria D. cf. parallela Klein Lehmhagen pit, Grimmen Specimens A stick insect , member of the family Aerophasmidae. Example of extant Phasmatodean , Durnovaria was probably similar
Blattodea Genus Species Stratigraphic position Material Notes Images Blattula B. langfeldti B. dubia Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Ahrensburg Erratics Specimens A cockroach , member of the family Blattulidae . Caloblattina C. mathildae Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A cockroach, type member of the family Caloblattinidae. Dipluroblattina D. scudderi Dobbertin, Mecklenburg (Greifswald collection) Specimens A cockroach, member of the family Blattulidae. Eublattula E. crassivena Dobbertin, Mecklenburg (Greifswald collection) Specimens Liadoblattina L. blakei Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A carnivorous cockroach, type member of the family Raphidiomimidae. Mesoblattina M. protypa Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A cockroach, type member of the family Mesoblattinidae .
Hemiptera Genus Species Stratigraphic position Material Notes Images Acromocoris A. similis Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens A water boatman, member of the family Corixidae . Example of extant Corixidae, Acromocoris was probably similar Anosmus A. spilopterus Dobbertin, Mecklenburg (Greifswald collection) Specimens Shore bugs, member of the family Archegocimicidae. Aphlebocoris A. punctata A. nana Dobbertin, Mecklenburg (Greifswald collection) Specimens A creeping water bug, member of the family Naucoridae . Example of extant Naucoridae, extinct genera where probably similar Apopnus A. magniclavus Dobbertin, Mecklenburg (Greifswald collection) Apsicoria Apsicoria semideleta Dobbertin, Mecklenburg (Greifswald collection) Specimens True bugs, member of the family Pachymeridiidae. Ardela A. grimmenensis Klein Lehmhagen pit, Grimmen Specimens Leafhoppers of the family Archijassidae . Archegocimex A. geinitzi A. primitiva A. liadis Dobbertin, Mecklenburg (Greifswald collection) Specimens Shore bugs, member of the family Archegocimicidae. Example of extant Pentacora , Archegocimicidae genera where probably similar Archicercopis A. falcatus Dobbertin, Mecklenburg (Greifswald collection) Specimens Moss bugs of the family Progonocimicidae . Archijassus Archijassus heeri Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Leafhoppers of the family Archijassidae. Example of extant Membracidae , Archijassidae genera where probably similar Aphidulum A. ciliatum Klein Lehmhagen pit, Grimmen Specimens A Sternorrhynchan, member of the family Protopsyllidiidae. Archiconiopteryx A. liasina Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A Stemorrhynchan, type member of the family Archiconiopterygidae. Cathalus Cathalus alutaceus Dobbertin, Mecklenburg (Greifswald collection) Specimens True bugs, member of the family Pachymeridiidae. Cixiites C. liasinus Dobbertin, Mecklenburg (Greifswald collection) Specimens Planthoppers of the family Fulgoridiidae .Cuneocoris C. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens A true bug, member of the family Cuneocoridae. Diatillus Diatillus debilis Dobbertin, Mecklenburg (Greifswald collection) Specimens Shore bugs, member of the family Archegocimicidae. Eocercopis E. ancyloptera E. similis Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens Moss bugs of the family Progonocimicidae. Fulgoridium F. egens F. lapideum F. dubium F. latum F. venosum F. pallidum F. geinitzi F. spilographum F. oligoneurum F. regulare F. graphipterum F. inconspicuum F. quadrisignatum F. breviradiatum F. modestum F. anale F. marginepunctatum F. picturatum F. anomalum F. stigmaticum F. punctatum F. mortuum F. alatum F. nodosum F. megapolitanum F. pulchrum F. remotum F. multivenosum F. debile F. trifurcatum F. litorale F. parvispilum F. retractum F. polyneurum F. oligospilum F. nubeculum F. obtusum F. grave F. fenestratum F. breve F. dilutum F. multipunctatum F. ampliatum F. clavatum F. nebulosum F. vicinum F. curvipenne F. elegantulum F. defunctum F. plicatum F. exhumatum F. angulosum F. acutum F. rotundatum F. ancylla F. bifurcatum F. inaequale F. brachyptilum F. intercalatum F. balticum Grimmen (EMAU collection) Klein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens Planthoppers of the family Fulgoridiidae.Example of extant Fulgoridae , Fulgoridiidae genera where probably similar Grimmenaphis G. magnifica Klein Lehmhagen pit, Grimmen Specimens An aphid, member of the family Oviparosiphidae . Example of extant Aphidoidea , Oviparosiphidae genera where probably similar Hadrocoris H. scutellaris Dobbertin, Mecklenburg (Greifswald collection) Specimens A true bug, member of the family Hadrocoridae. Hypocimex Hypocimex membranaceus Dobbertin, Mecklenburg (Greifswald collection) Specimens True bugs, member of the family Pachymeridiidae. Liadopsylla L. geinitzi L. obtusa Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A jumping plant louse, member of the family Liadopsyllidae . Liassocercopis L. schnicki Klein Lehmhagen pit, Grimmen Specimens A cicadomorphan, member of the family Hylicellidae. Liasocoris L. hainmuelleri Schwinzer brickyard, near Goldberg Specimens A true bug, incertae sedis inside Hemiptera . Margaroptilon M. germanicum Dobbertin, Mecklenburg (Greifswald collection) Specimens Planthoppers of the family Fulgoridiidae.Mesoledra M. pachyneura Klein Lehmhagen pit, Grimmen Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection) Specimens Leafhoppers of the family Archijassidae. Metafulgoridium M. spilotum M. ampliatum M. graptum Dobbertin, Mecklenburg (Greifswald collection) Specimens Planthoppers of the family Fulgoridiidae.Pachymeridium P. dubium Dobbertin, Mecklenburg (Greifswald collection) Specimens True bugs, member of the family Pachymeridiidae. Probascanion P. megacephalum Dobbertin, Mecklenburg (NHMW collection) Specimens A Coleorrhynchan, member of the family Probascanionidae. Procercopis P. alutacea P. coriacea P. liasina P. similis P. jurassica Dobbertin, Mecklenburg (Greifswald collection) Specimens Froghoppers of the family Procercopidae . Procercopina P. frenzeli Klein Lehmhagen pit, Grimmen Specimens Progonocimex P. jurassicus P. liasinus Dobbertin, Mecklenburg (Greifswald collection) Specimens Moss bugs of the family Progonocimicidae. Example of extant Peloridiidae , Progonocimicidae genera where probably similar Progonocoris P. pictus Dobbertin, Mecklenburg (Greifswald collection) Specimens Shore bugs, member of the family Archegocimicidae. Sisyrocoris S. rudis Dobbertin, Mecklenburg (Greifswald collection) Specimens True bugs, member of the family Pachymeridiidae. Toarcopsyllidium T. ernsti Klein Lehmhagen pit, Grimmen Isolated Wings A Sternorrhynchan, member of the family Protopsyllidiidae.
Hymenoptera Genus Species Stratigraphic position Material Notes Images Brigittepterus B. brauckmanni Dobbertin, Mecklenburg (Zessin collection) Isolated Wings Wasps, member of the family Ephialtitidae. Grimmaratavites G. mirabilis Klein Lehmhagen pit, Grimmen Specimens A Wood Wasp, member of the family Karatavitidae . Liadobracona L. raduhna Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Isolated Wings Wasps, member of the family Ephialtitidae. Xyelula X. benderi Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Specimens A pseudo-Wasp , member of the family Sepulcidae .
Megaloptera Genus Species Stratigraphic position Material Notes Images Dobbertinia D. reticulata Dobbertin, Mecklenburg (Greifswald collection) Specimens An alderfly, member of the family Sialidae . Example of extant Sialidae , Dobbertinia was probably similar
Neuroptera Genus Species Stratigraphic position Material Notes Images Actinophlebia A. megapolitana A. aenea A. parallela Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Lacewings of the family Prohemerobiidae Apeirophlebia A. grandis Dobbertin, Mecklenburg (Greifswald collection) Specimens A silky lacewing, member of the family Psychopsidae . Example of extant Psychopsidae, Apeirophlebia was probably similar Epigambria E. longipennis Dobbertin, Mecklenburg (Greifswald collection) Specimens A lacewing, member of the family Epigambriidae. Example of extant Ithonidae , Epigambriidae members where probably similar Liassochrysa L. stigmatica Dobbertin, Mecklenburg (Ansorge coll) Specimens A Neuropteran of the family Mantispidae . Example of extant Mantispidae, Liassochrysa was probably similar Loxophleps L. costalis Dobbertin, Mecklenburg (Greifswald collection) Specimens A lacewing, incertae sedis inside Neuroptera . Melaneimon M. dubium Dobbertin, Mecklenburg (Greifswald collection) Specimens Melamnous M. indistinctus Dobbertin, Mecklenburg (Greifswald collection) Specimens Mesoleon M. dobbertinianus Dobbertin, Mecklenburg (Greifswald collection) Specimens Mesosmylina M. falcifera Klein Lehmhagen pit, Grimmen Specimens A lance lacewing, member of the family Osmylidae . Polyosmylus P. excelsus Klein Lehmhagen pit, Grimmen Specimens A lacewing, member of the family Epigambriidae. Prohemerobius P. prodromus P. ovatus P. anomalus P. costalis P. persimilis P. geinitzi P. liasinus P. major P. parvulus P. geinitzianus P. dilaroides P. chryseus P. latus Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Lacewings of the family Prohemerobiidae. Solenoptilon S. kochi Dobbertin, Mecklenburg (Greifswald collection) Specimens A lacewing, type member of the family Solenoptilidae. Toarciconiopteryx T. dipterosimilis Klein Lehmhagen pit, Grimmen Specimens A dustywing, member of the subfamily Toarciconiopteryginae. Example of extant Coniopterygidae , Toarciconiopteryx was probably similar
Coleoptera Genus Species Stratigraphic position Material Notes Images Allognosis A. nitens Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera . Anancaeon A. microcephalum Dobbertin, Mecklenburg (NHMW collection) Specimens Anomerus A. punctifer Dobbertin, Mecklenburg (Greifswald collection) Specimens Anypostatus A. taurus Dobbertin, Mecklenburg (Greifswald collection) Specimens Apioderes A. punctatus Dobbertin, Mecklenburg (Greifswald collection) Specimens Apsychus A. alutaceus Dobbertin, Mecklenburg (Greifswald collection) Specimens Bareus B. strigipennis Dobbertin, Mecklenburg (Greifswald collection) Specimens Bathygerus B. bellus B. divergens Dobbertin, Mecklenburg (Greifswald collection) Specimens Carabites C. dubius C. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens A ground beetle, member of the family Carabidae . Example of extant Carabidae , Carabites was probably similar Clinomerus C. laticollis Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Coptogyrinus C. scutellatus Dobbertin, Mecklenburg (Greifswald collection) Specimens A whirligig beetle, member of the family Gyrinidae . Example of extant Gyrinidae , Coptogyrinus was probably similar Ecthlimma E. forficuloides Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Enamma E. striatum Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Eurynucha E. pseudobuprestis Dobbertin, Mecklenburg (Greifswald collection) Specimens A jewel beetle, member of the family Buprestidae . Example of extant Buprestidae , Eurynucha was probably similar Gyrinulopsis G. nanu Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Hydrobiites H. dobbertinensis H. punctatostriatus Dobbertin, Mecklenburg (NHMW collection) Specimens A beetle, member of the family Permosynidae . Keleusticus K. zirkeli Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Liassodites L. obsti Klein Lehmhagen pit, Grimmen Specimens A False Ground Beetle of the family Trachypachidae . Extant member of Trachypachidae , Liassodites was probably similar Masselytron M. quinquestriatum Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Nannoodes N. pseudocistela Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Nebrioides N. dobbertinensis Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Onkedodimus O. discicollis Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Oxytoroptera O. mediocris Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Paracurculium P. punctatum Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Parnidium P. frechi P. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Periboloptera P. rotunda Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Plastobuprestites P. elegans Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Plastonebria P. scudderi Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Polypamon P. byrrhoides Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Pseudocyphon P. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Pseudoprionites P. liasinus Dobbertin, Mecklenburg (Greifswald collection) Specimens Beetles, incertae sedis inside Coleoptera. Thoracotes T. dubius Dobbertin, Mecklenburg (Greifswald collection) Specimens A bark gnawing beetle, member of the family Trogossitidae . Example of extant Trogossitidae , Thoracotes was probably similar
Trichoptera Genus Species Stratigraphic position Material Notes Images Archiptilia A. ovata Dobbertin, Mecklenburg (Greifswald collection) Specimens Caddisflies, member of the family Necrotauliidae . Epididontus E. geinitzianus Dobbertin, Mecklenburg (Greifswald collection) Specimens Liadotaulius L. maior Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A finger-net caddisfly, member of the family Philopotamidae . Example of extant Philopotamidae , Liadotaulius was probably similar Mesotrichopteridium M. intermedium M. pusillum Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens Caddisflies, member of the family Necrotauliidae. Metarchitaulius M. longus Dobbertin, Mecklenburg (Greifswald collection) Specimens Nannotrichopteron N. gracile Dobbertin, Mecklenburg (Greifswald collection) Specimens Necrotaulius N. parvulus Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens Palaeotaulius P. vicinus Dobbertin, Mecklenburg (Greifswald collection) Specimens Parataulius P. jurassicus Dobbertin, Mecklenburg (Greifswald collection) Specimens Paratrichopteridium P. efossum P. costale Dobbertin, Mecklenburg (Greifswald collection) Specimens Pararchitaulius P. ovalis Dobbertin, Mecklenburg (Greifswald collection) Specimens Pseudorthophlebia P. platyptera Dobbertin, Mecklenburg (Greifswald collection) Specimens
Lepidoptera Genus Species Stratigraphic position Material Notes Images Eolepidopterigidae Eolepidopterigidae indet. A Eolepidopterigidae indet. B Eolepidopterigidae indet. C Klein Lehmhagen pit, Grimmen Isolated wings Isolated wing scales A moth, member of the family Eolepidopterigidae.
Mecoptera Genus Species Stratigraphic position Material Notes Images Mesobittacus M. minutus Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Hangingflies, member of the family Bittacidae . Mesochorista M. sinuata Dobbertin, Mecklenburg (Greifswald collection) Specimens A Scorpionfly, member of the family Permochoristidae. Mesopanorpa M. maculata Dobbertin, Mecklenburg (Greifswald collection) Specimens Hangingflies, member of the family Orthophlebiidae . Neorthophlebia N. maculipennis N. megapolitana N. pallida N. debilis N. simillima N. medialis N. stigmatica N. acutipennis Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Hangingflies, member of the family Bittacidae. Example of extant Bittacidae, extinct members where probably similar Orthophlebia O. germanica O. elongata O. fuscipennis O. limnophila O. cf. limnophila O. radialis O. liadis O. maculata O. vicina O. brunsvicensis Klein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (Greifswald collection) Specimens Hangingflies, member of the family Orthophlebiidae. Parabittacus P. analis P. lingula Dobbertin, Mecklenburg (Greifswald collection) Specimens Hangingflies, member of the family Bittacidae. Pseudopolycentropus P. triangularis P. perlaeformis P. sp. Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A scorpionfly, type member of the family Pseudopolycentropodidae .
Diptera Genus Species Stratigraphic position Material Notes Images Aenne A. liasina Schönenwald railway cutting, Grimmen Klein Lehmhagen pit, Grimmen Specimens The oldest known non biting midgess , members of the Chironomidae Example of extant Chironomidae, extinct members where probably similar Antefungivora A. germanica Klein Lehmhagen pit, Grimmen Specimens Flies , member of the family Antefungivoridae .Archibio A. mycetophilinus Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Archipleciomima A. germanica A. sp. Klein Lehmhagen pit, Grimmen Specimens A fly , incertae sedis inside Diptera Archirhyphus A. geinitzi Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Flies , member of the family Protorhyphidae.Architipula A. stigmatica A. latipennis A. elegans A. seebachi A. nana A. pusilla A. pulla A. debilis A. simplex A. intermedia A. obliqua A. areolata A. maior A. conspicua A. clara A. dubia Dobbertin, Mecklenburg (Greifswald collection) Specimens A crane fly of the family Limoniidae . Example of extant Limoniidae, extinct members where probably similar Crenoptychoptera C. dobbertinensis Dobbertin, Mecklenburg (Ansorge coll) Specimens A Phantom crane fly, member of the family Ptychopteridae . Example of extant Ptychopteridae, extinct members where probably similar Eoditomyia E. primitiva Klein Lehmhagen pit, Grimmen Specimens A fly, type member of the family Eoditomyidae. Eolimnobia E. geinitzi Dobbertin, Mecklenburg (Greifswald collection) Specimens A phantom crane fly, member of the family Ptychopteridae. Eoptychoptera E. simplex E. eximia Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens Eotipula E. defuncta E. mortua E. coarctata Dobbertin, Mecklenburg (Greifswald collection) Specimens A crane fly of the family Limoniidae. Grimmenia G. prima G. secunda Lehmhagen, near Grimmen (Zessin collection) Specimens A crane fly of the family Limoniidae. Grimmyia G. baltica Klein Lehmhagen pit, Grimmen Specimens A snipe fly, member of the family Rhagionidae . Example of extant Rhagionidae, extinct members where probably similar Heterorhyphus H. triangularis Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A fly, type member of the family Heterorhyphidae. Lehmhagenia L. ansorgei Klein Lehmhagen pit, Grimmen Specimens A fly, member of the family Protobrachyceridae. Liassobrachyceron L. kotejai Dobbertin, Mecklenburg (Ansorge coll) Specimens A snipe fly, member of the family Rhagionidae. Liassopsychodina L. pommerana Klein Lehmhagen pit, Grimmen Specimens A Moth fly , member of the family Psychodidae . Example of extant Psychodidae, extinct members where probably similar Mailotrichocera M. mikereichi M. variabilis M. zessini Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Specimens A Winter Crane Fly, member of the family Trichoceridae . Example of extant Trichoceridae, extinct members where probably similar Mesorhyphus M. nanus Dobbertin, Mecklenburg (Greifswald collection) Specimens A wood gnat, member of the family Anisopodidae . Example of extant Anisopodidae, extinct members where probably similar Mesotipula M. siggiae M. vicina M. cf. vicina M. mirabilis M. curvata M. geinitzi M. minuta M. lapidaria M. parva Lehmhagen, near Grimmen (Zessin collection) Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection) Specimens A Crane Fly of the family Limoniidae. Metatrichopteridium M. confusum Dobbertin, Mecklenburg (Greifswald collection) Specimens A fly, member of the family Hennigmatidae. Nannotanyderus N. grimmenensis N. krzeminskii Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Specimens A primitive Crane Fly , member of the family Tanyderidae . Oryctochlus O. toarciensis Klein Lehmhagen pit, Grimmen Specimens The oldest known non biting midges, members of the Chironomidae. Rhaetomyia R. herrigi Dobbertin, Mecklenburg (Greifswald collection) Specimens A Phantom midge of the family Chaoboridae . Example of extant Chaoboridae, extinct members where probably similar Palaeobrachyceron P. willmanni Klein Lehmhagen pit, Grimmen Specimens A Snipe fly, member of the family Rhagionidae. Phryganidium P. minimum Dobbertin, Mecklenburg (Greifswald collection) Specimens A Crane Fly of the family Limoniidae. Pleciofungivora P. liassica Klein Lehmhagen pit, Grimmen Specimens A fly, member of the family Pleciofungivoridae. Podonomius P. tumidus Klein Lehmhagen pit, Grimmen Specimens The oldest known non biting midges, members of the Chironomidae. Praemacrochile P. decipiens P. dobbertinensis Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Specimens A primitive Crane Fly , member of the family Tanyderidae. Protobrachyceron P. liasinum Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Specimens A fly, member of the family Protobrachyceridae. Protoplecia P. klafacki P. liasina Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens A fly, type member of the family Protopleciidae. Protorhyphus P. simplex Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection) Specimens Flies , member of the family Protorhyphidae.Tanypsycha T. connexa Klein Lehmhagen pit, Grimmen Specimens A Moth fly , member of the family Psychodidae.
Echinodermata In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%. In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.
Genus Species Stratigraphic position Material Notes Images Crinoidea Indeterminate Dobbertin Columnar ossicles Incertae sedis Holothuriida Indeterminate Dobbertin Holothurian wheels Incertae sedis Sinosura S. brodiei S . spp.Dobbertin Isolated and semiarticulated specimens A brittle star, member of Ophioleucidae inside Ophioleucida. Identified as Ophiura longivertebralis .
Vertebrates Fishes Genus Species Stratigraphic position Material Notes Images Dapedium D. pholidotum Dobbertin Incomplete skull Member of the Dapediidae . Dapedium Grimmenichthys G. ansorgei Cf. G. ansorgei Klein Lehmhagen pit, Grimmen Articulated, but incompletely preserved specimens Member of the group Pholidophoriformes. This genus was a small 'pholidophoriform' teleost. Grimmenodon G. aureum Klein Lehmhagen pit, Grimmen Almost complete left prearticular with dentition Member of the order Pycnodontiformes . Is of comparatively small size, suggesting a hypothetical small standard length of the fish of about 7–10 cm at the time of death. Gyrosteus G. mirabilis Ahrensburg Erratic Assemblage GPIH 4864, Hyomandibula Member of the Chondrosteidae inside Acipenseriformes . Hybodus H. hauffianus H . spp.Klein Lehmhagen pit, Grimmen Ahrensburg erratic assemblage Fragmentary Fin spine Supposed coprolithes Teeth Member of the Hybodontiformes . Hybodus hauffianus Lepidotes L. buelowianus L. elvensis L . spp.Klein Lehmhagen pit, Grimmen Dobbertin Ahrensburg Erratic Assemblage Various specimens Member of the family Lepidotidae inside Lepisosteiformes . L. buelowianus Lepisosteidae Indeterminate Grimmen GG 439/4-7 scales The oldest confirmed true Gar , member of the Lepisosteidae. Lepisosteus extant example of GarLeptolepidae Indeterminate Former clay pit of Dobbertin Isolated head with missing postcranial bones Assigned Teeth Member of the family Leptolepidae. Leptolepis L. bronni L. coryphaenoides L. normandica L. jaegeri L . spp.Klein Lehmhagen pit, Grimmen Dobbertin clay pit Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade Ahrensburg Slightly disarticulated, incomplete head; Subadult, almost complete; fragmentary head; isolated head with pectoral girdle; incomplete head Isolated remains in form of "Grätensandsteine" Type member of the family Leptolepidae. The most abundant vertebrate recovered on the formation, including 3D preserved specimens, as well, is the main component of the Fishbone sandstones from the upper layers. Leptolepis Lycopteroidarum L . sp. A L . sp. BAhrensburg Erratic Assemblage Otoliths Otoliths of the family Lycopteroidea inside Lycopteriformes. Mengius M. acutidens Klein Lehmhagen pit, Grimmen Schwinzer Ziegelei ca 3 km E of Dobbertin Holotype: Articulated, incomplete fish Referred articulated, semiarticulated and fragmentary specimens Member of the family Lepidotidae inside Lepisosteiformes. Stomach content is also preserved on a specimen from Dobbertin, and is composed by arthropod cuticles. Pachycormiformes Indeterminate Klein Lehmhagen pit, Grimmen Unknown, only cited. Member of the family Pachycormiformes . Pholidophorus P . sp.Ahrensburg Erratic Assemblage Assigned Fragmentary remains Member of the family Pholidophoriformes. Pholidophoriformes Indeterminate Klein Lehmhagen pit, Grimmen Isolated skull roof with articulated ethmoid region, parasphenoid, and partial braincase Member of the family Pholidophoriformes. Resembles Pholidophoroides crenulata and P. limbata . Proleptolepis P . sp.Klein Lehmhagen pit, Grimmen MV 202612, fragmentary head . Assigned Teeth Member of the family Leptolepidae. Specimens attributed to Proleptolepis have previously been reported only from the Sinemurian of western Europe. Saurorhynchus S. hauffi S . sp.Klein Lehmhagen pit, Grimmen Dobbertin Diganostic Incomplete skull Incomplete skull The youngest representative of the family Saurichthyidae . Distinctive for its large jaws, similar to modern Belonidae . Saurorhynchus Tetragonolepis T. semicincta Klein Lehmhagen pit, Grimmen Incomplete specimen Isolated Jaw Member of the family Dapediidae. Tetragonolepis
Amniotes Genus Species Stratigraphic position Material Notes Images Ichthyosauria Indeterminate Klein Lehmhagen pit, Grimmen Ahrensburg Erratic Assemblage Presacral centrum 2 ?sacral centrum Isolated vertebrae Incertae sedis Macrospondylus M. bollensis Klein Lehmhagen pit, Grimmen Incomplete skull and associated osteoderm A marine crocodrylomorph, member of the family Machimosauridae . GG 422/6 can be clearly referred to an immature individual. Macrospondylus Megalosauridae Indeterminate Forst Hagen gravel pit Dorsal vertebrae A theropod, member of the family Megalosauridae . Was referred to Megalosaurus . The vertebrae centrum measures 80 mm, implying a medium-sized theropod (~5 m long). Marshosaurus , example of basal Orionides Mesoeucrocodylia Indeterminate Klein Lehmhagen pit, Grimmen Cervical vertebra A crocodrilian, member of Mesoeucrocodylia . Is considered to come from an osteologically immature individual. Meyerasaurus M . sp.Ahrensburg Erratic Assemblage Incomplete coracoid Associated rib A pliosauroid, member of the family Rhomaleosauridae . Meyerasaurus Mystriosaurus M . sp.Klein Lehmhagen pit, Grimmen Partial rostrum with teeth A marine crocodrylomorph, member of the family Teleosauridae . Is indicative of a juvenile individual, with expected around 200–250 mm long skull and the entire animal about 1.50 m. Neoichthyosauria Indeterminate Clay Pit of Dobbertin Four articulated tail vertebrae. An indeterminate ichthyosaur, has been assigned to the species Stenopterygius longifrons . Ophthalmosauridae ?Indeterminate Klein Lehmhagen pit, Grimmen Partial skull and associated postcranial elements preserved in a concretion It has an expanded basipterygoid process on the basisphenoid, only currently known in members of the Ophthalmosauridae Plesiosauria Indeterminate Klein Lehmhagen pit, Grimmen Dobbertin, falciferum zone Three caudal vertebrae and other postcranial elements Gastralia Phalanx Cervical rib and other postcranial elements Incertae sedis Plesiosauroidea Indeterminate Klein Lehmhagen pit, Grimmen Cervical centrum Caudal centrum Incertae sedis Pterosauria ?Indeterminate Klein Lehmhagen pit, Grimmen GG 510, fragmentary tibiofibular(?) A possible pterosaur bone, with features similar to the tibiofibular of coeval taxon Dorygnathus banthensis. Alternatively it can be another type of bone and belong to stem-turtles, plesiosaurs, dinosaurs or even anurans. Dorygnathus banthensis, know from inner localities, has shared features that may suggest GG 510 is from a PterosaurSeeleyosaurus S .? sp.Ahrensburg Erratic Assemblage Three articulated dorsal vertebrae A plesiosaur, member of the family Plesiosauridae . Seeleysaurus Stenopterygius S. cf. quadriscissus Clay pit of Dobbertin Posterior left half of the cranium. An ichthyosaur of the family Stenopterygiidae . Restoration Teleosauridae Indeterminate Ahrensburg Erratic Assemblage Osteoderms Isolated non described remains Incertae sedis Temnodontosaurus T. cf. platyodon Ahrensburg Erratic Assemblage Skull Coracoid Associated rib fragment An icthyosaur, type member of Temnodontosauridae . Assigned to Ichthyosaurus sp., but also suggested affinities to "Leptopterygius " (= Temnodontosaurus ) platyodon . Restoration
Flora Genus Species Stratigraphic position Material Notes Images Botryococcus B. sp. Grimmen clay pit Freshwater algae Indicates freshwater influx in palynofacies assemblage IV, associated with coastal or deltaic settings. Equisetites E . sp. E. cf. Münsteri E. columnare Glacial Erratic Near Grimmen Dobbertin Ahrensburg area Erratic deposit near Lüttow Isolated stems, rhizomes Equisetalean stems of the family Equisetidae . Dominant in mono-to oligotypic stands near water bodies, indicative of high terrestrial input in palynofacies assemblages I and IV. Erlansonisporites E. sparassis E . sp.Grimmen clay pit Megaspores Affinities with the Selaginellaceae . Transported from northern sources (e.g., Fennoscandia), indicative of humid climates. Hughesisporites H. pustulatus Grimmen clay pit Megaspores Affinities with the Selaginellaceae. Likely transported via suspension load plumes, reflecting freshwater influx. Minerisporites M. institus M. volucris Grimmen clay pit Megaspores Affinities with the Selaginellaceae. Associated with warm, humid conditions and high atmospheric CO2. Pagiophyllum P . sp. P. cf. Kurri Grimmen clay pit Leaves, driftwood Conifer macrofossils of the family Cheirolepidiaceae or Araucariaceae . High phytoclasts and cuticles in palynofacies assemblages I, II, and IV support conifer presence. Paxillitriletes P. phyllicus Grimmen clay pit Reinberg 1E Megaspores Affinities with the Isoetales . Transported from Fennoscandia, indicates humid climate, biostratigraphic marker (Paxillitriletes phyllicus Zone, also seen in the Ciechocinek Formation ) Extant Isoetes , typical example of Isoetales . Paxillitriletes probably come from a similar or a related Plant Striatriletes S. excavatus Grimmen clay pit Megaspores Affinities with the Selaginellaceae. Reflects freshwater influx from northern sources. Trileites T . spp.Grimmen clay pit Megaspores Affinities with lycophytes or ferns. Indicates diverse megaspore flora transported from northern sources.
See also