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The Sorthat Formation is a geologic formation on Bornholm, Denmark, and the Rønne Graben, Baltic Sea, from the Latest Pliensbachian to Late Toarcian. It holds plant fossils and invertebrate traces, overlain by fluvial and lacustrine deposits of the Aalenian-BathonianBagå Formation. Initially classified as part of the Bagå Formation until 2003, it spans the Latest Pliensbachian to Early Aalenian. It reflects a deltaic to marine setting with eastern river systems forming in the Toarcian. Early Pliensbachian volcanism from southern Sweden extended across the North Sea. The Central Skåne Volcanic Province and Egersund Basin contributed volcanic material, affecting tectonics. Early Jurassic porphyriticnephelinite lavas in the Egersund Basin, akin to those in the formation's clay pits, suggest fluvial sediment transport to the Grimmen Formation and Ciechocinek Formation. The Grimmen Formation is its sister unit.
Description
Sorthat Formation layers, mainly mudstones, claystones, and sandstone bands
Bornholm's Lower-Middle Jurassic includes the Rønne (Hettangian–Sinemurian), Hasle (Early–Late Pliensbachian), Sorthat, and Bagå Formations. Coal-bearing clays and sands overlie the Hasle Formation, divided between the Sorthat and Bagå. The Sorthat Formation aligns with the Röddinge Formation, sharing a fluvial system, and correlates with the Ciechocinek Formation, Fjerritslev Formation, and Rya Formation. Originally termed Levka, Sorthat, and Bagå beds, its key exposure is the Korsodde section. Faulting and scarce marine fossils hinder stratigraphic clarity. Palynological data from the Levka-1 core and Korsodde section date it to Late Pliensbachian–Toarcian, possibly Early Aalenian. Megaspores indicate Toarcian–Aalenian strata. It features bioturbated sands, heteroliths, clays, coal veins, and dinoflagellates, suggesting brackish to marine settings, capped by Bagå deposits.
The Sorthat Formation's lithology varies. The Levka-1 well reveals fining-upward units, 3–14 m thick, with coarse sand, muddy, coal- and mica-rich sands, clays, and coal seams. Parallel lamination dominates, with minor cross-bedding. Abundant plant fragments and quartz, but no marine palynomorphs, indicate a coastal delta plain. This mirrors the Ciechocinek Formation's Toarcian–Bajocian deltaic shoreline. The North German Basin shows four sea-level fluctuations forming delta generations, with Toarcian regressive deltas depositing 40 m in Prignitz and Brandenburg. Palynomorphs tie to the Sorthat Formation.
The upper formation (~40 m) has bioturbated sands, heteroliths, syneresis cracks, pyrite nodules, and ichnofossils like Planolites and Teichichnus, reflecting nearshore lagoons and channels. The 93 m Korsodde section, with organic-rich sands, suggests fluvial channels tied to coastal lakes, with plant remains and ichnofossils like Diplocraterion. The top features fine, yellowish-brown sands and sandstones with bioturbated, wave-rippled beds.
At Korsodde, the environment includes the following:
Stratigraphy of the Korsodde section
Unit
Lithology
Thickness (metres)
Type of environment
Fossil flora
Fossil fauna
Unit A
Yellow, weakly cemented muscovitequartz sandstone, medium- to fine-grained in the lower part, fine-grained in the upper part.
0.45–2.3 m
Estuarine channel fill (upper or marginal, less energetic part)
Intercalations of muscovite quartz sandstones and dark mudstone drapes, with abundant heteroliths.
2.3–3.41 m
Upper tidal flat deposits surrounding an estuary
None recovered
Planolites isp. Rosselia isp. Palaeophycus isp. B. tortuosusD. parallelum
Unit C
Two main layers: a series of 20 cm dark mudstone with horizontal lamination and silt intercalations and a series of dark heteroliths with intercalated mudstones and ripple limestones.
3.41–3.7 m
Restricted bay passing into upper tidal flat deposits
None recovered
D. parallelum
Unit D
Yellow ripple cross sandstone with abundant muscovite, alternating with continuous and discontinuous dark mudstone with abundant organic material. There are pyrite concretions in the lower part.
3.7–4.7 m
Lower tidal flat within an estuary
Roots
Planolites isp. Palaeophycus isp. B. tortuosusD. parallelum
Unit E
Mostly fine-grained sediments with abundant organic matter.
A prominent erosional surface at the start, composed of yellow medium- to fine-grained cross-laminated sandstones with muscovite.
9.9–11.35 m
Estuarine bar
None reported
None reported
Unit H
Pale, fine-grained ripple and herringbone sandstones and mudstones, with intercalations of sandy mudstones and mudstone drapes with intense ferruginization, and some layers of mudstone–sandstone heteroliths
11.35–14.2 m
Marginal part of an estuary channel fill
None reported
Rosselia isp. Palaeophycus isp. Cylindrichnus isp. Skolithos isp. D. parallelum
The Sorthat Formation hosts a rich Pliensbachian–Toarcian flora, one of Europe's most complete for this period, with significant Jurassic palynological deposits. The flora, including gymnosperms, ferns (e.g., Dicksonia, Coniopteris, Osmundaceae), and thin-cutinised leaves like Podozamites and Equisetales, suggests a warm, humid climate conducive to diverse vegetation.
Environment
Late Pliensbachian–Early Toarcian Fennoscandinavia, with flora based on the Sorthat Formation and fauna from the Lehmhagen Member, Grimmen Formation and Drzewica Formation.Paleogeography of the North German basin in the Toarcian, showing the Grimmen Formation and Sorthat Formation extent.
The Sorthat Formation, deposited in the Rønne Graben and on Bornholm's coasts, reflects a deltaic to marine environment, with the Grimmen Formation as its sister unit. A modern analog is New Zealand's Northland humid coastal forests, where peat-forming woodlands thrive. The Stina-1 well shows sand, clay, and coal, indicating an emerged graben. High kaolinite and reworked Carboniferous palynomorphs suggest erosion of a Carboniferous regolith. A Late Pliensbachian regression allowed coal deposition, halted by an Early Toarcian transgression. The Levka-1 well and Korsodde section show lagoons, channels, and floodplains, with rapid subsidence in the Rønne Graben during the Toarcian. Shoreface deposits with bioturbation mark a deepening trend, correlating with the Fjerritslev Formation. Depositional settings include the Levka Beds, interpreted as fluvial channels, floodplains, and peat swamps. Marine palynomorphs (e.g., Nannoceratopsis) indicate lagoonal settings. The Sorthat Beds represent delta plain deposits with pyrite nodules and Arenicolites traces. The Korsodde Section was a fluvial channel sands with coal and rootlets, transitioning to lagoons and palaeosols, with dinoflagellates (e.g., Mendicodinium). Outside emerged Bornholm, offshore Rønne and Kolobrzeg grabens include mostly fluvial-coastal layers, fed by regional currents.
The Sorthat Formation, with the Grimmen Formation as its sister unit, records significant vegetation changes during the Early Toarcian, linked to carbon cycle perturbations and the Toarcian oceanic anoxic event, driven by large-scale volcanism. A modern analog is New Zealand's Northland humid coastal forests, supporting peat-forming woodlands.
The Sorthat Fm was a cold, humid y forested setting, whose best modern analogs can be found in New ZealandStratigraphic map of Bornholm
In the Late Pliensbachian, pollen assemblages, dominated by Cupressaceae (e.g., Perinopollenites) and cycads (Cycadopites), indicate a warm, humid Mediterranean climate. During the Toarcian event, spore-rich layers reflect a shift to ferns and lycophytes, suggesting increased humidity. Post-event, Hirmeriellaceae pollen (e.g., Corollina, Spheripollenites) dominates, indicating a drier, warmer climate. Woody vegetation shows changes in carbon isotopes, with pollen from Sciadopityaceae, Miroviaceae (Cerebropollenites), cycads (Chasmatosporites), and Corollina. Macroflora, mainly from the Hasle clay pit and Korsodde section, includes 68 species, 50% ferns. The Late Pliensbachian flora, rich in ferns and sphenophytes, grew in marshy floodplains along a meandering river, with Bennettites as shrubs and conifers (e.g., Pagiophyllum) and ginkgoaleans as trees, reflecting a warm, seasonal climate. In the Toarcian, Hirmeriellaceae conifers (95% of pollen), seed ferns, Bennettites, and Czekanowskiales dominate, with Pagiophyllum and Corollina torosus indicating high temperatures, aridity, and seasonal wildfires. Wood fragments, both macroscopic and microscopic (0.25–1 mm), are common in Korsodde's nearshore deposits.
Coal seams, mainly from Levka-1 and Korsodde, formed in anoxic, nutrient-rich swamps. Peat accumulated rapidly (~1 mm/yr), akin to Central Kalimantan, in a warm, humid climate. High huminite content (e.g., Eu-ulminite, densinite) indicates anoxic conditions. Wildfires, evidenced by charcoal, increased during the Toarcian oceanic anoxic event, reflecting drier conditions. Coals can be found at Levka-1 (112 m of coal, sand, and clay with abundant coalified wood), representing fluvial channels and floodplains with lagoons. At Korsodde six coal seams from a coastal lagoon setting can be found, with huminite-rich coal and dinoflagellates like Mendicodinium reticulatum.
Fungal spores of uncertain classification. The three uppermost samples of the Korssode section are poor in diversity, but fungal spores are common in at least one sample; these have not been recorded from the samples below.
ExtantGeastrum campestre specimen, found linked with plant matter. Spores recovered on the Sorthat Formation may be derived from similar fungi.
Phytoplankton
In the Lower Jurassic of Bornholm there were several successions of nearshore peat formations with dinoflagellates. Coal-bearing strata were deposited in an overall coastal plain environment during the Hettangian–Sinemurian, and then during the Early Pliensbachian deposition was interrupted until the late Pliensbachian–Lowermost Toarcian due to a sea regression.
Type genus of the Botryococcaceae in the Trebouxiales. A colonial green microalga of freshwater and brackish ponds and lakes around the world, where it often can be found in large floating masses. Sorthat Formation Botryococcus lived in an environment interpreted as a coastal lake, permanently vegetated and shallow, that was occasionally flooded by the sea.
Extant specimen
Chomotriletes
C. minor
Korsodde section Levka-1 borehole Sorthat beds
Miospores
Affinities with the family Zygnemataceae. A genus derived from freshwater filamentous or unicellular, uniseriate (unbranched) green algae.
N. senexN. gracilisN. ridinguiN. triangulataN. tricerasN. dictyanbonisN. sp.
Korsodde section Levka-1 borehole Sorthat beds
Cysts
A dinoflagellate, member of the family Nannoceratopsiaceae. It is characteristic of marine deposits. The presence of N. gracilis, N. senex and N. triceras, and common occurrence of Botryococcus is interpreted as indicating a lagoon succession overlying a transgressive surface and signals a rise in relative sea level.
Ovoidites
O. sp. A O. sp. B O. sp. C O. spp.
Korsodde section Levka-1 borehole
Miospores
Affinities with the family Zygnemataceae. A genus derived from freshwater filamentous or unicellular, uniseriate (unbranched) green algae.
Extant Spirogyra; Ovoidites may be derived from a similar genus
Pterospermella
P. spp.
Korsodde section
Miospores
Affinities with the family Pterospermataceae.
Rotundus
R. granulatus
Sorthat beds
Miospores
An algal palynomorph unique to the setting and probably related to freshwater red algae; similar to extant Batrachospermales.
A foraminifer, type genus of the Spirillinidae in the Spirillinida.
Striatella
S. jurassicaS. parvaS. seebergensisS. scanica
Korsodde section Levka-1 borehole Sorthat beds
Miospores
Brown algae, type genus of the family Striatellaceae in the Striatellales. These brown algae diatoms are associated with either brackish or marginal marine environments.
Tasmanites
T. sp.
Korsodde section Levka-1 borehole Sorthat beds
Miospores
Affinities with the family Pyramimonadaceae. Found on shoreface and shoreface–offshore transition zone deposits.
Incertae sedis; affinities with Bryophyta. This spore is found in Jurassic sediments associated with the polar regions. The Sorthat Formation is among its southernmost locations.
Affinities with the family Encalyptaceae in the Bryopsida. Branching moss spores, indicating high water-depleting environments.
Extant Encalypta specimens; Staplinisporites probably come from similar genera
Stereisporites
S. antiquasporitesS. stereoides
Levka-1 borehole Sorthat beds Korsodde section
Spores
Affinities with the family Sphagnaceae in the Sphagnopsida. "Peat moss" spores, related to genera such as Sphagnum that can store large amounts of water.
Extant Sphagnum specimens; Stereisporites, Sculptisporis and Rogalskaisporites probably come from similar genera
Affinities with the Selaginellaceae in the Lycopsida. Herbaceous lycophyte flora, similar to ferns, found in humid settings. This family of spores are also the most diverse in the formation.
Extant Selaginella, typical example of Selaginellaceae. Genera like Anapiculatisporites or Densoisporites probably come from a similar or a related Plant
Affinities with Selaginellaceae and Lycopodiidae in the Lycopodiales. It was originally described as Lycopodites falcatus. The leaves of this species are more prominently anisophyllous than in the Raheto-Hettangian S. coburgensis from Franconia.
Sestrosporites
S. pseudoalveolatus
Sorthat beds Korsodde section
Spores
Affinities with the family Lycopodiaceae in the Lycopodiopsida. Lycopod spores, related to herbaceous to arbustive flora common in humid environments.
Extant Lycopodium specimens. Genera like Sestrosporites, Camarozonosporites, Retitriletes, Lycopodiumsporites and Semiretisporis probably come from a similar plant
Affinities with the Calamitaceae in the Equisetales. Horsetails are herbaceous flora found in humid environments and are flooding-tolerant. In the sections of the formation such as Korsodde, this genus has small peaks in abundance in the layers where more Equisetites stems are found.
Reconstruction of the genus Calamites, found associated with Calamospora
Affinities with Equisetaceae in the Equisetales. Related equisetalean stems are found in the Hettangian strata along Skane, Sweden. In the lagoonar sections there is correlation between bioturbation and transported Equisetites stems. Local Equisetales reached a considerable size, comparable to modern subtropical bamboos, close to lakes and in the wettest environments.
Three incomplete axes Isolated Incomplete fragments
Affinities with Calamitaceae in the Equisetales. Related equisetalean stems are found in strata of the same age along Skane, Sweden. Based on analogies with morphologically similar extant Equisetum species, it is interpreted to represent a plant of consistently moist habitats, such as marshes, lake margins or forest understorey, normally developing dense thickets.
Affinities with the genus Saccoloma, type representative of the family Saccolomataceae. This fern spore resembles those of the living genus Saccoloma, being probably from a pantropical genus found in wet, shaded forest areas.
Extant Saccoloma specimens; Annulispora probably comes from similar genera or maybe a species in the genus
Baculatisporites
B. comaumensisB. primariusB. wellmaniiB. sp.
Korsodde section Levka-1 borehole Sorthat beds
Spores
Affinities with the family Osmundaceae in the Polypodiopsida. Near fluvial current ferns, related to the modern Osmunda regalis.
Extant Osmunda specimens; Baculatisporites and Todisporites probably come from similar genera or maybe a species from the genus
Affinities with Osmundaceae in the Osmundales. Specimens assigned to this morphothype have been found in the Middle Jurassic flora of Yorkshire, associated with Todites miospores, and were originally described as Asplenites cladophleboides.
Affinities with Dicksoniaceae in the Cyatheales. It show similarities with Sphenopteris longipinnata in the morphological outline of the leaflets and the keels of the pinnate axis.
D. acutilobiumD. munsteriD. bartholliniD. cf. nilssoniiD. cf. spectabile
Vellengsby Bagagraven clay pit Hasle clay pit
Isolated pinnae
Affinities with Dipteridaceae in the Polypodiales. Dictyophyllum is a common dipteridacean genus of the mid-Mesozoic.
Dictyophyllumnilssonii specimen from an unknown location
Eboracia
E. lobifoliaE. sp.
Bagagraven clay pit Hasle clay pit Vellengsby
Isolated pinnae
Affinities with Dicksoniaceae in the Cyatheales. The Lund material is dominated by ferns belonging to the genus Eboracia (28 specimens of E. lobifolia and 14 of another Eboracia sp.). The latter has smaller pinnules than E. lobifolia.
Extant Cyathea; Zebrasporites and Cibotiumspora probably come from similar genera
"Peltaspermales"/Indet. Spermatophytes
Genus
Species
Stratigraphic position
Material
Notes
Images
Alisporites
A. grandisA. radialisA. robustusA. thomasiiA. microsaccusA. diaphanus
Levka-1 borehole Sorthat beds Korsodde section
Pollen
Affinities with the families Peltaspermaceae, Corystospermaceae or Umkomasiaceae in the Peltaspermales. Pollen of uncertain provenance that can be derived from any of the members of the Peltaspermales. The lack of distinctive characters and poor conservation make this pollen difficult to classify. Arboreal to arbustive seed ferns.
From the family Caytoniaceae in the Caytoniales. Caytoniaceae are a complex group of Mesozoic fossil floras that may be related to both Peltaspermales and Ginkgoaceae.
Type pollen of the Erdtmanithecales, related to the Gnetales. Thick tectum, infratectum of small granules, indistinct or absent foot layer. Originally thought to come from angiosperms, then suggested to come from arbustive Bennettites. It was recently found to come from Eucommiitheca, a member of the enigmatic Erdtmanithecales, reinterpreted as an unusual gymnosperm grain with a single distal colpus flanked by two subsidiary lateral colps. Is very similar to the pollen of the extant Ephedra and Welwitschia (mainly on the basis of the granular structure of the exine).
Cycadophyta
Genus
Species
Stratigraphic position
Material
Notes
Images
Butefia
B. ensiformis
Bagagraven clay pit
Leaflets
Affinities with Cycadales in the Cycadopsida. Originally described as Podozamites ensiformis.
Chasmatosporites
C. apertusC. elegansC. hiansC. majorC. minor
Levka-1 borehole Korsodde section
Pollen
Affinities with the family Zamiaceae in the Cycadales. It is among the most abundant flora recovered on the upper section of the coeval Rya Formation, and was found to be similar to the pollen of the extant Encephalartos laevifolius.
Affinities with the family Cycadaceae in the Cycadales. The structure of the exine of Clavatipollenites hughesii from Jurassic deposits is fundamentally different from that of Cretaceous grains referred to the same species, confirming observations made previously on the basis of analysis under the light microscope and suggesting a possible derivation from cycadalean rather than angiospermous plants.
Extant Cycas platyphylla. Clavatipollenites may come from a related plant
Affinities with the family Cycadaceae and Bennettitaceae. It has been found associated with the Bennetite pollen cone Bennettistemon. It increases towards the Toarcian section.
Affinities with Williamsoniaceae in the Bennettitales. Insufficient and incomplete material prevents certain assignment of Otozamites cf. reglei and Otozamites cf. mimetes
Affinities with Czekanowskiales in the Ginkgoales. This genus is related to flora from the Rhaetian–Hettangian boundary of Jameson Land, but also present in Romania.
Hartzia
H. tenuisH. sp.
Korsodde section
Leaf compressions; Cuticles
Affinities with Czekanowskiales in the Ginkgoales. Linked to the Lower Liassic flora of Greenland.
Affinities with Ginkgoaceae in the Ginkgoales. Seven species assigned to either Ginkgo or Ginkgoites have been reported from Latest Triassic to middle Jurassic strata of southern Sweden.
Extant Ginkgo, the only surviving member of the Ginkgoaceae. Monosulcites pollen is similar to the pollen of this extant species.
Solenites
S. murrayana
Korsodde section
Leaf compressions; Cuticles
Affinities with Czekanowskiales in the Ginkgoales. This species was described on the basis of individuals collected in Greenland from the Triassic–Jurassic boundary.
Affinities with Hirmeriellaceae or Araucariaceae in the Pinales. Originally Araucarioxylon württembergica. This genus is usually associated with leaf-bearing twigs referred to as Pagiophyllum, abundant in the Sorthat Formation.
Fragmentary axis compressions with preserved leaves
Affinities with Taxaceae in the Pinales. Was first identified in Bornholm. Is similar to the cretaceous Taxus huolingolensis and extant Taxus in leaf gross morphology and has papillate abaxial cuticles, probably being close to this genus.
C. dampieriC. turbatusC. microvelatusC. segmentatus
Sorthat beds Korsodde section
Pollen
Affinities with the family Araucariaceae in the Pinales. Conifer pollen from medium to large arboreal plants.
Extant Araucaria. Callialasporites may come from a related plant
Cerebropollenites
C. macroverrucosusC. thiergartii
Levka-1 borehole Sorthat beds Korsodde section
Pollen
Affinities with both Sciadopityaceae and Miroviaceae in the Pinopsida. This pollen's resemblance to extant Sciadopitys suggest that Miroviaceae may be an extinct lineage of Sciadopityaceae-like plants.
Extant Sciadopitys. Cerebropollenites likely come from a related plant
Affinities with Cupressoideae in the Cupressales. It matches with the Middle Jurassic Cyparissidium blackii from Yorkshire, England.
Dactylethrophyllum
D. ramonensis
Korsodde section
Coalified fragments; Cuticles
Affinities with Hirmeriellaceae in the Pinales. It is related to other representatives of the genus of the Toarcian of Italy and Lower Jurassic of Israel. Spheripollenites co-occurs with cuticles of Dactylethrophyllumramonensis, and the species S. psilatus may be produced by the conifer genus Dactylethrophyllum.
Affinities with the family Cupressaceae in the Pinopsida. Pollen that resembles that of extant genera such as the genus Actinostrobus and Austrocedrus, probably derived from dry environments.
Extant Austrocedrus. Exesipollenites and Perinopollenites maybe come from a related plant
Affinities with Hirmeriellaceae in the Pinales. The main genus of the Hirmeriellaceae, found in dry environments and probably fire tolerant.
Quadraeculina
Q. anellaeformis
Levka-1 borehole Sorthat beds Korsodde section
Pollen
Affinities originally suggested with the family Podocarpaceae in the Pinopsida. Quadraeculina is not comparable to pollen of any modern gymnosperm family.
Fragmentary axis compressions with preserved leaves; Coalified fragments; Cuticles
Affinities with Araucariaceae or Hirmeriellaceae in the Pinales. P. kurrii (originally P. steenstrupi) is preferred as this species is characterised by relatively broad leaves inserted at high angles to the stem. P. peregrinum has been found on the Hettangian Rønne Formation associated with hirmeriellidaceous wood of Simplicioxylon. On the Toarcian levels, is the most common plant cuticle recovered locally.
Affinities with Palissyaceae in the Palissyales. Descriptions of Palissya come mostly from coeval deposits in the Northern Hemisphere, based on a very few specimens from Sweden, Germany and America.
Affinities with Schizolepisaceae in the Pinaceae. This genus is found associated with Schizolepis on many places, making diverse authors to put both on Pinaceae.
Affinities with the Hirmeriellaceae in the Pinopsida. Spheripollenites psilatus composes up to 95% of the Lower Toarcian section and is correlated with Toarcian carbon cycle anomalies including the oceanic anoxic event, suggesting dry climates.
Stachyotaxus
S. septentrionalis
Hasle clay pit
Fragmentary axis compressions with preserved leaves
Fragmentary axis compressions with preserved leaves
Affinities with Taxaceae in the Pinales. Known only from Bornholm and belongs to an extant genus. This species is related to the Middle Jurassic floras of Yorkshire.
B. Eske Koch corroborated the presence of amber drops in the Sorthat Formation. This record represents one of the few worldwide from Jurassic layers. This amber was quoted as derived from Coniferales indet.
Thalassinoides burrowing structures, with modern related fauna, showing the ecological convergence and the variety of animals that left this Ichnogenus.