The Moltrasio Formation (also known as the Lombardische Kieselkalk Formation, Saltrio Formation, Broccatello Formation and Alpe Perino Limestone) is a geological formation in Italy and Switzerland. This Formation mostly developed in the Sinemurian stage of the Lower Jurassic, where on the Lombardian basin tectonic activity modified the current marine and terrestrial habitats. Here it developed a series of marine-related depositional settings, represented by an outcrop of 550–600 m of grey Calcarenites and Calcilutites with chert lenses and marly interbeds, that recovers the Sedrina, Moltrasio and Domaro Formations. This was mostly due to the post-Triassic crisis, that was linked locally to tectonics. The Moltrasio Formation is considered a continuation of the Sedrina Limestone and the Hettangian Albenza Formation, and was probably a shallow water succession, developed on the passive margin of the westernmost Southern Alps. It is known due to the exquisite preservation observed on the Outcrop in Osteno, where several kinds of marine biota have been recovered.

Utilities

Salnova quarry in Saltrio

The first extraction activities of the famous Saltrio & Moltrasio stone give back to the times of the ancient Romans, with modern reports of activity in Salnova quarry since 1400. In the Monte Oro area, on the southern slope of Monte Orsa, there were numerous trench quarries which were used to extract this precious rock, used both for structural constructions and for the production of artefacts and artistic works. In more recent times the mining activity has been transformed and we have moved from the extraction of stone for construction to the extraction for the production of stabilized and split crushed stone, useful for the production of motorway foundations and mixtures for the production of asphalt. To date it is the only active quarry where Saltrio stone is extracted.

Structural scheme illustrating the syn-rift context of the western Lombardian Basin during late Triassic and early Jurassic.

Above, the Moltrasio limestone emerges, a greyish-brown limestone composed of biocalcarenite and containing widespread nodules of spongolitic silica. This rock is rarely fossiliferous except in the contact areas between the Formations. At the roof of the Moltrasio Fm, a whitish yellow limestone emerges, again of marine-pelagic origin, where there is a lot of micro-diffused silica within the sediment.

Since the early 1900s, fossil finds have been known in the Salnova Quarry and in the various quarry sites present in the surroundings of this site. The first written testimonies, and subsequent revisions, are reported starting from the sixties by Giulia Sacchi Vialli. The scholar describes the fossil faunas of Saltrio by listing and detailing various taxa belonging to ammonoids, nautiloids, gastropods, crinoids, brachiopods and bivalves.

In that period, the great phase of extraction of ornamental stone using manual-mechanical methods had just ended in the quarry. Paleontologists could only recover fossils from the waste flakes near the quarry and therefore the possibility of seeing more specimens was limited to the length of manual operations. In those years, however, the quarry was acquired by Salnova SPA (1969): the purpose of the extracted material, and therefore the extraction method and processing, changed. From classic and manual extraction we move on to the use of heavy mechanical means and extraction with explosives: the moved rubble increases considerably, making it easier to observe other specimens, new lithologies and above all different faunas.

The fauna present at the base of the Moltrasio Formation is condensed and includes ammonoids of species attributed to the entire Sinemurian. The taxa attributable to the Lower Sinemurian found in the Saltrio quarries probably come from the base of the formation or have been reworked. The Formation includes taxa indicative of all the biozones between the Bucklandi Zone (Lower Sinemurian) and the Obtusum Zone, and possibly also of the Oxynotum Zone of the Upper Sinemurian, present at the base of the Formation. The contact between the Main Dolomite and the Moltrasio Formation also contains selachian teeth, glauconite and phosphated internal models of ammonites.

Description

Exposed layers at Parco delle Gole della Breggia, Southern Switzerland

The Moltrasio Limestones record a transgressive episode during which the sea flooded a fault-segmented carbonate platform. Sedimentation was slow and often interrupted, producing condensed successions with bored hardgrounds, glauconite coatings, and local phosphatic grains.

On structural highs, crinoid-rich grainstones and packstones formed encrinitic carpets that were later bored and glauconitized. In nearby grabens, coarse epiclastic calcarenites and rudites accumulated, composed of reworked Triassic platform grains later mixed with crinoid debris. Lower-lying areas preserve thin condensed horizons overlain by dark laminated micrites with sponge spicules, scattered bioclasts, and minor terrigenous and phosphatic material. Along the southern margin, the transgression is marked by reddish calcarenites and litharenites derived from basement rocks.

Basinward, these deposits grade into the spiculitic Kieselkalk, a sponge-rich lime mudstone with interbedded bioclastic and fine terrigenous layers. The Saltrio environment was complex, with different layers showing distinct conditions. In some areas, the Saltrio layers blend with the "Broccatello d'Arzo", a related limestone formation, but they can still be separated based on differences in their structure and fossil content. The region also experienced sedimentary discontinuities, where layers were not deposited continuously, likely due to tectonic activity or submarine erosion.

The stratigraphic sequence at the Galli quarry, located at an elevation of approximately 700 meters on the southeastern flank of the ridge above Saltrio, represents one of the most detailed exposures of the Saltrio Formation. This section, reaching a total thickness of about 17 meters, begins atop underlying dolomite and consists of a series of carbonate-dominated layers that reflect varying depositional conditions in a marine setting.

Closeup of the "Saltrio Limestones"

At the base, a thin dolomitic breccia layer (up to 1 meter thick) contains angular dolomite fragments embedded in a lighter calcareous-dolomitic matrix. This is overlain by a 0.3-meter-thick marly limestone with minor detrital components, displaying an olive-gray to greenish hue and iron oxide stains. Next is a 0.8-meter saccharoidal limestone with sparse marl, glauconite, and quartz grains, followed by a thin (0.1–0.2 meter) reddish-brown clay horizon.

Above this, a 1-meter oolitic limestone features intact and broken ooids in a compact calcareous cement, grading from white to grayish-yellow. This transitions into a 3-meter unit of finely to coarsely detrital marly limestone rich in organic fragments, shifting from gray-pink at the base to yellowish upward, with iron oxide patches. A 0.8-meter calcareous breccia with diverse clasts and ooids follows, exhibiting irregular surfaces.

The upper part includes a 5.5-meter marly limestone with minimal detritus, progressing from gray-ashy at the base to dark smoky due to bituminous content. The sequence concludes with a 5-meter dark limestone containing chert nodules, which become more abundant and marly toward the top, before passing into overlying cherty limestones approximately 200 meters thick.

Paleoenvironment

The nearby area at Monte Nudo basin margin was notorious for the dominance of shallow subtidal to intertidal settings with Cheirolepidiaceae conifer forests.
Paleogeography of the northwestern Neotethys showing the location of Osteno (red star)

During the late Hettangian to early Sinemurian, the western Lombardy Basin formed part of the Southern Alps area, passive margin of Adria and formed part of an evolving rift system linked to the western Tethys, where horst-and-graben tectonics created alternating shallow platforms and subsiding troughs. Structural highs such as Monte Campo dei Fiori, the Varese-Arbostora swell, and Monte Nudo defined the basin architecture, producing shallow carbonate platforms, emergent land, and subsiding depocenters. Nearby emerged areas are known at Briançonnais with the bauxite rich “Siderolitico” linked to long emersion phase (Ladinian-Bathonian) and Western Tauern with graphitic schists and quartzites & sandy marble, overall pointing to the Lombardian sector acting as a possible land bridge with Laurasia, connected with the Calcari Grigi di Noriglio Formation in the Trento carbonate Platform, based on Theropod tracks of similar size to Saltriovenator. The main inner land was the Malossa-Zandobbio palaeohigh system in the Po Plain, tied with the Saltrio area by a regional belt of positive blocks. This Highs, if assumed as a single unit probably got 1,000-3,000 km² for the inner lands and 22,500 km² in total of intermittently exposed terrain (similar in size to modern Sicily). Indicators of subaerial conditions are seen at Castello Cabiaglio-Orino, thick "terra rossa" paleosols developed directly above the Rhaetian Zu Limestone, showing rhizoliths, alveolar structures, and meteoric diagenesis. Palynological assemblages from these horizons indicate Hirmeriellaceae-dominated forests and understory of Lycopsid-Ferns, as well potential Characeae, adapted to marsh or ponded settings in a Tropical subhumid climate. These emerged lands bordered a gulf-shaped embayment, open northward, where shallow-marine carbonate platforms alternated with rapidly subsiding basins. Towards the Early Sinemurian the Arbostora swell submerged into a shallow open sea (ramp-slope), still bordered south and southwest by emerged land supported by terrigenous sands from eroded igneous/metamorphic rocks and terrestrial plants in the limestones.

Within the Moltrasio Formation, the shallowest deposits belong to the "Alpe Perino Limestone" ("Gozzano-type" marginal onlap), a small carbonate platform developed on structural highs and fault-bounded grabens. Its basal beds of Ostracod-rich mudstones and marls, with reworked Triassic lithoclasts and local plant remains (Castello Cabiaglio–Orino section), reflect restricted lagoons or marsh-like ponds on the inner platform. Then the facies evolve in repeated shallowing-upward cycles with stromatolitic-rich tidal flats and fossil-rich (gastropods, bivalves, echinoids, Dasycladales, and Foraminifera) shoals, marking a dominance of shallow subtidal to intertidal settings in the Monte Nudo basin margin, with limited terrigenous input but clear evidence of proximity to land. This unit is either part of the "Late Hettangian hypothesis" (shallower section being flooded by the Saltrio Beds) or is part the "Early Sinemurian hypothesis" (overlap with earliest Saltrio Beds on tectonic blocks and be diachronous shallow-platform vs. outer-ramp).

Artistic reconstruction of a hypothetical scene related to the Osteno Lagerstätte.
Exposed Moltrasio Fm at Monte Cornizzolo

Early marine flooding in fault-bounded grabens produced the “Viggiù-type” facies, consisting of cross-bedded coarse epiclastic calcarenites-rudites, rich in rounded ooids, algal-encrusted bioclasts, dolomitic pelites, and lithoclasts derived from exposed Triassic platforms (Hauptdolomit), later mixed with echinoderm debris. These record short-transport input from adjacent structural highs during the first phase of transgression.

On neighboring highs, pale crinoidal packstones-grainstones (“Saltrio-type”) formed as autochthonous crinoid meadows, with encrusting sponge reefs, bivalvia, brachiopods, bryozoans, and foraminiferans. Fragmentary Ichthyosaur remains and bioeroded dinosaur bones (e.g., Saltriovenator zanellai) suggest transport from nearby terrestrial sources into a proximal slope or ramp, that is, an open subtidal zone some dozen of meters depth reached by the effects of storm waves and with constant bottom currents. Pauses in sedimentation generated bored hardgrounds coated with glauconite and phosphatic crusts, indicating slow accumulation under open-marine conditions.

In intervening lows, condensed horizons (“Poaggia-type”) developed on firmgrounds, hosting abundant ammonites and other pelagic organisms, overlain by thin encrinitic calcarenites and laminated spiculitic micrites that contain resedimented crinoid grains, sponge spicules, fish-phosphate, and fine terrigenous silt, reflecting hemipelagic deposition and early slope development. At the platform margins, such as Gozzano and Monte Fenera, reddish calcarenites and litharenites with basement-derived quartz and rhyolite mark true transgressive onlap onto exposed highs. These later give way to sponge-dominated carbonate mounds (locally “Broccatello-type”), reflecting a benthic, sponge-reef-dominated carbonate factory in deeper, low-light environments during progressive platform drowning.

Regional studies link this to platform drowning amid rifting, with carbon-isotope excursions implying volcanic influences and ocean perturbations. A modern analogue is the Bahama Banks, featuring oolitic shoals and lagoons in a subtropical passive-margin setting.

Osteno Lagerstätte

Apart from the Eocene of Monte Bolca, the Sinemurian of Osteno is the only fossil deposit in Italy in which soft bodies are preserved. The Osteno site was discovered in 1964. It was recovered from a series of 6 metres (20 ft) package of fine laminated, gray, spongiolitic, micritic limestone. Coroniceras bisulcatum allowed to date the outcrop as the Bucklandi zone, lower Sinemurian. The outcrop is a good documentation of a particularly complete fauna and flora of the Lower Jurassic which is not exactly common in the Southern Alps. The Osteno outcrop, part of the formation, is worldwide known due to the exceptional preservation of mostly marine biota, including rare fossilized components, helping to understand the ecosystems of the local Sinemurian margin of the Monte Generoso Basin. The high local variety of fossils found is most likely due to unique conditions of preservation, where phosphatized soft tissues have not been observed in any fish or polychaetes, but they are common in crustaceans (33%) and also occur in a smaller percentage of teuthids (14%). Soft part preservation through phosphatization in this deposit includes the muscles and branchia of Crustaceans, fish tissues, and the digestive tracts of coleoids, polychaetes, and nematodes. These fossils are interpreted as having been preserved in a stagnant, restricted basin with anoxic conditions likely within the sediment pore waters.

Biota

Foranimifera

TaxaSpeciesLocalityMaterialNotesImages
AmmobaculitesA. sp.Castello Cabiaglio-Orino M. Sasso del FerroTestsSimple agglutinated Ammomarginulininae benthic foraminifera
InvolutinaI. cf. liassicaCastello Cabiaglio-OrinoTestsEarly Jurassic Involutinidae, typical of outer-platform wackestones
LenticulinaL. sp.Widespread in Alpe Perino & Moltrasio-Saltrio faciesTestsHyaline benthic foraminifera of Nodosariidae; typical in open-subtidal facies
ReophaxR. sp.Alpe PerinoTestsSimple agglutinated benthic foraminifera of Reophacidae
SiphovalvulinaS. sp.Alpe PerinoTestsShallow-subtidal benthic foraminifera of Siphovalvulinidae
VidalinaV. martanaAlpe Perino Moltrasio-Saltrio faciesTestsEarly Jurassic benthic foraminifera of Involutinidae-like affinity; common on Liassic platforms

Porifera

GenusSpeciesLocalityMaterialNotesImages
CorynellaC. spp.Broccatello of ArzoSpicules & ImprintsA Calcareous sponge of the family Endostomatidae.
EndostomaE. spp.Broccatello of ArzoSpicules & ImprintsA Calcareous sponge of the family Endostomatidae.
StellispongiaS. spp.Broccatello of ArzoSpicules & ImprintsA Calcareous sponge of the family Stellispongiidae.
NeuroporaN. cf. undulata N. spp.Cava Salnova quarry Galli QuarrySpicules & ImprintsA Demosponge of the family Neuroporidae. A notorious reef developing genus

Brachiopoda

GenusSpeciesLocalityMaterialNotesImages
ArzonellinaA. exoticaBroccatello of ArzoShellsA Terebratulidan of the family Arzonellinidae.
AulacothyrisA. waterhousiBroccatello of ArzoShellsA Terebratulidan of the family Zeilleriidae.Aulacothyris
CirpaC. fronto C. briseis C. variabilisBroccatello of Arzo Cava Salnova quarryShellsA Rhynchonellidan of the family Wellerellidae. Identified originally as "Rhynchonella variabilis".
FurcirhynchiaF. cf. melvilleiBroccatello of ArzoShellsA Rhynchonellidan of the family Rhynchonellidae.
LiospiriferinaL. rostrataBroccatello of Arzo Cava Salnova quarryShellsA Rhynchonellidan of the family Spiriferinidae. Was identified originally as "Spiriferina haasi".Liospiriferina specimens
LobothyrisL. punctata L. andleri L. subpunctata L. ovatissinia L. sp.Broccatello of Arzo Cava Salnova quarry Galli QuarryShellsA Terebratulidan of the family Lobothyrididae. Was identified originally as "Terebratula punctata".Lobothyris
PrionorhynchiaP. belemnitica P. greppini P. flabellum P. ssp.Broccatello of ArzoShellsA Rhynchonellidan of the family Prionorhynchiidae.
RimirhynchiaR. aff. anglicaBroccatello of ArzoShellsA Rhynchonellidan of the family Rhynchonellidae.
RhynchonellaR. acanthica R. binodosa R. belemnitica R. latifrons R. rimata R. cf.polyptychaBroccatello of Arzo Cava Salnova quarry Galli QuarryShellsA Rhynchonellidan of the family Wellerellidae.Rhynchonella
RhynchonellinaR. alpina R. ssp.Broccatello of ArzoShellsA Rhynchonellidan of the family Dimerellidae.
TetrarhynchiaT. spp.Broccatello of ArzoShellsA Rhynchonellidan of the family Tetrarhynchiidae.
TerebratulaT. scissa T. rhomboedrica T. prumusCava Salnova quarry Galli QuarryShellsA Terebratulidan of the family Terebratulidae.
SpiriferinaS. expansa S. hartmanni S. oxigona S. walcotti S. rupestris S. rostrata S. haasi S. tumida S. cf. spirigeroidesBroccatello of Arzo Cava Salnova quarryShellsA Rhynchonellidan of the family Spiriferinidae.Spiriferina
SulcirostraS. alpinaBroccatello of ArzoShellsA Rhynchonellidan of the family Dimerellidae.
ViallithyrisV. gozzanensis V. adnetensisBroccatello of Arzo Cava Salnova quarryShellsA Rhynchonellidan of the family Rhynchonellidae.
ZeilleriaZ. sarthacensis Z. choffati Z. cor Z. ewaldi Z. stapia Z. cornuta Z. subnumismalis Z. (Zeilleria) indentataBroccatello of Arzo Cava Salnova quarry Galli QuarryShellsA Terebratulidan of the family Zeilleriidae.Zeilleria

Bryozoa

GenusSpeciesLocalityMaterialNotesImages
CerioporaC.? cf. orbignyiCava Salnova quarry Galli QuarryImprintsA Cyclostomatidan of the family Cerioporidae.

Nematoida

GenusSpeciesLocationMaterialNotesImages
EophasmaE. jurasicumOstenoSix specimens with soft partsA nematode, type member of the family Eophasmidae. A rare find, since nematode fossils are absent in most Mesozoic marine rocks.Eophasma

Annelida

GenusSpeciesLocationMaterialNotesImages
MelanoraphiaM. maculataOstenoEleven specimens, complete with soft partsA polychaete, member of the group Phyllodocemorpha with incertae sedis assignation, suggested to be a member of Lumbrineridae.Melanoraphia
TerebellaT. lapilloides T. spp.Broccatello of Arzo Cava Salnova quarryTubiform structuresA terebellid, member of the family TerebellidaeTerebella specimen

Arthropoda

GenusSpeciesLocationMaterialNotesImages
AegerA. foesteri A. muensteri A. robustus A. rostrospinatus A. elongatus A. macropusOstenoMultiple specimens fairly preservedA decapod of the family Aegeridae. This genus is the most abundant local crustacean, and was a shrimp-like creature that was probably necrophagous.Aeger foersteri
ColeiaC. vialiii C. medilerranea C. cf. antiqua C. pinnnaiOstenoMultiple specimens fairly preservedA decapod of the family Coleiidae. Probably a bottom dweller predatory crustacean.Coleia viallii
ErymaE. sinemurianaOsteno41 specimens, fairly preservedA decapod of the family Erymidae. The species was originally identified as Pustulina sinemuriana.
GlypheaG. tricarinataOsteno137 complete and fragmentary specimensA decapod of the family Erymidae.
MecochirusM. germariOsteno81 complete and fragmentary specimens,A decapod of the family Mecochiridae.Mecochirus germari
OstenocarisO. cypriformisOstenoMultiple specimens, complete and incompleteA thylacocephalan of the family Ostenocarididae. It is the most common fossil of the formation, and the main identified thylacocephalan from the formation.Ostenocaris
OstenolimulusO. latusOstenoA nearly complete specimenA horseshoe crab of the family Limulidae. This genus represents the oldest Jurassic limulid and only the second xiphosuran known from Italy. Also indicates that this family were a diverse and widespread clade during the Jurassic.
OstenosculdaO. teruzziiOstenoSingle Specimen fairly preservedAn early unipeltatan mantis shrimp
PalaeastacusP. meyeriOstenoMultiple specimens fairly preservedA decapod of the family Erymidae. The species was originally included in the genus Eryma as E. meyeri, although the specimens reveal morphological characters diagnostic of Palaeastacus.
PhlyctisomaP. sinemurianaOsteno10 specimens, in a fairly good state of preservationA decapod of the family Erymidae. The species was originally identified as Pustulina sinemuriana.
PseudoglypheaP. ancylochelisOsteno7 complete specimens, in a fairly good stateA decapod of the family Litogastroidae. Probably a bottom dweller predatory crustacean.
TeruzzichelesT. popeyeiOstenoVarious specimens fairly preservedA decapod of the family Polychelidae

Mollusks

GenusSpeciesLocalityMaterialNotesImages
AgassicerasA. nodosaries A. aff. scipionianumCava Salnova quarryShellsAn Ammonitidan of the family Arietitidae.
AngulaticerasA. sp.OstenoShellsAn ammonitidan of the family Schlotheimiidae.
AnticonulusA. profunde-umbilicatusCava Salnova quarry Galli QuarryShellsA Snail of the family Trochidae.
ArietitesA. bucklandi A. raricostatus A. rapidecrescens A. ceratitoides A. kridioides A. dimorphus A. arnoui A. conybeari A. sauzeanusCava Salnova quarry OstenoShellsAn Ammonitidan of the family Arietitidae.Arietites bucklandi
ArniocerasA. cf. bodleyiCava Salnova quarry Galli QuarryShellsAn Ammonitidan of the family Arietitidae.
AstarteA. praeobliquaCava Salnova quarryShellsA Clam of the family Astartidae. Some shells identified as Cardium probably belong to this genus.Astarte
AsterocerasA. obtusum A. stellare A. retusum A. confusumCava Salnova quarry Burgioli quarry Galli quarryShellsAn Ammonitidan of the family Arietitidae.Asteroceras
AulacocerasA. ("Ausseites") italicusCava Salnova quarry Galli QuarryPhagmoconesA Belemnite of the family Aulacoceratidae.
AviculaA. sinemuriensis A. (Oxytoma) dumortieriCava Salnova quarry Galli QuarryShellsA Pearl Oyster of the family Pteriidae.
BathrotomariaB. cf. trocheataCava Salnova quarry Galli QuarryShellsA Snail of the family Pleurotomariidae.
CalvaentoliumC. hehliiCava Salnova quarryShellsA Scallop of the family Pectinoidae. It was identified originally as "Pecten (Pseudamusium) hehlii".
CardiniaC. hybrida C. similis C. rugosa C. hybrida C. concinna C. depressa C. spp.Cava Salnova quarry Galli QuarryShellsA Clam of the family Cardiniidae.
CardiumC.? italicum C. cf. multicostatum C. spp.Cava Salnova quarryShellsA Clam of the family Cardiniidae.
CenocerasC. amorettii C. stoppanii C. intermedium C. arare C. spreaficoi C. breislacki C. striatum C. sturi C. balsamocrivellii C. affinis C. brancoi C. ornatusCava Salnova quarry Galli QuarryShellsA Nautilidan of the family Cenoceratidae. Cenoceras was identified as member of the genus Nautilus originally.Cenoceras reconstruction
CoronicerasC. cf.gmuendense C. rotiformis C. bucklandi C. orbiculatus C. bisulcatusCava Salnova quarry OstenoShellsAn Ammonitidan of the family Arietitidae.Coroniceras specimen
CrucilobicerasC. cf. tardarmatumGalli QuarryShellsAn Ammonitidan of the family Eoderoceratidae.
EctocentritesE. sp.OstenoShellsAn ammonitidan of the family Lytoceratidae.
EntoliumE. hehli E. calvum E. disciformeCava Salnova quarry Galli QuarryShellsA Scallop of the family Pectinoidae.
EparietitesE. impendensCava Salnova quarry Burgioli quarry Galli QuarryShellsAn Ammonitidan of the family Arietitidae
EuasterocerasE. cf.turneriCava Salnova quarry Galli QuarryShellsAn Ammonitidan of the family Arietitidae
ChlamysC. textoriaCava Salnova quarry Galli QuarryShellsA Scallop of the family Pectinoidae It was identified as "Pecten (Chlamys) textorius".Chlamys
CoelostylinaC. lepontinaCava Salnova quarry Galli QuarryShellsA Snail of the family Coelostylinidae.
DiscohelixD. sp.Broccatello of ArzoShellsA Snail of the family Discohelicidae.Discohelix
DiscotomaD.? aff. ornataCava Salnova quarryShellsA Snail of the family Pleurotomariidae.
FimbriaF. semireticulata F. (Sphaeriola) sp.Cava Salnova quarryShellsA Clam of the family Lucinidae.Fimbria
GresslyaG. lunulataCava Salnova quarry Galli QuarryShellsA Clam of the family Pleuromyidae.
GoniomyaG. verbanaCava Salnova quarryShellsA Clam of the family Lucinidae.
GryphaeaG. arcuataCava Salnova quarryShellsAn Oyster of the family Gryphaeidae.Fimbria
LimaL. (Radula) succincta L. (Radula) valmariannae L. (Radula) hettangiensis L. spp.Broccatello of Arzo Cava Salnova quarryShellsA File Clam of the family Limidae.Lima
LucinaL. ? liasinaCava Salnova quarryShellsA Clam of the family Lucinidae.Lucina
LytocerasL. sp.Galli QuarryShellsAn Ammonitidan of the family Lytoceratidae
MactromyaM. arenacea M. cingulata M. problematicaCava Salnova quarry Galli QuarryShellsAn Adapedont of the family Edmondiidae.
MicroderocerasM. birchiiCava Salnova quarryShellsAn Ammonitidan of the family Eoderoceratidae.Microderoceras specimen
ModiolusM. vomer M. cf.scalpraCava Salnova quarry Galli QuarryShellsA Mussel of the family Mytilidae. Identified as the genus "Modiola", now junior synonym of Modiolus.Modiolus
MyoconchaM. scabra M. rugosa M. ssp.Cava Salnova quarry Galli QuarryShellsA Clam of the family Pleuromyidae.
NannobelusN. cf. acutusCava Salnova quarry Galli QuarryPhagmoconesA Belemnite of the family Belemnitidae.
OstenoteuthisO. siroiOstenoSeveral specimens with decent preservationA coleoid of the family Ostenoteuthidae. A unique group of coeloids, only found at this location.Ostenoteuthis
OstreaO. cf. chillyensis O. electraBroccatello of Arzo Cava Salnova quarryShellsAn Oyster of the family Ostreidae.Ostrea
OxynoticerasO. oxynotum O. actinotumCava Salnova quarry Burgioli quarry Galli Quarry Osteno OutcropShellsAn Ammonitidan of the family Oxynoticeratidae.Oxynoticeras specimen
PaltechiocerasP. (Plesechioceras) demissum P.(Plesechioceras) delicatumBurgioli quarryShellsAn Ammonitidan of the family Echioceratidae.
ParacoronicerasP. cf. gmuendenseCava Salnova quarryShellsAn Ammonitidan of the family Arietitidae.
ParadasycerasP. stellaCava Salnova quarry Galli QuarryShellsAn Ammonitidan of the family Juraphyllitidae.
ParallelodonP. (Grammatodon) cypriniformeCava Salnova quarry Galli QuarryShellsA Clam of the family Parallelodontidae.
PectenP. (Pseudamusium) diblasii P. (Pseudamusium) hehlii P. (Chlamys) subalpinus P. (Chlamys) textorius P. spp.Broccatello of Arzo Cava Salnova quarryShellsA Scallop of the family Pectinoidae.Pecten
PholadomyaP. sp.Cava Salnova quarryShellsA Clam of the family Pholadomyidae.Pholadomya
PlagiostomaP. giganteum P. stabilei P. spp.Broccatello of Arzo Cava Salnova quarryShellsA File Clam of the family Limidae. Identified originally as "Lima (Plagiostoma) gigantea".Plagiostoma
PleuromyaP. cf. angusta P. saltriensis P. galatheaCava Salnova quarry Galli QuarryShellsA Clam of the family Pleuromyidae.
PleurotomariaP. oblita P. anglica P. escheri P. granulatocincta P. hettangiensis P. saltriensis P. cf. gigas P. sp.Broccatello of Arzo Cava Salnova quarryShellsA Snail of the family Pleurotomariidae.Pleurotomaria
PromicrocerasP. planicostaBurgioli quarryShellsAn Ammonitidan of the family Eoderoceratidae.
PtychomphalusP. expansusCava Salnova quarryShellsA Snail of the family Eotomariidae.
PyrgotrochusP. basilicus P. cf.basilicusCava Salnova quarry Galli QuarryShellsA Snail of the family Pleurotomariidae.
SchlotheimiaS.(Charmasseiceras?) sp.Galli QuarryShellsAn Ammonitidan of the family Schlotheimiidae
TerquemiaT. hebertiCava Salnova quarryShellsA Clam of the family Prospondylidea.
TrochusT. spBroccatello of Arzo Cava Salnova quarryShellsA Snail of the family Trochidae.Trochus
TrochotomaT. subturritaCava Salnova quarryShellsA Snail of the family Pleurotomariidae.
UncinoteuthisU. cuvieriOstenoAn incomplete specimen consisting of the arm crown onlyA coleoid of the family Ostenoteuthidae.Uncinoteuthis
VermicerasV. spiratissimum V. cf.spiratissimumCava Salnova quarryShellsAn Ammonitidan of the family Arietitidae.Vermiceras specimen

Echinoderms

GenusSpeciesLocationMaterialNotesImages
HispidocrinusH. scalarisPradalungaMultiple ossiclesA Crinoidean, member of the family Isocrininae inside Isocrinida.
IsocrinusI. tuberculatus I. spp.Broccatello of Arzo Cava Salnova quarry M. Campo dei Fiori Ubiale-ClanezzoMultiple ossiclesA Crinoidean, member of the family Isocrininae inside Isocrinida.
MillericrinusM. cf. adneticusM. di GroneMultiple ossiclesA Crinoidean, member of the family Millericrinidae inside Millericrinida.
MiocidarisM. amaltheiCava Salnova quarrySpines & AmbulacrumAn Echinoidean of the family Miocidaridae.
PalaeocomaP. milleriMount Campo dei Fiori Ubiale-ClanezzoMSNVI 044/017, dorsally-ventrally oriented ophiuroidAn Ophiuridan of the family Ophiodermatidae. Extant tropical species like Ophioderma are benthic predators and scavengers that show the same short spines seen in Palaeocoma.Palaeocoma milleri
PentacrinitesP. fasciculosus P. tuberculosus P. subangularis P. spp.Broccatello of Arzo Cava Salnova quarryMultiple ossiclesA Sea lily of the family Pentacrinitidae.Pentacrinites
PlegiocidarisP. falsaniCava Salnova quarry Galli QuarrySpines & AmbulacrumAn Echinoidean of the family Cidaridae.Plegiocidaris

Hemichordata

GenusSpeciesLocationMaterialNotesImages
MegaderaionM. sinemurienseOstenoSix specimens with soft partsAn acorn worm, member of the family Harrimaniidae inside Enteropneusta. This genus was probably benthic, linked with burrowing systems.Modern relative, Saccoglossus

Chondrichthyes

GenusSpeciesLocationMaterialNotesImages
HybodontiformesIndeterminateOstenoTeethA shark, member of the family Hybodontiformes.
MyriacanthidaeGen et sp. novOstenoVirtually complete skull with associated parts of the branchial skeletonA chimaerid, member of the family Myriacanthidae inside Chimaeriformes.
NotidanoidesN. arzoensisBroccatello of Arzo Cava Salnova quarryTeethA Crassodontidanidae Hexanchiform
OstenoselacheO. stenosomaOstenoOne complete articulated specimenA euselachiian of the family Ostenoselachidae. A genus only found in the Osteno Formation. With an elongated eel-shaped morphology, the specimens found measure at least 240 mm in length, with a neurocranium with a long, complex rostrum. It has been suggested it was an animal able to produce electric shocks, probably to kill prey.Ostenoselache
PalaeospinaxP. pinnaiOstenoA single specimen in four contiguous pieces and including parts of the dentition and squamationA shark of the family Palaeospinacidae.
SqualorajaS. sp.OstenoOne complete articulated specimenA chimaerid of the family Squalorajidae. This genus belongs to a lineage of ray-like chimaeras.Squaloraja
SphenodusS. helveticus S. cf. alpinus S. stschurowskiiBroccatello of Arzo Cava Salnova quarryTeethAn Orthacodontidae Synechodontiform

Actinopterygii

GenusSpeciesLocationMaterialNotesImages
DapediumD. sp.OstenoMSNM V620 -MSNM V630, caudal fin with associated scalesHolostei of the family Dapediidae.Dapedium
DorsetichthysD. becheiOstenoMSNM V409-MSNM V638: nearly complete articulated skeleton; MSNM V547-575: patch of articulated scales; MSNM V558: two isolated head skeletons; MSNM V572: poorly preserved head skeleton; MSNM V573: isolated caudal and caudal peduncle; MSNM V595: incomplete head skeleton; MSNM V604: patch of articulated scales; MSNM V629: nearly complete articulated skeleton; MSNM V633: incomplete articulated skeleton; MSNM V650: isolated head skeleton, in part and counterpart; MSNM V654: patch of articulated scales. MSNM V658: largely incomplete articulated skeleton; MSNM V662: partially complete articulated skeleton; MSNM V664: nearly complete articulated skeleton.A member of the family Dorsetichthyidae. It is the main fish recovered on the formation.Dorsetichthys
FuroF. ("Eugnathus") sp.OstenoMSNM V623, an incomplete articulated skeleton; MSNM V634, an incomplete partially articulated skeleton.A member of the family Furidae.Furo
OstenolepisO. marianiiOstenoMSNM V617, an almost complete, articulated skeleton; MSNM V538, largely incomplete specimen; MSNM V554, incomplete articulated skeleton; MSNM V652, incomplete twisted articulated skeleton; MSNM V618-19, skull & toracic section specimen. Other 3 SpecimensA member of the family Palaeoniscimorpha. Previously referred to Cosmolepis and Pteroniscus.Ostenolepis
PeripeltopleurusP. jurassicusOstenoMSNM V659, almost complete specimen lacking the tailA member of the family Wushaichthyidae. Represents the only known Jurassic survivor of the order PeltopleuriformesPeripeltopleurus
PholidolepisP. teruzziiOstenoMSNM V564-V621: almost complete, articulated skeleton. MSNM V529, MSNM V562, MSNM V533, MSNM V565, MSNM V596, MSNM V645, MSNM V648, MSNM V581, MSNM V649: isolated caudals; MSNM V540: incomplete articulated skeleton; MSNM V556, MSNM V559, MSNM V567, MSNM V568, MSNM V579, MSNM V594: isolated head skeleton; MSNM V563: disarticulated and largely incomplete skeleton; MSNM V570: incomplete articulated skeleton; MSNM V613: almost com-plete articulated skeleton; MSNM V625: incomplete articulated skeleton; MSNM V641: complete, articulated skeleton; MSNM V646: incomplete articulated skeleton MSNM V656: isolated pectoral girdle.A member of the family Pholidophoridae.Pholidolepis

Sarcopterygii

GenusSpeciesLocationMaterialNotesImages
HolophagusH. cf. guloOstenoMSNM V530, V561, V587, V586, V588, V589, V591, V592, V593, V597, V598, V601, V599, V600, V667, V602, V607, V666, V668, isolated scales; MSNM V543, isolated urohyal.A Coelacanth of the family Latimeriidae. Isolated 20 mm scales suggests a taxon up to 75 cm long.Holophagus

Icthyosaurs

In 2016 new vertebrate remains were discovered in the Salnova quarry, the remains are being studied to understand if it is a new dinosaur or some other creature. Latter has been confirmed to be Marine Diapsid material.

GenusSpeciesLocalityMaterialNotesImages
IchthyosaurusI. spp.Galli Quarry3 Vertebrae ImprintsA Neoichthyosaurian of the family Ichthyosauridae.Ichthyosaurus
TemnodontosaurusT. cf.platyodonCava Salnova quarryIsolated Tooth Isolated VertebraeA Neoichthyosaurian of the family Temnodontosauridae. Quoted on the 1880s, specimen that apparently has never been described or figured and whose present repository is unknownTemnodontosaurus platyodon

Pterosaurs

GenusSpeciesLocalityMaterialNotesImages
PterosauriaIndeterminateCava Salnova quarryIsolated ToothA Pterosaur, referred originally to "Pterodactylus longirostris"Bergamodactylus, an older (Triassic) Pterosaur from Italy

Dinosaurs

GenusSpeciesLocalityMaterialNotesImages
SaltriovenatorS. zanellaiCava Salnova quarryMSNM V3664, very fragmentary and disarticulated skeleton MSNM V3659, one maxillary or dentary tooth (Referred material)A potential Ceratosaur. Traces on the bones show that the dinosaur carcass remained exposed to the water-sediment interface for months or years, long enough to first be defleshed by mobile scavengers, then colonized by a microbial community that spanned the bone–water interface, which in turn attracted slow-moving grazers and epibionts.Saltriovenator holotype

Algae

Potential Dasycladaceae are seen on intertidal facies, while Characeae are seen at the base.

TaxaSpeciesLocalityMaterialNotesImages
CayeuxiaC. spp.Alpe Perino; Castello Cabiaglio-OrinoImprintsCalcareous green alga of Udoteaceae; typical of shallow subtidal facies
ThaumatoporellaT. spp.Alpe Perino LimestoneImprintsEncrusting calcified alga, typical of Liassic shallow-marine platforms

Plants

GenusSpeciesLocationMaterialNotesImages
BrachyphyllumB. saportum B. spp.Osteno Dintorni di MoltrasioBranched ShootsA coniferophytan of the family Cheirolepidiaceae.
CallialasporitesC. spp.Castello Cabiaglio-OrinoPollenAraucariaceae ConiferExtant Araucaria.
CerebropollenitesC. macroverrucosusCastello Cabiaglio-OrinoPollenSciadopityaceae ConiferExtant Sciadopitys.
ClassopollisC. torosusCastello Cabiaglio-OrinoPollenCheirolepidiaceae Conifer, dominant palynomorph
ClathropterisC. platyphyllaCarate-LarioFrondsA Fern of the family Dipteridaceae
ConcavisporitesC. crassexiniusCastello Cabiaglio-OrinoSporesTrilete spore of Lycophyte affinity
CtenopterisC. grandisMoltrasioFrondsA Fern of the family Polypodiaceae
EquisititesE. bunburyanusOstenoStemsA Horsetail of the family Equisetaceae.
GliscopollisC. meyerianaCastello Cabiaglio-OrinoPollenTypical of Jurassic-Cretaceous Cheirolepidiaceae
GranulatisporitesG. punctatusCastello Cabiaglio-OrinoSporesTrilete spore of Lycopodiales
LomatopterisL. jurensisCarate-LarioFrondsA "Seed Fern", affinities with Cyclopteridaceae.Lomatopteris jurensis
LycopodiaciditesL. rugulatusCastello Cabiaglio-OrinoSporesTrilete spore of LycopodiaceaeExtant Lycopodium.
MicroreticulatisporitesM. fuscusCastello Cabiaglio-OrinoSporesTrilete spore of probable Lycophyte origin
OtozamitesO. bunburyanus O. bechei O. latior O. brevifolius O. pterophylloides O. sp.Osteno Moltrasio Carate-LarioLeafletsA spermatopsidan, member of Bennettitales.
PachypterisP. cf. rhomboidalisOstenoFrondsA "Seed Fern" of the family Corystospermaceae.
PagiophyllumP. peregrinum P. sp.Osteno Carate-LarioBranched ShootsA coniferophytan of the family Araucariaceae.
PorcellisporaP. longdonensisCastello Cabiaglio-OrinoSporesAffinities with Bryophyta or Marchantiophyta, potentially Riellaceae.
ThinnfeldiaT. rhomboidalis T. obtusaCarate-LarioFrondsA "Seed Fern" of the family Corystospermaceae.
UvaesporitesU. argenteaeformisCastello Cabiaglio-OrinoSporesTrilete spore of Selaginellaceae.Extant Selaginella.
WilliamsoniaW. sp.OstenoReproductive OrganA spermatopsidan, member of Bennettitales.Williamsonia
WiddringtonitesW. baehmanniMombelloBranched ShootsA coniferophytan of the family Araucariaceae.
ZamitesZ. sp.OstenoLeafletsA spermatopsidan, member of Bennettitales.

See also