The Grimmen Formation is a Lower Jurassic (Toarcian) geological formation in northeastern Germany, primarily exposed in the Grimmen and Klein Lehmhagen clay pits and documented in wells such as Reinberg 1E. Formally established in 2025, it was previously part of the informal "Green Series" of the Ciechocinek Formation. It represents a prodelta to brackish-marine depositional system in the eastern North German Basin (NGB), shaped by sea-level changes and the Toarcian anoxic event.

History

Studies of Toarcian strata near Grimmen began in 1874 with the discovery of fossiliferous clays in a railway cutting near Schönenwalde, 5 km north of Grimmen, initially misidentified as Middle Jurassic due to ammonite finds. In 1909, the succession was reclassified as Lower Toarcian (Lias epsilon), distinct from the Posidonia Shale Formation. From 1959 to 1995, the Grimmen clay pit was excavated, revealing a glacially dislocated raft of Liassic clays and sands deformed by Pleistocene ice advances. Exploration wells (e.g., Reinberg 1E, Kb Barth 10, Kb Grambow 5) since the 1950s provided extensive core data, with Reinberg 1E designated as the reference section in 2025. The clay pit, abandoned and water-filled since 1995, was established as the type section in 2025, with 2016 and 2020 excavations refining an 18 m composite log spanning the Pliensbachian-Toarcian boundary to the elegans Subzone.

Sedimentology/Lithology

Profile at Grimmen

The Grimmen Formation comprises greenish to bluish claystones, siltstones, and fine-grained sandstones, with siderite and carbonate concretions, reaching up to 100 m thick in the eastern North German Basin. The type section in the Grimmen clay pit forms an 18 m composite log, transitioning from Pliensbachian sandy deposits of the Wolgast Formation to Toarcian organo-detrital clays and heteroliths of the Lehmhagen Member (Posidonia Shale Formation), followed by the Reinberg Member of the Grimmen Formation. The Reinberg Member (~10 m thick) starts with bluish clays, grading to greenish, pellet-laminated clays with exaratum and lower elegans marker beds (marly limestones with fecal pellets). Pellet laminae, up to 5 mm thick, decrease upsection, while silty/sandy intercalations and siderite concretions increase, forming a coarsening-upward trend with pyrite-filled burrows. The upper Grimmen Formation features greenish clays with thin silt/sand laminae in 8–15 m symmetric cycles, capped by the Glashütte Formation's deltaic sandstones. Heavy minerals (zircon, rutile, tourmaline) and smectite-group clay minerals indicate volcanogenic input, while kaolinite and chlorite are common. Glacially dislocated Eocene greensand and clay are locally intercalated. Organic-rich layers contain charcoal fragments, suggesting wildfires.

Toarcian material found in glacial Erratics in Ahrensburg and the Hagen Forest have been in controversy due to its dubious origin, being linked with the Rya Formation and Sorthat Formation, as well this unit. They were originally considered or local or Baltic in derivation, but that changued with the recovery of erratic concretions in the Baltic sea cliffs near Lübeck, being found as part of the Weichselian Glacial Maximum. Liassic–Cretaceous sediments in the assemblage are most probably associated with the tectonic Sorgenfrei-Tornquist Zone. The origin of this erratics from southwestern Baltic, Poland or Danish archipelago is unlikely, as those zones are dominated by Late Cretaceous–Paleocene strata, suggesting that this Toarcian assamblages should come from south/SW between STZ–TTZ and the German Baltic coast. The most clear hint link this deposits with the Grimmen Fm, as they're identical in fauna and facies composition of Grimmen and Dobbertin, also affected by subglacial erosion and thrusting, suggesting a close stratigraphic and palaeogeographical origin.

Paleoenvironment

Pliensbachian–Toarcian plankton communities in the NGB
Simplified Paleogeography of the NGB in the Toarcian, with the extent of the Grimmen Formation and adjacent units

The Grimmen Formation was deposited in a prodelta to brackish-marine environment in the eastern North German Basin, influenced by sea-level changes and the Toarcian anoxic event. The Reinberg Member reflects a transition from organo-detrital Posidonia Shale Formation to brackish-marine clays, with high total organic carbon (TOC, ~3 wt%) at the base decreasing to <1 wt% upsection, indicating a shift to dysoxic conditions. Basal bluish clays (exaratum Subzone) represent a restricted marine basin, followed by greenish clays with pellet laminae, suggesting increased fluvial input and suspension-load plumes during wetter phases. The coarsening-upward trend and onset of bioturbation (pyrite-filled burrows) reflect prodelta progradation from Fennoscandia, culminating in the Glashütte Formation's deltaic systems. High kaolinite and smectite content indicate a warm, humid climate with biochemical weathering and volcanogenic influence, akin to modern tropical settings. The absence of major ripples suggests minimal wave action, with sedimentation driven by fluvial input and storm events. The formation extends from western Poland to the Ringkøbing-Fyn-Møn-Arkona High, interfingering with the Ciechocinek Formation in the east. The Grimmen Formation's biostratigraphy is primarily based on ammonite zones, with the falciferum Zone (elegantulum, exaratum, elegans, falciferum subzones) for the lower Toarcian. The type section spans the Pliensbachian-Toarcian boundary (tenuicostatum Subzone) to the elegans Subzone, with key fossils preserved in concretions and clays.

Reinberg Member biota

Ammonites define subzones, with Dactylioceras semicelatum and possible D. tenuicostatum at the boundary, followed by Lobolytoceras siemensi and Tiltoniceras antiquum in the Lehmhagen Member (semicelatum Subzone). Elegantulum concretions yield abundant Eleganticeras elegantulum and rare Hildaites murleyi, while exaratum concretions in the Reinberg Member contain Cleviceras exaratum, Phylloceras heterophyllum, and Lytoceras crenatum. Belemnites (Passaloteuthis bisulcata) are common in the semicelatum Subzone but absent from the elegantulum Subzone due to anoxia. Teuthoids and coleoids occur in elegantulum and exaratum concretions.

Holoplanktonic gastropod Coelodiscus minutus and larval 'Inoceramus' dubius are abundant, with adults in elegantulum and exaratum concretions, often in regurgitalites. Ostracods and foraminifera are absent in black shales but occur above the Dörnten Member. Calcareous nannoplankton (e.g., Rhombolithion) are diverse in the elegantulum Subzone, correlating with NJ5b–NJ6 zones. Palynofacies show a dinoflagellate cyst blackout in the elegantulum-exaratum Subzones, with amorphous organic matter (AOM) dominance and micro-charcoal. Megaspores (e.g., Paxillitriletes phyllicus) from Fennoscandia indicate a humid climate.

Reinberg Member Fossil Record

The Reinberg Member, notably at Grimmen and Dobbertin, is a Konservatlagerstätte with exceptional marine and terrestrial fossils in exaratum concretions. Ammonites (e.g., Cleviceras exaratum), Coelodiscus minutus, and 'Inoceramus' dubius are preserved in calcite, with predation traces. Decapod crustaceans (e.g., Palaeoastacus-like), coleoid gladii, and fish (leptolepids, Grimmenodon aureum) occur in calcium phosphate. Coprolites, likely from fish, sharks, and marine reptiles (e.g., ichthyosaurs, plesiosaurs), are common. Terrestrial insects, especially in exaratum concretions types 1 and 2, include well-preserved wings and rare 3D specimens, comparable to Dobbertin's fauna. These fossils, formed in anoxic, fine-grained limestone below the storm wave base, reflect a nutrient-rich environment fertilized by insect carcasses.

Biota

Color key Taxon Reclassified taxon Taxon falsely reported as present Dubious taxon or junior synonym Ichnotaxon Ootaxon MorphotaxonNotes Uncertain or tentative taxa are in small text; crossed out taxa are discredited.
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon

Foraminifera

GenusSpeciesLocationMaterialNotesImages
BerthelinellaB. paradoxaDobbertinTestsForaminiferan of the family Plectofrondiculariidae
BolivinaB. liasicaDobbertinTestsForaminiferan of the family Bolivinidae
CornuspiraC. orbiculaDobbertinTestsForaminiferan of the subfamily Cornuspirinae
DentalinaD. deslongchampsi D. primaeva D. pseudocommunis D. submucronata D. subtenuicollis D. tenuistriata D. torta D. vetustaDobbertinTestsForaminiferan of the family Nodosariidae
EoguttulinaE. bilocularis E. liassica E. cf.simplexDobbertinTestsForaminiferan of the subfamily Polymorphininae
FalsopalmulaF. insignisDobbertinTestsForaminiferan of the family Robuloididae
FrondiculariaF. bicostata F. sulcata F. terquemi F. sp.DobbertinTestsForaminiferan of the family Nodosariidae
GlomospiraG. gordialis G. sp.DobbertinTestsForaminiferan of the subfamily Usbekistaniinae
LagenaL. globosa L. oxystoma L. globosa L. tenuicostata L. sp.DobbertinTestsForaminiferan of the family Nodosariidae
LenticulinaL. acutiangulata L. müensteri L. varians L. sublaevisDobbertinTestsForaminiferan of the family Vaginulinidae
LingulinaL. teneraDobbertinTestsForaminiferan of the subfamily Lingulininae
MarginulinaM. prima M. simplex M. oolithicaDobbertinTestsForaminiferan of the subfamily Marginulininae
NodosariaN. dispar N. agglutinans N. elongata N. germanica N. hirsuta N. dispar N. mitis N. regularis N. simplexDobbertinTestsForaminiferan of the family Nodosariidae
OphtalmidiumO. orbiculare O. carinatum O. sp.DobbertinTestsForaminiferan of the family Ophthalmidiidae
RectoglandulinaR. oviformis R. sexcostataDobbertinTestsForaminiferan of the family Nodosariidae
StilostomellaS.? sp.DobbertinTestsForaminiferan of the family Stilostomellidae
TrochamminoidesT. sp.DobbertinTestsForaminiferan of the family Trochamminoidae
TurrispirillinaT. conoideaDobbertinTestsForaminiferan of the family Spirillinidae
VaginulinaV. anomalaDobbertinTestsForaminiferan of the subfamily Vaginulininae

Ichnofossils

GenusSpeciesTypeLocationOriginImages
ChondritesC. bollensisFodinichniaDobbertin Clay PitSipunculans Polychaetes Enteropneustans EchiuransIllustration of Chondrites bollensis
MegagraptonM. isp.PascichniaDobbertin Clay PitPolychaetes Sipunculans Enteropneustans Echiurans
PlanolitesP. montanus P. beverleyensisPascichniaKozłowice Clay Pit "Boroszów" BrickyardPolychaetes InsectsExample of Planolites fossil
SkolithosS. isp.DomichniaMüssentin gravel pitPolychaetes Phoronidans
TaenidiumT. serpentinumFodinichniaDobbertin Clay PitDeposit-feeding Sipuncula Polychaetes Phoronidans

Brachiopoda

GenusSpeciesStratigraphic positionMaterialNotesImages
DiscinaD. cornucopiae D. papyracea D. sp.Klein Lehmhagen pit, Grimmen DobbertinShellsA saltwater brachiopod, member of Discinidae inside Discinida.

Bivalvia

GenusSpeciesStratigraphic positionMaterialNotesImages
BositraB. buchii B. radiataKlein Lehmhagen pit, Grimmen Dobbertin AhrensburgShellsA saltwater oyster, member of Posidoniidae.Specimens on a matrix
GoniomyaG. rhombiferaKlein Lehmhagen pit, Grimmen DobbertinShellsA saltwater clam, member of Pholadomyidae.
GresslyaG. sp.AhrensburgShellsA saltwater clam, member of Ceratomyidae.
LucinaL. planaKlein Lehmhagen pit, Grimmen DobbertinShellsA saltwater clam, member of Lucinidae.Lucina
MeleagrinellaM. substriataKlein Lehmhagen pit, Grimmen Dobbertin clay pitShellsA saltwater scallop, member of Oxytomidae.Meleagrinella
NuculaN. caeciliaDobbertinShellsA saltwater clam, member of Nuculidae.Nucula
ParainoceramyaP. dubia P. amygdaloides P. sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow ConsradeShellsA saltwater/brackish clam, member of Inoceramidae.Parainoceramya
ParallelodonP. cypriniformisAhrensburgShellsA saltwater clam, member of Parallelodontidae.
StalagminaS. koeneniKlein Lehmhagen pit, Grimmen DobbertinShellsA saltwater clam, member of Inoceramidae.
SteinmanniaS. bronni S. radiataKlein Lehmhagen pit, Grimmen DobbertinShellsA saltwater oyster, member of Posidoniidae.Specimens on a matrix
TancrediaT. elegansAhrensburgShellsA saltwater clam, member of Tancrediidae.

Gastropoda

GenusSpeciesStratigraphic positionMaterialNotesImages
CerithiumC. armatum C. roeceriKlein Lehmhagen pit, GrimmenShellsA sea snail, member of Cerithiinae.Cerithium
CoelodiscusC. minutusClay pit of Dobbertin Klein Lehmhagen pit, Grimmen Müssentin gravel pit AhrensburgShellsA holoplanktonic snail, type member of the family Coelodiscidae
TurritellaT. sp.DobbertinShellsA sea snail, member of Turritellidae.

Cephalopoda

GenusSpeciesStratigraphic positionMaterialNotesImages
AcrocoelitesA. pyramidalisKlein Lehmhagen pit, Grimmen Müssentin gravel pitMultiple specimens.A Belemnoidea member of the family MegateuthididaeAcrocoelites
BeloteuthisB. schubleriKlein Lehmhagen pit, GrimmenMultiple specimensA Mesoteuthoidea, member of the family Beloteuthidae.Beloteuthis
DactyliocerasD. athleticum D. temperatum D. attenuatum D. tenuicostatum D. semicelatum D. cf. aequistriatum D. eickenbergi D. ernstiKlein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgShellsAn ammonite, type member of the subfamily Dactylioceratinae.Dactylioceras
EleganticerasE. elegantulum E. sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgShellsAn ammonite, member of the family Hildoceratidae.Eleganticeras
HildaitesH. subserpentinusAhrensburgShellsAn ammonite, member of the family Hildoceratidae.
JeletzkyteuthisJ. coriaceusKlein Lehmhagen pit, GrimmenTwo specimensA Loligosepiidae Loligosepiida (Vampyromorpha).
LobolytocerasL. siemensi L. sp.Klein Lehmhagen pit, Grimmen Dobbertin Thüringen Brandenburg Braunschweig Boulder from AhrensburgShellsAn ammonite, member of the family Lytoceratidae.Lobolytoceras
NodicoelocerasN. crassoidesAhrensburgShellsAn ammonite, type member of the subfamily Dactylioceratinae.
PeronocerasP. fibulatumKlein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgShellsAn ammonite, member of the subfamily Dactylioceratinae.
PseudoliocerasP. lythenseKlein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgShellsAn ammonite, member of the subfamily Harpoceratinae.Pseudolioceras
PhyllocerasP. heterophyllumAhrensburgShellsAn ammonite, type member of the subfamily Phylloceratinae.Dactylioceras
TeudopsisT. bollensisKlein Lehmhagen pit, GrimmenFew specimensA vampire squid, member of Teudopsidae.Teudopsis
TiltonicerasT. antiquum T. costatum T. acutum T. schroederi T. costatum T. capillatum T. sp.Klein Lehmhagen pit, Grimmen Dobbertin Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgShellsAn ammonite, member of the family Hildoceratidae.Tiltoniceras
WhitbyicerasW. pingueAhrensburgShellsAn ammonite, member of the family Phymatoceratidae.

Crustacea

Small indeterminate shrimps, sometimes found associated in great numbers, are recovered on several layers at Grimmen.

GenusSpeciesStratigraphic positionMaterialNotesImages
BairdiaB. hildaDobbertinValvesA marine/brackish ostracod of the family Bairdiidae.
BythocyprisB. elongataDobbertinValvesA marine/brackish ostracod of the family Bythocyprididae.
CytherelloideaIndeterminateDobbertinValvesA marine/brackish ostracod of the superfamily Cytherelloidea.
EuestheriaE. opalina E. sp.Klein Lehmhagen pit, Grimmen DobbertinValvesA freshwater clam shrimp (Phyllopodan) of the family Lioestheriidae. Originally identified as a bivalve of the genus Posidonia and latter a member of the genus Lioestheria.
GlypheaG. liasinaDobbertinIsolated molts Isolated chelaeA marine decapod, type member of the family Glypheidae.Glyphea
HungarellaH. adenticulata H. sp.DobbertinValvesA marine/brackish ostracod of the family Healdiidae
ParacyprisP. semidiscaDobbertinValvesA marine/brackish ostracod of the subfamily Paracypridinae.
PolycopeP. cincinnata P. sp.DobbertinValvesA marine/brackish ostracod of the family Polycopidae.
ProcytherideaP. sp.DobbertinValvesA marine/brackish ostracod of the family Protocytheridae.
ProeryonP. hartmanni cf.P. sp.Klein Lehmhagen pit, Grimmen DobbertinIsolated molts Single specimen with preserved upper thorax and chelaeA marine decapod, member of the subfamily Proeryoninae.Proeryon
TrachycythereT. tubulosaDobbertinValvesA marine/brackish ostracod of incertae sedis affinity

Arachnida

GenusSpeciesStratigraphic positionMaterialNotesImages
SeppoS. koponeniKlein Lehmhagen pit, GrimmenSingle incomplete specimenA spider, possible member of the superfamily Palpimanoidea. It is the first confirmed spider from the lower Jurassic, and a rare find, probably washed to the sea due to a hard wind related to hurricane action. Probably a ground-dwelling predator that hunted the abundant insect fauna present on the layers.

Insecta

Insects are common terrestrial animals that were probably drifted to the sea due to Moonsonal conditions present on the Ciechocinek Formation. In Klein Lehmhagen insects are found as part of calcareous nodules in the exaratum-elegantulum subzones, with specimens also found in living chambers of Eleganticeras elegantulum macrochonchs and in fish coprolites which are the most frequent fossils at all. In the elegantulum the insect fauna is dominated by beetle elytra, indicating strong fluvial input and a nearshore deltaic complex. On Dobbertin, insects are present in the exaratum nodules, where fluvial input is seen thanks to the phyllopod abundance and whole bedding planes covered by algae substituted by Ca-phosphat, being the layers where insects are most abundant.

Notoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
GeinitziaG. schlieffeni G. debilis G. minorDobbertin, Mecklenburg (Greifswald collection)SpecimensA notopteran, member of the family Geinitziidae.
NeleN. jurassicaKlein Lehmhagen pit, GrimmenSpecimensA notopteran, member of the family Bajanzhargalanidae.Grylloblatta, extant ice crawler, Nele was probably similar
ProsepididontusP. calopteryxDobbertin, Mecklenburg (Greifswald collection)SpecimensA notopteran, member of Geinitziidae.

Odonatoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
ObotritagrionO. tenuiformum O. petersiKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection)SpecimensAn odonatopteran, member of the family Protomyrmeleontidae.
ProtomyrmeleonP. brunonis P. quadriordinumKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)Specimens

Odonata

GenusSpeciesStratigraphic positionMaterialNotesImages
AnisozygopteronA. geinitzianumDobbertin, Mecklenburg (Greifswald collection)SpecimensA dragonfly, member of the family Myopophlebiidae.
AnomothemisA. brevistigmaDobbertin, Mecklenburg (Greifswald collection)Specimensincertae sedis
ArchithemisA. brodieiDobbertin, Mecklenburg (Greifswald collection)Specimens
DialothemisD. dubiaDobbertin, Mecklenburg (Greifswald collection)SpecimensA dragonfly, member of the family Selenothemistidae.
EosagrionE. risiDobbertin, Mecklenburg (Greifswald collection)SpecimensA dragonfly, member of the family Eosagrionidae.
GrimmenopteronG. elegantulumKlein Lehmhagen pit, GrimmenSpecimensA dragonfly, member of the family Liassophlebiidae.
HeterophlebiaH. buckmani H. megapolitanaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection)SpecimensA dragonfly, member of the family Heterophlebiidae.
HeterothemisH. brodiei H. germanicaDobbertin, Mecklenburg (Greifswald collection) Ahrensburg erraticsSpecimensA dragonfly, member of the family Liassogomphidae.
LiadothemisL. hydrodictyon L. major L. geinitzi L. insignisDobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Greifswald collection)SpecimensIncertae sedis. Magnasupplephlebia represents a very large dragonfly, with a wingspan of 13 cm. Other odonatan remains are unable to being referred to a concrete group due to their incomplete status.
MagnasupplephlebiaM. kallweita M. intercalariaDobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll)Specimens
ParelthothemisP. dobbertinensisDobbertin, Mecklenburg (Greifswald collection)Specimens
PetrothemisP. singularisDobbertin, Mecklenburg (Greifswald collection)Specimens
PycnothemisP. densaDobbertin, Mecklenburg (Greifswald collection)Specimens
RhabdothemisR. strigivenaDobbertin, Mecklenburg (Greifswald collection)Specimens
SelenothemisS. liadisDobbertin, Mecklenburg (NHMW collection)SpecimensA dragonfly, type member of the family Selenothemistidae.
TemnostigmaT. singulareDobbertin, Mecklenburg (Greifswald collection)SpecimensIncertae sedis.
TrigonophlebiaT. zessiniKlein Lehmhagen pit, GrimmenSpecimensA dragonfly, member of the family Heterophlebiidae.
TuranopteronT. pommeranaKlein Lehmhagen pit, GrimmenSpecimensA dragonfly, member of the family Asiopteridae. Sphenophlebia pommerana is a junior synonym.

Paraneoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
ArchipsyllaA. primitivaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA paraneopteran, member of the family Archipsyllidae.
LiassopsocusL. lanceolatusKlein Lehmhagen pit, GrimmenSpecimensA paraneopteran, member of the family Psocidiidae.

Eoblattida

GenusSpeciesStratigraphic positionMaterialNotesImages
GriphopteronG. molleKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensAn Eoblattidan, member of the family Blattogryllidae.

Thysanoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
LiadoptiliaL. miseraKlein Lehmhagen pit, GrimmenSpecimensThysanopterans, members of the family Lophioneurida. Aeroplankton is extraordinarily well preserved in Grimmen, with the most abundant representatives of the aeroplankton (around 3 mm) being Lophioneurids, specially Undacypha europaea.Extant Thysanopteran, extinct genera were probably similar
TrichorthophlebiaT. piliferaDobbertin, Mecklenburg (Greifswald collection)
UndacyphaU. europaeaDobbertin, Mecklenburg (Greifswald collection)

Plecoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
DicronemouraD. furcataKlein Lehmhagen pit, GrimmenSpecimensA stonefly, member of the family Perlariopseidae.
DobbertiniopteryxD. capniomimusDobbertin, Mecklenburg (Ansorge coll)SpecimensA winter stonefly, member of the family Capniidae.extant Capniidae, Dobbertiniopteryx was probably similar

Orthoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
LocustopsisL. gyra L. bernstorffi L. sippeli L. dubia L. bernstorffi L. nana L. reducta L. elongata L. elegans L. pulchella L. procera L. mecklenburgica L. maculosa L. sp 1 L. sp 2 L. sp 3 L. sp 4Grimmen (Krempien collection) Lehmhagen, near Grimmen (Zessin collection) Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensGrasshoppers of the family Locustopsidae.
LocustophanesL. rhipidophorusDobbertin, Mecklenburg (Greifswald collection)SpecimensA locust, type member of the family Locustidae.Acanthacris, extant Locust, Locustophanes was probably similar
PanorpidiumP. geinitzi P. magna P. minima P. media P. reticulata P. mesostenaGrimmen (Krempien collection) Lehmhagen, near Grimmen (Zessin collection) Klein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (Zessin collection) Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA grasshopper, member of the family Elcanidae.
ParelcanaP. tenuisKlein Lehmhagen pit, Grimmen Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection)Specimens
PlesioschwinziaP. thalassophila P. laceraDobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensGrasshoppers of the family Locustopsidae.
ProtochaetaP. masculina P. lanceolataKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA grasshopper, member of the family Regiatidae.
ProtogryllusP. dobbertinensis P. germanicus P. sp. P. vicinus P. liadis P. major P. stenobasis P. femina P. acutipennis P. minor P. irregularisKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (BMNH collection) Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection) Ahrensburg ErraticsSpecimensA cricket-like grasshopper, type member of the family Protogryllidae.
SchwinziaS. solaDobbertin, Mecklenburg (Zessin collection)SpecimensGrasshoppers of the family Locustopsidae.
SynelcanaS. muelleriDobbertin, Mecklenburg (Zessin collection)SpecimensA grasshopper, member of the family Elcanidae.
ZalmonitesZ. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensA grasshopper, incertae sedis.

Phasmatodea

GenusSpeciesStratigraphic positionMaterialNotesImages
DurnovariaD. cf. parallelaKlein Lehmhagen pit, GrimmenSpecimensA stick insect, member of the family Aerophasmidae.Example of extant Phasmatodean, Durnovaria was probably similar

Blattodea

GenusSpeciesStratigraphic positionMaterialNotesImages
BlattulaB. langfeldti B. dubiaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection) Ahrensburg ErraticsSpecimensA cockroach, member of the family Blattulidae.
CaloblattinaC. mathildaeDobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA cockroach, type member of the family Caloblattinidae.
DipluroblattinaD. scudderiDobbertin, Mecklenburg (Greifswald collection)SpecimensA cockroach, member of the family Blattulidae.
EublattulaE. crassivenaDobbertin, Mecklenburg (Greifswald collection)Specimens
LiadoblattinaL. blakeiDobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA carnivorous cockroach, type member of the family Raphidiomimidae.
MesoblattinaM. protypaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA cockroach, type member of the family Mesoblattinidae.

Hemiptera

GenusSpeciesStratigraphic positionMaterialNotesImages
AcromocorisA. similisDobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA water boatman, member of the family Corixidae.Example of extant Corixidae, Acromocoris was probably similar
AnosmusA. spilopterusDobbertin, Mecklenburg (Greifswald collection)SpecimensShore bugs, member of the family Archegocimicidae.
AphlebocorisA. punctata A. nanaDobbertin, Mecklenburg (Greifswald collection)SpecimensA creeping water bug, member of the family Naucoridae.Example of extant Naucoridae, extinct genera where probably similar
ApopnusA. magniclavusDobbertin, Mecklenburg (Greifswald collection)
ApsicoriaApsicoria semideletaDobbertin, Mecklenburg (Greifswald collection)SpecimensTrue bugs, member of the family Pachymeridiidae.
ArdelaA. grimmenensisKlein Lehmhagen pit, GrimmenSpecimensLeafhoppers of the family Archijassidae.
ArchegocimexA. geinitzi A. primitiva A. liadisDobbertin, Mecklenburg (Greifswald collection)SpecimensShore bugs, member of the family Archegocimicidae.Example of extant Pentacora, Archegocimicidae genera where probably similar
ArchicercopisA. falcatusDobbertin, Mecklenburg (Greifswald collection)SpecimensMoss bugs of the family Progonocimicidae.
ArchijassusArchijassus heeriKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensLeafhoppers of the family Archijassidae.Example of extant Membracidae, Archijassidae genera where probably similar
AphidulumA. ciliatumKlein Lehmhagen pit, GrimmenSpecimensA Sternorrhynchan, member of the family Protopsyllidiidae.
ArchiconiopteryxA. liasinaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA Stemorrhynchan, type member of the family Archiconiopterygidae.
CathalusCathalus alutaceusDobbertin, Mecklenburg (Greifswald collection)SpecimensTrue bugs, member of the family Pachymeridiidae.
CixiitesC. liasinusDobbertin, Mecklenburg (Greifswald collection)SpecimensPlanthoppers of the family Fulgoridiidae.
CuneocorisC. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensA true bug, member of the family Cuneocoridae.
DiatillusDiatillus debilisDobbertin, Mecklenburg (Greifswald collection)SpecimensShore bugs, member of the family Archegocimicidae.
EocercopisE. ancyloptera E. similisDobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection)SpecimensMoss bugs of the family Progonocimicidae.
FulgoridiumF. egens F. lapideum F. dubium F. latum F. venosum F. pallidum F. geinitzi F. spilographum F. oligoneurum F. regulare F. graphipterum F. inconspicuum F. quadrisignatum F. breviradiatum F. modestum F. anale F. marginepunctatum F. picturatum F. anomalum F. stigmaticum F. punctatum F. mortuum F. alatum F. nodosum F. megapolitanum F. pulchrum F. remotum F. multivenosum F. debile F. trifurcatum F. litorale F. parvispilum F. retractum F. polyneurum F. oligospilum F. nubeculum F. obtusum F. grave F. fenestratum F. breve F. dilutum F. multipunctatum F. ampliatum F. clavatum F. nebulosum F. vicinum F. curvipenne F. elegantulum F. defunctum F. plicatum F. exhumatum F. angulosum F. acutum F. rotundatum F. ancylla F. bifurcatum F. inaequale F. brachyptilum F. intercalatum F. balticumGrimmen (EMAU collection) Klein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection)SpecimensPlanthoppers of the family Fulgoridiidae.Example of extant Fulgoridae, Fulgoridiidae genera where probably similar
GrimmenaphisG. magnificaKlein Lehmhagen pit, GrimmenSpecimensAn aphid, member of the family Oviparosiphidae.Example of extant Aphidoidea, Oviparosiphidae genera where probably similar
HadrocorisH. scutellarisDobbertin, Mecklenburg (Greifswald collection)SpecimensA true bug, member of the family Hadrocoridae.
HypocimexHypocimex membranaceusDobbertin, Mecklenburg (Greifswald collection)SpecimensTrue bugs, member of the family Pachymeridiidae.
LiadopsyllaL. geinitzi L. obtusaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA jumping plant louse, member of the family Liadopsyllidae.
LiassocercopisL. schnickiKlein Lehmhagen pit, GrimmenSpecimensA cicadomorphan, member of the family Hylicellidae.
LiasocorisL. hainmuelleriSchwinzer brickyard, near GoldbergSpecimensA true bug, incertae sedis inside Hemiptera.
MargaroptilonM. germanicumDobbertin, Mecklenburg (Greifswald collection)SpecimensPlanthoppers of the family Fulgoridiidae.
MesoledraM. pachyneuraKlein Lehmhagen pit, Grimmen Dobbertin (Naturkundemuseum Coll.) Dobbertin, Mecklenburg (Greifswald collection)SpecimensLeafhoppers of the family Archijassidae.
MetafulgoridiumM. spilotum M. ampliatum M. graptumDobbertin, Mecklenburg (Greifswald collection)SpecimensPlanthoppers of the family Fulgoridiidae.
PachymeridiumP. dubiumDobbertin, Mecklenburg (Greifswald collection)SpecimensTrue bugs, member of the family Pachymeridiidae.
ProbascanionP. megacephalumDobbertin, Mecklenburg (NHMW collection)SpecimensA Coleorrhynchan, member of the family Probascanionidae.
ProcercopisP. alutacea P. coriacea P. liasina P. similis P. jurassicaDobbertin, Mecklenburg (Greifswald collection)SpecimensFroghoppers of the family Procercopidae.
ProcercopinaP. frenzeliKlein Lehmhagen pit, GrimmenSpecimens
ProgonocimexP. jurassicus P. liasinusDobbertin, Mecklenburg (Greifswald collection)SpecimensMoss bugs of the family Progonocimicidae.Example of extant Peloridiidae, Progonocimicidae genera where probably similar
ProgonocorisP. pictusDobbertin, Mecklenburg (Greifswald collection)SpecimensShore bugs, member of the family Archegocimicidae.
SisyrocorisS. rudisDobbertin, Mecklenburg (Greifswald collection)SpecimensTrue bugs, member of the family Pachymeridiidae.
ToarcopsyllidiumT. ernstiKlein Lehmhagen pit, GrimmenIsolated WingsA Sternorrhynchan, member of the family Protopsyllidiidae.

Hymenoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
BrigittepterusB. brauckmanniDobbertin, Mecklenburg (Zessin collection)Isolated WingsWasps, member of the family Ephialtitidae.
GrimmaratavitesG. mirabilisKlein Lehmhagen pit, GrimmenSpecimensA Wood Wasp, member of the family Karatavitidae.
LiadobraconaL. raduhnaDobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll)Isolated WingsWasps, member of the family Ephialtitidae.
XyelulaX. benderiKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll)SpecimensA pseudo-Wasp, member of the family Sepulcidae.

Megaloptera

GenusSpeciesStratigraphic positionMaterialNotesImages
DobbertiniaD. reticulataDobbertin, Mecklenburg (Greifswald collection)SpecimensAn alderfly, member of the family Sialidae.Example of extant Sialidae, Dobbertinia was probably similar

Neuroptera

GenusSpeciesStratigraphic positionMaterialNotesImages
ActinophlebiaA. megapolitana A. aenea A. parallelaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensLacewings of the family Prohemerobiidae
ApeirophlebiaA. grandisDobbertin, Mecklenburg (Greifswald collection)SpecimensA silky lacewing, member of the family Psychopsidae.Example of extant Psychopsidae, Apeirophlebia was probably similar
EpigambriaE. longipennisDobbertin, Mecklenburg (Greifswald collection)SpecimensA lacewing, member of the family Epigambriidae.Example of extant Ithonidae, Epigambriidae members where probably similar
LiassochrysaL. stigmaticaDobbertin, Mecklenburg (Ansorge coll)SpecimensA Neuropteran of the family Mantispidae.Example of extant Mantispidae, Liassochrysa was probably similar
LoxophlepsL. costalisDobbertin, Mecklenburg (Greifswald collection)SpecimensA lacewing, incertae sedis inside Neuroptera.
MelaneimonM. dubiumDobbertin, Mecklenburg (Greifswald collection)Specimens
MelamnousM. indistinctusDobbertin, Mecklenburg (Greifswald collection)Specimens
MesoleonM. dobbertinianusDobbertin, Mecklenburg (Greifswald collection)Specimens
MesosmylinaM. falciferaKlein Lehmhagen pit, GrimmenSpecimensA lance lacewing, member of the family Osmylidae.
PolyosmylusP. excelsusKlein Lehmhagen pit, GrimmenSpecimensA lacewing, member of the family Epigambriidae.
ProhemerobiusP. prodromus P. ovatus P. anomalus P. costalis P. persimilis P. geinitzi P. liasinus P. major P. parvulus P. geinitzianus P. dilaroides P. chryseus P. latusKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensLacewings of the family Prohemerobiidae.
SolenoptilonS. kochiDobbertin, Mecklenburg (Greifswald collection)SpecimensA lacewing, type member of the family Solenoptilidae.
ToarciconiopteryxT. dipterosimilisKlein Lehmhagen pit, GrimmenSpecimensA dustywing, member of the subfamily Toarciconiopteryginae.Example of extant Coniopterygidae, Toarciconiopteryx was probably similar

Coleoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
AllognosisA. nitensDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
AnancaeonA. microcephalumDobbertin, Mecklenburg (NHMW collection)Specimens
AnomerusA. punctiferDobbertin, Mecklenburg (Greifswald collection)Specimens
AnypostatusA. taurusDobbertin, Mecklenburg (Greifswald collection)Specimens
ApioderesA. punctatusDobbertin, Mecklenburg (Greifswald collection)Specimens
ApsychusA. alutaceusDobbertin, Mecklenburg (Greifswald collection)Specimens
BareusB. strigipennisDobbertin, Mecklenburg (Greifswald collection)Specimens
BathygerusB. bellus B. divergensDobbertin, Mecklenburg (Greifswald collection)Specimens
CarabitesC. dubius C. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensA ground beetle, member of the family Carabidae.Example of extant Carabidae, Carabites was probably similar
ClinomerusC. laticollisDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
CoptogyrinusC. scutellatusDobbertin, Mecklenburg (Greifswald collection)SpecimensA whirligig beetle, member of the family Gyrinidae.Example of extant Gyrinidae, Coptogyrinus was probably similar
EcthlimmaE. forficuloidesDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
EnammaE. striatumDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
EurynuchaE. pseudobuprestisDobbertin, Mecklenburg (Greifswald collection)SpecimensA jewel beetle, member of the family Buprestidae.Example of extant Buprestidae, Eurynucha was probably similar
GyrinulopsisG. nanuDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
HydrobiitesH. dobbertinensis H. punctatostriatusDobbertin, Mecklenburg (NHMW collection)SpecimensA beetle, member of the family Permosynidae.
KeleusticusK. zirkeliDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
LiassoditesL. obstiKlein Lehmhagen pit, GrimmenSpecimensA False Ground Beetle of the family Trachypachidae.Extant member of Trachypachidae, Liassodites was probably similar
MasselytronM. quinquestriatumDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
NannoodesN. pseudocistelaDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
NebrioidesN. dobbertinensisDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
OnkedodimusO. discicollisDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
OxytoropteraO. mediocrisDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
ParacurculiumP. punctatumDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
ParnidiumP. frechi P. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PeribolopteraP. rotundaDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PlastobuprestitesP. elegansDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PlastonebriaP. scudderiDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PolypamonP. byrrhoidesDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PseudocyphonP. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
PseudoprionitesP. liasinusDobbertin, Mecklenburg (Greifswald collection)SpecimensBeetles, incertae sedis inside Coleoptera.
ThoracotesT. dubiusDobbertin, Mecklenburg (Greifswald collection)SpecimensA bark gnawing beetle, member of the family Trogossitidae.Example of extant Trogossitidae, Thoracotes was probably similar

Trichoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
ArchiptiliaA. ovataDobbertin, Mecklenburg (Greifswald collection)SpecimensCaddisflies, member of the family Necrotauliidae.
EpididontusE. geinitzianusDobbertin, Mecklenburg (Greifswald collection)Specimens
LiadotauliusL. maiorKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA finger-net caddisfly, member of the family Philopotamidae.Example of extant Philopotamidae, Liadotaulius was probably similar
MesotrichopteridiumM. intermedium M. pusillumKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensCaddisflies, member of the family Necrotauliidae.
MetarchitauliusM. longusDobbertin, Mecklenburg (Greifswald collection)Specimens
NannotrichopteronN. gracileDobbertin, Mecklenburg (Greifswald collection)Specimens
NecrotauliusN. parvulusKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)Specimens
PalaeotauliusP. vicinusDobbertin, Mecklenburg (Greifswald collection)Specimens
ParatauliusP. jurassicusDobbertin, Mecklenburg (Greifswald collection)Specimens
ParatrichopteridiumP. efossum P. costaleDobbertin, Mecklenburg (Greifswald collection)Specimens
PararchitauliusP. ovalisDobbertin, Mecklenburg (Greifswald collection)Specimens
PseudorthophlebiaP. platypteraDobbertin, Mecklenburg (Greifswald collection)Specimens

Lepidoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
EolepidopterigidaeEolepidopterigidae indet. A Eolepidopterigidae indet. B Eolepidopterigidae indet. CKlein Lehmhagen pit, GrimmenIsolated wings Isolated wing scalesA moth, member of the family Eolepidopterigidae.

Mecoptera

GenusSpeciesStratigraphic positionMaterialNotesImages
MesobittacusM. minutusKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensHangingflies, member of the family Bittacidae.
MesochoristaM. sinuataDobbertin, Mecklenburg (Greifswald collection)SpecimensA Scorpionfly, member of the family Permochoristidae.
MesopanorpaM. maculataDobbertin, Mecklenburg (Greifswald collection)SpecimensHangingflies, member of the family Orthophlebiidae.
NeorthophlebiaN. maculipennis N. megapolitana N. pallida N. debilis N. simillima N. medialis N. stigmatica N. acutipennisKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensHangingflies, member of the family Bittacidae.Example of extant Bittacidae, extinct members where probably similar
OrthophlebiaO. germanica O. elongata O. fuscipennis O. limnophila O. cf. limnophila O. radialis O. liadis O. maculata O. vicina O. brunsvicensisKlein Lehmhagen pit, Grimmen Dobbertin (Breslau Museum Coll.) Dobbertin, Mecklenburg (Greifswald collection)SpecimensHangingflies, member of the family Orthophlebiidae.
ParabittacusP. analis P. lingulaDobbertin, Mecklenburg (Greifswald collection)SpecimensHangingflies, member of the family Bittacidae.
PseudopolycentropusP. triangularis P. perlaeformis P. sp.Klein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA scorpionfly, type member of the family Pseudopolycentropodidae.

Diptera

GenusSpeciesStratigraphic positionMaterialNotesImages
AenneA. liasinaSchönenwald railway cutting, Grimmen Klein Lehmhagen pit, GrimmenSpecimensThe oldest known non biting midgess, members of the ChironomidaeExample of extant Chironomidae, extinct members where probably similar
AntefungivoraA. germanicaKlein Lehmhagen pit, GrimmenSpecimensFlies, member of the family Antefungivoridae.
ArchibioA. mycetophilinusKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)Specimens
ArchipleciomimaA. germanica A. sp.Klein Lehmhagen pit, GrimmenSpecimensA fly, incertae sedis inside Diptera
ArchirhyphusA. geinitziKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensFlies, member of the family Protorhyphidae.
ArchitipulaA. stigmatica A. latipennis A. elegans A. seebachi A. nana A. pusilla A. pulla A. debilis A. simplex A. intermedia A. obliqua A. areolata A. maior A. conspicua A. clara A. dubiaDobbertin, Mecklenburg (Greifswald collection)SpecimensA crane fly of the family Limoniidae.Example of extant Limoniidae, extinct members where probably similar
CrenoptychopteraC. dobbertinensisDobbertin, Mecklenburg (Ansorge coll)SpecimensA Phantom crane fly, member of the family Ptychopteridae.Example of extant Ptychopteridae, extinct members where probably similar
EoditomyiaE. primitivaKlein Lehmhagen pit, GrimmenSpecimensA fly, type member of the family Eoditomyidae.
EolimnobiaE. geinitziDobbertin, Mecklenburg (Greifswald collection)SpecimensA phantom crane fly, member of the family Ptychopteridae.
EoptychopteraE. simplex E. eximiaDobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Greifswald collection)Specimens
EotipulaE. defuncta E. mortua E. coarctataDobbertin, Mecklenburg (Greifswald collection)SpecimensA crane fly of the family Limoniidae.
GrimmeniaG. prima G. secundaLehmhagen, near Grimmen (Zessin collection)SpecimensA crane fly of the family Limoniidae.
GrimmyiaG. balticaKlein Lehmhagen pit, GrimmenSpecimensA snipe fly, member of the family Rhagionidae.Example of extant Rhagionidae, extinct members where probably similar
HeterorhyphusH. triangularisKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA fly, type member of the family Heterorhyphidae.
LehmhageniaL. ansorgeiKlein Lehmhagen pit, GrimmenSpecimensA fly, member of the family Protobrachyceridae.
LiassobrachyceronL. kotejaiDobbertin, Mecklenburg (Ansorge coll)SpecimensA snipe fly, member of the family Rhagionidae.
LiassopsychodinaL. pommeranaKlein Lehmhagen pit, GrimmenSpecimensA Moth fly, member of the family Psychodidae.Example of extant Psychodidae, extinct members where probably similar
MailotrichoceraM. mikereichi M. variabilis M. zessiniKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Ansorge coll)SpecimensA Winter Crane Fly, member of the family Trichoceridae.Example of extant Trichoceridae, extinct members where probably similar
MesorhyphusM. nanusDobbertin, Mecklenburg (Greifswald collection)SpecimensA wood gnat, member of the family Anisopodidae.Example of extant Anisopodidae, extinct members where probably similar
MesotipulaM. siggiae M. vicina M. cf. vicina M. mirabilis M. curvata M. geinitzi M. minuta M. lapidaria M. parvaLehmhagen, near Grimmen (Zessin collection) Dobbertin, Mecklenburg (NHMW collection) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA Crane Fly of the family Limoniidae.
MetatrichopteridiumM. confusumDobbertin, Mecklenburg (Greifswald collection)SpecimensA fly, member of the family Hennigmatidae.
NannotanyderusN. grimmenensis N. krzeminskiiKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll)SpecimensA primitive Crane Fly, member of the family Tanyderidae.
OryctochlusO. toarciensisKlein Lehmhagen pit, GrimmenSpecimensThe oldest known non biting midges, members of the Chironomidae.
RhaetomyiaR. herrigiDobbertin, Mecklenburg (Greifswald collection)SpecimensA Phantom midge of the family Chaoboridae.Example of extant Chaoboridae, extinct members where probably similar
PalaeobrachyceronP. willmanniKlein Lehmhagen pit, GrimmenSpecimensA Snipe fly, member of the family Rhagionidae.
PhryganidiumP. minimumDobbertin, Mecklenburg (Greifswald collection)SpecimensA Crane Fly of the family Limoniidae.
PleciofungivoraP. liassicaKlein Lehmhagen pit, GrimmenSpecimensA fly, member of the family Pleciofungivoridae.
PodonomiusP. tumidusKlein Lehmhagen pit, GrimmenSpecimensThe oldest known non biting midges, members of the Chironomidae.
PraemacrochileP. decipiens P. dobbertinensisDobbertin, Mecklenburg (Zessin collection) Dobbertin, Mecklenburg (Ansorge coll)SpecimensA primitive Crane Fly, member of the family Tanyderidae.
ProtobrachyceronP. liasinumDobbertin, Mecklenburg (Ansorge coll) Dobbertin, Mecklenburg (Greifswald collection)SpecimensA fly, member of the family Protobrachyceridae.
ProtopleciaP. klafacki P. liasinaKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensA fly, type member of the family Protopleciidae.
ProtorhyphusP. simplexKlein Lehmhagen pit, Grimmen Dobbertin, Mecklenburg (Greifswald collection)SpecimensFlies, member of the family Protorhyphidae.
TanypsychaT. connexaKlein Lehmhagen pit, GrimmenSpecimensA Moth fly, member of the family Psychodidae.

Echinodermata

In Dobbertin, the echinoderm remains are rare in contrast to foraminifera, phyllopods and ostracods, yet in some places they attain a percentage of the total fauna between 0.7 and 26.5%. In the upper layers they're totally absent, as well on the erratics and in the whole Grimmen sequence.

GenusSpeciesStratigraphic positionMaterialNotesImages
CrinoideaIndeterminateDobbertinColumnar ossiclesIncertae sedis
HolothuriidaIndeterminateDobbertinHolothurian wheelsIncertae sedis
SinosuraS. brodiei S. spp.DobbertinIsolated and semiarticulated specimensA brittle star, member of Ophioleucidae inside Ophioleucida. Identified as Ophiura longivertebralis.

Vertebrates

Fishes

GenusSpeciesStratigraphic positionMaterialNotesImages
DapediumD. pholidotumDobbertinIncomplete skullMember of the Dapediidae.Dapedium
GrimmenichthysG. ansorgei Cf. G. ansorgeiKlein Lehmhagen pit, GrimmenArticulated, but incompletely preserved specimensMember of the group Pholidophoriformes. This genus was a small 'pholidophoriform' teleost.
GrimmenodonG. aureumKlein Lehmhagen pit, GrimmenAlmost complete left prearticular with dentitionMember of the order Pycnodontiformes. Is of comparatively small size, suggesting a hypothetical small standard length of the fish of about 7–10 cm at the time of death.
GyrosteusG. mirabilisAhrensburg Erratic AssemblageGPIH 4864, HyomandibulaMember of the Chondrosteidae inside Acipenseriformes.
HybodusH. hauffianus H. spp.Klein Lehmhagen pit, Grimmen Ahrensburg erratic assemblageFragmentary Fin spine Supposed coprolithes TeethMember of the Hybodontiformes.Hybodus hauffianus
LepidotesL. buelowianus L. elvensis L. spp.Klein Lehmhagen pit, Grimmen Dobbertin Ahrensburg Erratic AssemblageVarious specimensMember of the family Lepidotidae inside Lepisosteiformes.L. buelowianus
LepisosteidaeIndeterminateGrimmenGG 439/4-7 scalesThe oldest confirmed true Gar, member of the Lepisosteidae.Lepisosteus extant example of Gar
LeptolepidaeIndeterminateFormer clay pit of DobbertinIsolated head with missing postcranial bones Assigned TeethMember of the family Leptolepidae.
LeptolepisL. bronni L. coryphaenoides L. normandica L. jaegeri L. spp.Klein Lehmhagen pit, Grimmen Dobbertin clay pit Karziger Ufer Müssentin Broock Hohenbarnekow Grenzin Groß Roge Elmenhorst Klütz Höved Pinnow Consrade AhrensburgSlightly disarticulated, incomplete head; Subadult, almost complete; fragmentary head; isolated head with pectoral girdle; incomplete head Isolated remains in form of "Grätensandsteine"Type member of the family Leptolepidae. The most abundant vertebrate recovered on the formation, including 3D preserved specimens, as well, is the main component of the Fishbone sandstones from the upper layers.Leptolepis
LycopteroidarumL. sp. A L. sp. BAhrensburg Erratic AssemblageOtolithsOtoliths of the family Lycopteroidea inside Lycopteriformes.
MengiusM. acutidensKlein Lehmhagen pit, Grimmen Schwinzer Ziegelei ca 3 km E of DobbertinHolotype: Articulated, incomplete fish Referred articulated, semiarticulated and fragmentary specimensMember of the family Lepidotidae inside Lepisosteiformes. Stomach content is also preserved on a specimen from Dobbertin, and is composed by arthropod cuticles.
PachycormiformesIndeterminateKlein Lehmhagen pit, GrimmenUnknown, only cited.Member of the family Pachycormiformes.
PholidophorusP. sp.Ahrensburg Erratic AssemblageAssigned Fragmentary remainsMember of the family Pholidophoriformes.
PholidophoriformesIndeterminateKlein Lehmhagen pit, GrimmenIsolated skull roof with articulated ethmoid region, parasphenoid, and partial braincaseMember of the family Pholidophoriformes. Resembles Pholidophoroides crenulata and P. limbata.
ProleptolepisP. sp.Klein Lehmhagen pit, GrimmenMV 202612, fragmentary head . Assigned TeethMember of the family Leptolepidae. Specimens attributed to Proleptolepis have previously been reported only from the Sinemurian of western Europe.
SaurorhynchusS. hauffi S. sp.Klein Lehmhagen pit, Grimmen DobbertinDiganostic Incomplete skull Incomplete skullThe youngest representative of the family Saurichthyidae. Distinctive for its large jaws, similar to modern Belonidae.Saurorhynchus
TetragonolepisT. semicinctaKlein Lehmhagen pit, GrimmenIncomplete specimen Isolated JawMember of the family Dapediidae.Tetragonolepis

Amniotes

GenusSpeciesStratigraphic positionMaterialNotesImages
IchthyosauriaIndeterminateKlein Lehmhagen pit, Grimmen Ahrensburg Erratic AssemblagePresacral centrum 2 ?sacral centrum Isolated vertebraeIncertae sedis
MacrospondylusM. bollensisKlein Lehmhagen pit, GrimmenIncomplete skull and associated osteodermA marine crocodrylomorph, member of the family Machimosauridae. GG 422/6 can be clearly referred to an immature individual.Macrospondylus
MegalosauridaeIndeterminateForst Hagen gravel pitDorsal vertebraeA theropod, member of the family Megalosauridae. Was referred to Megalosaurus. The vertebrae centrum measures 80 mm, implying a medium-sized theropod (~5 m long).Marshosaurus, example of basal Orionides
MesoeucrocodyliaIndeterminateKlein Lehmhagen pit, GrimmenCervical vertebraA crocodrilian, member of Mesoeucrocodylia. Is considered to come from an osteologically immature individual.
MeyerasaurusM. sp.Ahrensburg Erratic AssemblageIncomplete coracoid Associated ribA pliosauroid, member of the family Rhomaleosauridae.Meyerasaurus
MystriosaurusM. sp.Klein Lehmhagen pit, GrimmenPartial rostrum with teethA marine crocodrylomorph, member of the family Teleosauridae. Is indicative of a juvenile individual, with expected around 200–250 mm long skull and the entire animal about 1.50 m.
NeoichthyosauriaIndeterminateClay Pit of DobbertinFour articulated tail vertebrae.An indeterminate ichthyosaur, has been assigned to the species Stenopterygius longifrons.
Ophthalmosauridae?IndeterminateKlein Lehmhagen pit, GrimmenPartial skull and associated postcranial elements preserved in a concretionIt has an expanded basipterygoid process on the basisphenoid, only currently known in members of the Ophthalmosauridae
PlesiosauriaIndeterminateKlein Lehmhagen pit, Grimmen Dobbertin, falciferum zoneThree caudal vertebrae and other postcranial elements Gastralia Phalanx Cervical rib and other postcranial elementsIncertae sedis
PlesiosauroideaIndeterminateKlein Lehmhagen pit, GrimmenCervical centrum Caudal centrumIncertae sedis
Pterosauria?IndeterminateKlein Lehmhagen pit, GrimmenGG 510, fragmentary tibiofibular(?)A possible pterosaur bone, with features similar to the tibiofibular of coeval taxon Dorygnathus banthensis. Alternatively it can be another type of bone and belong to stem-turtles, plesiosaurs, dinosaurs or even anurans.Dorygnathus banthensis, know from inner localities, has shared features that may suggest GG 510 is from a Pterosaur
SeeleyosaurusS.? sp.Ahrensburg Erratic AssemblageThree articulated dorsal vertebraeA plesiosaur, member of the family Plesiosauridae.Seeleysaurus
StenopterygiusS. cf. quadriscissusClay pit of DobbertinPosterior left half of the cranium.An ichthyosaur of the family Stenopterygiidae.Restoration
TeleosauridaeIndeterminateAhrensburg Erratic AssemblageOsteoderms Isolated non described remainsIncertae sedis
TemnodontosaurusT. cf. platyodonAhrensburg Erratic AssemblageSkull Coracoid Associated rib fragmentAn icthyosaur, type member of Temnodontosauridae. Assigned to Ichthyosaurus sp., but also suggested affinities to "Leptopterygius" (= Temnodontosaurus) platyodon.Restoration

Flora

GenusSpeciesStratigraphic positionMaterialNotesImages
BotryococcusB. sp.Grimmen clay pitFreshwater algaeIndicates freshwater influx in palynofacies assemblage IV, associated with coastal or deltaic settings.
EquisetitesE. sp. E. cf. Münsteri E. columnareGlacial Erratic Near Grimmen Dobbertin Ahrensburg area Erratic deposit near LüttowIsolated stems, rhizomesEquisetalean stems of the family Equisetidae. Dominant in mono-to oligotypic stands near water bodies, indicative of high terrestrial input in palynofacies assemblages I and IV.
ErlansonisporitesE. sparassis E. sp.Grimmen clay pitMegasporesAffinities with the Selaginellaceae. Transported from northern sources (e.g., Fennoscandia), indicative of humid climates.
HughesisporitesH. pustulatusGrimmen clay pitMegasporesAffinities with the Selaginellaceae. Likely transported via suspension load plumes, reflecting freshwater influx.
MinerisporitesM. institus M. volucrisGrimmen clay pitMegasporesAffinities with the Selaginellaceae. Associated with warm, humid conditions and high atmospheric CO2.
PagiophyllumP. sp. P. cf. KurriGrimmen clay pitLeaves, driftwoodConifer macrofossils of the family Cheirolepidiaceae or Araucariaceae. High phytoclasts and cuticles in palynofacies assemblages I, II, and IV support conifer presence.
PaxillitriletesP. phyllicusGrimmen clay pit Reinberg 1EMegasporesAffinities with the Isoetales. Transported from Fennoscandia, indicates humid climate, biostratigraphic marker (Paxillitriletes phyllicus Zone, also seen in the Ciechocinek Formation)Extant Isoetes, typical example of Isoetales. Paxillitriletes probably come from a similar or a related Plant
StriatriletesS. excavatusGrimmen clay pitMegasporesAffinities with the Selaginellaceae. Reflects freshwater influx from northern sources.
TrileitesT. spp.Grimmen clay pitMegasporesAffinities with lycophytes or ferns. Indicates diverse megaspore flora transported from northern sources.

See also