Moltrasio Formation
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The Moltrasio Formation (also known as the Lombardische Kieselkalk Formation, Saltrio Formation, Broccatello Formation and Alpe Perino Limestone) is a geological formation in Italy and Switzerland. This Formation mostly developed in the Sinemurian stage of the Lower Jurassic, where on the Lombardian basin tectonic activity modified the current marine and terrestrial habitats. Here it developed a series of marine-related depositional settings, represented by an outcrop of 550–600 m of grey Calcarenites and Calcilutites with chert lenses and marly interbeds, that recovers the Sedrina, Moltrasio and Domaro Formations. This was mostly due to the post-Triassic crisis, that was linked locally to tectonics. The Moltrasio Formation is considered a continuation of the Sedrina Limestone and the Hettangian Albenza Formation, and was probably a shallow water succession, developed on the passive margin of the westernmost Southern Alps. It is known due to the exquisite preservation observed on the Outcrop in Osteno, where several kinds of marine biota have been recovered.
Utilities

The first extraction activities of the famous Saltrio & Moltrasio stone give back to the times of the ancient Romans, with modern reports of activity in Salnova quarry since 1400. In the Monte Oro area, on the southern slope of Monte Orsa, there were numerous trench quarries which were used to extract this precious rock, used both for structural constructions and for the production of artefacts and artistic works. In more recent times the mining activity has been transformed and we have moved from the extraction of stone for construction to the extraction for the production of stabilized and split crushed stone, useful for the production of motorway foundations and mixtures for the production of asphalt. To date it is the only active quarry where Saltrio stone is extracted.

Above, the Moltrasio limestone emerges, a greyish-brown limestone composed of biocalcarenite and containing widespread nodules of spongolitic silica. This rock is rarely fossiliferous except in the contact areas between the Formations. At the roof of the Moltrasio Fm, a whitish yellow limestone emerges, again of marine-pelagic origin, where there is a lot of micro-diffused silica within the sediment.
Since the early 1900s, fossil finds have been known in the Salnova Quarry and in the various quarry sites present in the surroundings of this site. The first written testimonies, and subsequent revisions, are reported starting from the sixties by Giulia Sacchi Vialli. The scholar describes the fossil faunas of Saltrio by listing and detailing various taxa belonging to ammonoids, nautiloids, gastropods, crinoids, brachiopods and bivalves.
In that period, the great phase of extraction of ornamental stone using manual-mechanical methods had just ended in the quarry. Paleontologists could only recover fossils from the waste flakes near the quarry and therefore the possibility of seeing more specimens was limited to the length of manual operations. In those years, however, the quarry was acquired by Salnova SPA (1969): the purpose of the extracted material, and therefore the extraction method and processing, changed. From classic and manual extraction we move on to the use of heavy mechanical means and extraction with explosives: the moved rubble increases considerably, making it easier to observe other specimens, new lithologies and above all different faunas.
The fauna present at the base of the Moltrasio Formation is condensed and includes ammonoids of species attributed to the entire Sinemurian. The taxa attributable to the Lower Sinemurian found in the Saltrio quarries probably come from the base of the formation or have been reworked. The Formation includes taxa indicative of all the biozones between the Bucklandi Zone (Lower Sinemurian) and the Obtusum Zone, and possibly also of the Oxynotum Zone of the Upper Sinemurian, present at the base of the Formation. The contact between the Main Dolomite and the Moltrasio Formation also contains selachian teeth, glauconite and phosphated internal models of ammonites.
Description

The Moltrasio Limestones record a transgressive episode during which the sea flooded a fault-segmented carbonate platform. Sedimentation was slow and often interrupted, producing condensed successions with bored hardgrounds, glauconite coatings, and local phosphatic grains.
On structural highs, crinoid-rich grainstones and packstones formed encrinitic carpets that were later bored and glauconitized. In nearby grabens, coarse epiclastic calcarenites and rudites accumulated, composed of reworked Triassic platform grains later mixed with crinoid debris. Lower-lying areas preserve thin condensed horizons overlain by dark laminated micrites with sponge spicules, scattered bioclasts, and minor terrigenous and phosphatic material. Along the southern margin, the transgression is marked by reddish calcarenites and litharenites derived from basement rocks.
Basinward, these deposits grade into the spiculitic Kieselkalk, a sponge-rich lime mudstone with interbedded bioclastic and fine terrigenous layers. The Saltrio environment was complex, with different layers showing distinct conditions. In some areas, the Saltrio layers blend with the "Broccatello d'Arzo", a related limestone formation, but they can still be separated based on differences in their structure and fossil content. The region also experienced sedimentary discontinuities, where layers were not deposited continuously, likely due to tectonic activity or submarine erosion.
The stratigraphic sequence at the Galli quarry, located at an elevation of approximately 700 meters on the southeastern flank of the ridge above Saltrio, represents one of the most detailed exposures of the Saltrio Formation. This section, reaching a total thickness of about 17 meters, begins atop underlying dolomite and consists of a series of carbonate-dominated layers that reflect varying depositional conditions in a marine setting.

At the base, a thin dolomitic breccia layer (up to 1 meter thick) contains angular dolomite fragments embedded in a lighter calcareous-dolomitic matrix. This is overlain by a 0.3-meter-thick marly limestone with minor detrital components, displaying an olive-gray to greenish hue and iron oxide stains. Next is a 0.8-meter saccharoidal limestone with sparse marl, glauconite, and quartz grains, followed by a thin (0.1–0.2 meter) reddish-brown clay horizon.
Above this, a 1-meter oolitic limestone features intact and broken ooids in a compact calcareous cement, grading from white to grayish-yellow. This transitions into a 3-meter unit of finely to coarsely detrital marly limestone rich in organic fragments, shifting from gray-pink at the base to yellowish upward, with iron oxide patches. A 0.8-meter calcareous breccia with diverse clasts and ooids follows, exhibiting irregular surfaces.
The upper part includes a 5.5-meter marly limestone with minimal detritus, progressing from gray-ashy at the base to dark smoky due to bituminous content. The sequence concludes with a 5-meter dark limestone containing chert nodules, which become more abundant and marly toward the top, before passing into overlying cherty limestones approximately 200 meters thick.
Paleoenvironment


During the late Hettangian to early Sinemurian, the western Lombardy Basin formed part of the Southern Alps area, passive margin of Adria and formed part of an evolving rift system linked to the western Tethys, where horst-and-graben tectonics created alternating shallow platforms and subsiding troughs. Structural highs such as Monte Campo dei Fiori, the Varese-Arbostora swell, and Monte Nudo defined the basin architecture, producing shallow carbonate platforms, emergent land, and subsiding depocenters. Nearby emerged areas are known at Briançonnais with the bauxite rich “Siderolitico” linked to long emersion phase (Ladinian-Bathonian) and Western Tauern with graphitic schists and quartzites & sandy marble, overall pointing to the Lombardian sector acting as a possible land bridge with Laurasia, connected with the Calcari Grigi di Noriglio Formation in the Trento carbonate Platform, based on Theropod tracks of similar size to Saltriovenator. The main inner land was the Malossa-Zandobbio palaeohigh system in the Po Plain, tied with the Saltrio area by a regional belt of positive blocks. This Highs, if assumed as a single unit probably got 1,000-3,000 km² for the inner lands and 22,500 km² in total of intermittently exposed terrain (similar in size to modern Sicily). Indicators of subaerial conditions are seen at Castello Cabiaglio-Orino, thick "terra rossa" paleosols developed directly above the Rhaetian Zu Limestone, showing rhizoliths, alveolar structures, and meteoric diagenesis. Palynological assemblages from these horizons indicate Hirmeriellaceae-dominated forests and understory of Lycopsid-Ferns, as well potential Characeae, adapted to marsh or ponded settings in a Tropical subhumid climate. These emerged lands bordered a gulf-shaped embayment, open northward, where shallow-marine carbonate platforms alternated with rapidly subsiding basins. Towards the Early Sinemurian the Arbostora swell submerged into a shallow open sea (ramp-slope), still bordered south and southwest by emerged land supported by terrigenous sands from eroded igneous/metamorphic rocks and terrestrial plants in the limestones.
Within the Moltrasio Formation, the shallowest deposits belong to the "Alpe Perino Limestone" ("Gozzano-type" marginal onlap), a small carbonate platform developed on structural highs and fault-bounded grabens. Its basal beds of Ostracod-rich mudstones and marls, with reworked Triassic lithoclasts and local plant remains (Castello Cabiaglio–Orino section), reflect restricted lagoons or marsh-like ponds on the inner platform. Then the facies evolve in repeated shallowing-upward cycles with stromatolitic-rich tidal flats and fossil-rich (gastropods, bivalves, echinoids, Dasycladales, and Foraminifera) shoals, marking a dominance of shallow subtidal to intertidal settings in the Monte Nudo basin margin, with limited terrigenous input but clear evidence of proximity to land. This unit is either part of the "Late Hettangian hypothesis" (shallower section being flooded by the Saltrio Beds) or is part the "Early Sinemurian hypothesis" (overlap with earliest Saltrio Beds on tectonic blocks and be diachronous shallow-platform vs. outer-ramp).


Early marine flooding in fault-bounded grabens produced the “Viggiù-type” facies, consisting of cross-bedded coarse epiclastic calcarenites-rudites, rich in rounded ooids, algal-encrusted bioclasts, dolomitic pelites, and lithoclasts derived from exposed Triassic platforms (Hauptdolomit), later mixed with echinoderm debris. These record short-transport input from adjacent structural highs during the first phase of transgression.
On neighboring highs, pale crinoidal packstones-grainstones (“Saltrio-type”) formed as autochthonous crinoid meadows, with encrusting sponge reefs, bivalvia, brachiopods, bryozoans, and foraminiferans. Fragmentary Ichthyosaur remains and bioeroded dinosaur bones (e.g., Saltriovenator zanellai) suggest transport from nearby terrestrial sources into a proximal slope or ramp, that is, an open subtidal zone some dozen of meters depth reached by the effects of storm waves and with constant bottom currents. Pauses in sedimentation generated bored hardgrounds coated with glauconite and phosphatic crusts, indicating slow accumulation under open-marine conditions.
In intervening lows, condensed horizons (“Poaggia-type”) developed on firmgrounds, hosting abundant ammonites and other pelagic organisms, overlain by thin encrinitic calcarenites and laminated spiculitic micrites that contain resedimented crinoid grains, sponge spicules, fish-phosphate, and fine terrigenous silt, reflecting hemipelagic deposition and early slope development. At the platform margins, such as Gozzano and Monte Fenera, reddish calcarenites and litharenites with basement-derived quartz and rhyolite mark true transgressive onlap onto exposed highs. These later give way to sponge-dominated carbonate mounds (locally “Broccatello-type”), reflecting a benthic, sponge-reef-dominated carbonate factory in deeper, low-light environments during progressive platform drowning.
Regional studies link this to platform drowning amid rifting, with carbon-isotope excursions implying volcanic influences and ocean perturbations. A modern analogue is the Bahama Banks, featuring oolitic shoals and lagoons in a subtropical passive-margin setting.
Osteno Lagerstätte
Apart from the Eocene of Monte Bolca, the Sinemurian of Osteno is the only fossil deposit in Italy in which soft bodies are preserved. The Osteno site was discovered in 1964. It was recovered from a series of 6 metres (20 ft) package of fine laminated, gray, spongiolitic, micritic limestone. Coroniceras bisulcatum allowed to date the outcrop as the Bucklandi zone, lower Sinemurian. The outcrop is a good documentation of a particularly complete fauna and flora of the Lower Jurassic which is not exactly common in the Southern Alps. The Osteno outcrop, part of the formation, is worldwide known due to the exceptional preservation of mostly marine biota, including rare fossilized components, helping to understand the ecosystems of the local Sinemurian margin of the Monte Generoso Basin. The high local variety of fossils found is most likely due to unique conditions of preservation, where phosphatized soft tissues have not been observed in any fish or polychaetes, but they are common in crustaceans (33%) and also occur in a smaller percentage of teuthids (14%). Soft part preservation through phosphatization in this deposit includes the muscles and branchia of Crustaceans, fish tissues, and the digestive tracts of coleoids, polychaetes, and nematodes. These fossils are interpreted as having been preserved in a stagnant, restricted basin with anoxic conditions likely within the sediment pore waters.
Biota
Foranimifera
| Taxa | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Ammobaculites | A. sp. | Castello Cabiaglio-Orino M. Sasso del Ferro | Tests | Simple agglutinated Ammomarginulininae benthic foraminifera | |
| Involutina | I. cf. liassica | Castello Cabiaglio-Orino | Tests | Early Jurassic Involutinidae, typical of outer-platform wackestones | |
| Lenticulina | L. sp. | Widespread in Alpe Perino & Moltrasio-Saltrio facies | Tests | Hyaline benthic foraminifera of Nodosariidae; typical in open-subtidal facies | |
| Reophax | R. sp. | Alpe Perino | Tests | Simple agglutinated benthic foraminifera of Reophacidae | |
| Siphovalvulina | S. sp. | Alpe Perino | Tests | Shallow-subtidal benthic foraminifera of Siphovalvulinidae | |
| Vidalina | V. martana | Alpe Perino Moltrasio-Saltrio facies | Tests | Early Jurassic benthic foraminifera of Involutinidae-like affinity; common on Liassic platforms |
Porifera
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Corynella | C. spp. | Broccatello of Arzo | Spicules & Imprints | A Calcareous sponge of the family Endostomatidae. | |
| Endostoma | E. spp. | Broccatello of Arzo | Spicules & Imprints | A Calcareous sponge of the family Endostomatidae. | |
| Stellispongia | S. spp. | Broccatello of Arzo | Spicules & Imprints | A Calcareous sponge of the family Stellispongiidae. | |
| Neuropora | N. cf. undulata N. spp. | Cava Salnova quarry Galli Quarry | Spicules & Imprints | A Demosponge of the family Neuroporidae. A notorious reef developing genus |
Brachiopoda
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Arzonellina | A. exotica | Broccatello of Arzo | Shells | A Terebratulidan of the family Arzonellinidae. | |
| Aulacothyris | A. waterhousi | Broccatello of Arzo | Shells | A Terebratulidan of the family Zeilleriidae. | Aulacothyris |
| Cirpa | C. fronto C. briseis C. variabilis | Broccatello of Arzo Cava Salnova quarry | Shells | A Rhynchonellidan of the family Wellerellidae. Identified originally as "Rhynchonella variabilis". | |
| Furcirhynchia | F. cf. melvillei | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Rhynchonellidae. | |
| Liospiriferina | L. rostrata | Broccatello of Arzo Cava Salnova quarry | Shells | A Rhynchonellidan of the family Spiriferinidae. Was identified originally as "Spiriferina haasi". | Liospiriferina specimens |
| Lobothyris | L. punctata L. andleri L. subpunctata L. ovatissinia L. sp. | Broccatello of Arzo Cava Salnova quarry Galli Quarry | Shells | A Terebratulidan of the family Lobothyrididae. Was identified originally as "Terebratula punctata". | Lobothyris |
| Prionorhynchia | P. belemnitica P. greppini P. flabellum P. ssp. | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Prionorhynchiidae. | |
| Rimirhynchia | R. aff. anglica | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Rhynchonellidae. | |
| Rhynchonella | R. acanthica R. binodosa R. belemnitica R. latifrons R. rimata R. cf.polyptycha | Broccatello of Arzo Cava Salnova quarry Galli Quarry | Shells | A Rhynchonellidan of the family Wellerellidae. | Rhynchonella |
| Rhynchonellina | R. alpina R. ssp. | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Dimerellidae. | |
| Tetrarhynchia | T. spp. | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Tetrarhynchiidae. | |
| Terebratula | T. scissa T. rhomboedrica T. prumus | Cava Salnova quarry Galli Quarry | Shells | A Terebratulidan of the family Terebratulidae. | |
| Spiriferina | S. expansa S. hartmanni S. oxigona S. walcotti S. rupestris S. rostrata S. haasi S. tumida S. cf. spirigeroides | Broccatello of Arzo Cava Salnova quarry | Shells | A Rhynchonellidan of the family Spiriferinidae. | Spiriferina |
| Sulcirostra | S. alpina | Broccatello of Arzo | Shells | A Rhynchonellidan of the family Dimerellidae. | |
| Viallithyris | V. gozzanensis V. adnetensis | Broccatello of Arzo Cava Salnova quarry | Shells | A Rhynchonellidan of the family Rhynchonellidae. | |
| Zeilleria | Z. sarthacensis Z. choffati Z. cor Z. ewaldi Z. stapia Z. cornuta Z. subnumismalis Z. (Zeilleria) indentata | Broccatello of Arzo Cava Salnova quarry Galli Quarry | Shells | A Terebratulidan of the family Zeilleriidae. | Zeilleria |
Bryozoa
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Ceriopora | C.? cf. orbignyi | Cava Salnova quarry Galli Quarry | Imprints | A Cyclostomatidan of the family Cerioporidae. |
Nematoida
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Eophasma | E. jurasicum | Osteno | Six specimens with soft parts | A nematode, type member of the family Eophasmidae. A rare find, since nematode fossils are absent in most Mesozoic marine rocks. | Eophasma |
Annelida
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Melanoraphia | M. maculata | Osteno | Eleven specimens, complete with soft parts | A polychaete, member of the group Phyllodocemorpha with incertae sedis assignation, suggested to be a member of Lumbrineridae. | Melanoraphia |
| Terebella | T. lapilloides T. spp. | Broccatello of Arzo Cava Salnova quarry | Tubiform structures | A terebellid, member of the family Terebellidae | Terebella specimen |
Arthropoda
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Aeger | A. foesteri A. muensteri A. robustus A. rostrospinatus A. elongatus A. macropus | Osteno | Multiple specimens fairly preserved | A decapod of the family Aegeridae. This genus is the most abundant local crustacean, and was a shrimp-like creature that was probably necrophagous. | Aeger foersteri |
| Coleia | C. vialiii C. medilerranea C. cf. antiqua C. pinnnai | Osteno | Multiple specimens fairly preserved | A decapod of the family Coleiidae. Probably a bottom dweller predatory crustacean. | Coleia viallii |
| Eryma | E. sinemuriana | Osteno | 41 specimens, fairly preserved | A decapod of the family Erymidae. The species was originally identified as Pustulina sinemuriana. | |
| Glyphea | G. tricarinata | Osteno | 137 complete and fragmentary specimens | A decapod of the family Erymidae. | |
| Mecochirus | M. germari | Osteno | 81 complete and fragmentary specimens, | A decapod of the family Mecochiridae. | Mecochirus germari |
| Ostenocaris | O. cypriformis | Osteno | Multiple specimens, complete and incomplete | A thylacocephalan of the family Ostenocarididae. It is the most common fossil of the formation, and the main identified thylacocephalan from the formation. | Ostenocaris |
| Ostenolimulus | O. latus | Osteno | A nearly complete specimen | A horseshoe crab of the family Limulidae. This genus represents the oldest Jurassic limulid and only the second xiphosuran known from Italy. Also indicates that this family were a diverse and widespread clade during the Jurassic. | |
| Ostenosculda | O. teruzzii | Osteno | Single Specimen fairly preserved | An early unipeltatan mantis shrimp | |
| Palaeastacus | P. meyeri | Osteno | Multiple specimens fairly preserved | A decapod of the family Erymidae. The species was originally included in the genus Eryma as E. meyeri, although the specimens reveal morphological characters diagnostic of Palaeastacus. | |
| Phlyctisoma | P. sinemuriana | Osteno | 10 specimens, in a fairly good state of preservation | A decapod of the family Erymidae. The species was originally identified as Pustulina sinemuriana. | |
| Pseudoglyphea | P. ancylochelis | Osteno | 7 complete specimens, in a fairly good state | A decapod of the family Litogastroidae. Probably a bottom dweller predatory crustacean. | |
| Teruzzicheles | T. popeyei | Osteno | Various specimens fairly preserved | A decapod of the family Polychelidae |
Mollusks
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Agassiceras | A. nodosaries A. aff. scipionianum | Cava Salnova quarry | Shells | An Ammonitidan of the family Arietitidae. | |
| Angulaticeras | A. sp. | Osteno | Shells | An ammonitidan of the family Schlotheimiidae. | |
| Anticonulus | A. profunde-umbilicatus | Cava Salnova quarry Galli Quarry | Shells | A Snail of the family Trochidae. | |
| Arietites | A. bucklandi A. raricostatus A. rapidecrescens A. ceratitoides A. kridioides A. dimorphus A. arnoui A. conybeari A. sauzeanus | Cava Salnova quarry Osteno | Shells | An Ammonitidan of the family Arietitidae. | Arietites bucklandi |
| Arnioceras | A. cf. bodleyi | Cava Salnova quarry Galli Quarry | Shells | An Ammonitidan of the family Arietitidae. | |
| Astarte | A. praeobliqua | Cava Salnova quarry | Shells | A Clam of the family Astartidae. Some shells identified as Cardium probably belong to this genus. | Astarte |
| Asteroceras | A. obtusum A. stellare A. retusum A. confusum | Cava Salnova quarry Burgioli quarry Galli quarry | Shells | An Ammonitidan of the family Arietitidae. | Asteroceras |
| Aulacoceras | A. ("Ausseites") italicus | Cava Salnova quarry Galli Quarry | Phagmocones | A Belemnite of the family Aulacoceratidae. | |
| Avicula | A. sinemuriensis A. (Oxytoma) dumortieri | Cava Salnova quarry Galli Quarry | Shells | A Pearl Oyster of the family Pteriidae. | |
| Bathrotomaria | B. cf. trocheata | Cava Salnova quarry Galli Quarry | Shells | A Snail of the family Pleurotomariidae. | |
| Calvaentolium | C. hehlii | Cava Salnova quarry | Shells | A Scallop of the family Pectinoidae. It was identified originally as "Pecten (Pseudamusium) hehlii". | |
| Cardinia | C. hybrida C. similis C. rugosa C. hybrida C. concinna C. depressa C. spp. | Cava Salnova quarry Galli Quarry | Shells | A Clam of the family Cardiniidae. | |
| Cardium | C.? italicum C. cf. multicostatum C. spp. | Cava Salnova quarry | Shells | A Clam of the family Cardiniidae. | |
| Cenoceras | C. amorettii C. stoppanii C. intermedium C. arare C. spreaficoi C. breislacki C. striatum C. sturi C. balsamocrivellii C. affinis C. brancoi C. ornatus | Cava Salnova quarry Galli Quarry | Shells | A Nautilidan of the family Cenoceratidae. Cenoceras was identified as member of the genus Nautilus originally. | Cenoceras reconstruction |
| Coroniceras | C. cf.gmuendense C. rotiformis C. bucklandi C. orbiculatus C. bisulcatus | Cava Salnova quarry Osteno | Shells | An Ammonitidan of the family Arietitidae. | Coroniceras specimen |
| Crucilobiceras | C. cf. tardarmatum | Galli Quarry | Shells | An Ammonitidan of the family Eoderoceratidae. | |
| Ectocentrites | E. sp. | Osteno | Shells | An ammonitidan of the family Lytoceratidae. | |
| Entolium | E. hehli E. calvum E. disciforme | Cava Salnova quarry Galli Quarry | Shells | A Scallop of the family Pectinoidae. | |
| Eparietites | E. impendens | Cava Salnova quarry Burgioli quarry Galli Quarry | Shells | An Ammonitidan of the family Arietitidae | |
| Euasteroceras | E. cf.turneri | Cava Salnova quarry Galli Quarry | Shells | An Ammonitidan of the family Arietitidae | |
| Chlamys | C. textoria | Cava Salnova quarry Galli Quarry | Shells | A Scallop of the family Pectinoidae It was identified as "Pecten (Chlamys) textorius". | Chlamys |
| Coelostylina | C. lepontina | Cava Salnova quarry Galli Quarry | Shells | A Snail of the family Coelostylinidae. | |
| Discohelix | D. sp. | Broccatello of Arzo | Shells | A Snail of the family Discohelicidae. | Discohelix |
| Discotoma | D.? aff. ornata | Cava Salnova quarry | Shells | A Snail of the family Pleurotomariidae. | |
| Fimbria | F. semireticulata F. (Sphaeriola) sp. | Cava Salnova quarry | Shells | A Clam of the family Lucinidae. | Fimbria |
| Gresslya | G. lunulata | Cava Salnova quarry Galli Quarry | Shells | A Clam of the family Pleuromyidae. | |
| Goniomya | G. verbana | Cava Salnova quarry | Shells | A Clam of the family Lucinidae. | |
| Gryphaea | G. arcuata | Cava Salnova quarry | Shells | An Oyster of the family Gryphaeidae. | Fimbria |
| Lima | L. (Radula) succincta L. (Radula) valmariannae L. (Radula) hettangiensis L. spp. | Broccatello of Arzo Cava Salnova quarry | Shells | A File Clam of the family Limidae. | Lima |
| Lucina | L. ? liasina | Cava Salnova quarry | Shells | A Clam of the family Lucinidae. | Lucina |
| Lytoceras | L. sp. | Galli Quarry | Shells | An Ammonitidan of the family Lytoceratidae | |
| Mactromya | M. arenacea M. cingulata M. problematica | Cava Salnova quarry Galli Quarry | Shells | An Adapedont of the family Edmondiidae. | |
| Microderoceras | M. birchii | Cava Salnova quarry | Shells | An Ammonitidan of the family Eoderoceratidae. | Microderoceras specimen |
| Modiolus | M. vomer M. cf.scalpra | Cava Salnova quarry Galli Quarry | Shells | A Mussel of the family Mytilidae. Identified as the genus "Modiola", now junior synonym of Modiolus. | Modiolus |
| Myoconcha | M. scabra M. rugosa M. ssp. | Cava Salnova quarry Galli Quarry | Shells | A Clam of the family Pleuromyidae. | |
| Nannobelus | N. cf. acutus | Cava Salnova quarry Galli Quarry | Phagmocones | A Belemnite of the family Belemnitidae. | |
| Ostenoteuthis | O. siroi | Osteno | Several specimens with decent preservation | A coleoid of the family Ostenoteuthidae. A unique group of coeloids, only found at this location. | Ostenoteuthis |
| Ostrea | O. cf. chillyensis O. electra | Broccatello of Arzo Cava Salnova quarry | Shells | An Oyster of the family Ostreidae. | Ostrea |
| Oxynoticeras | O. oxynotum O. actinotum | Cava Salnova quarry Burgioli quarry Galli Quarry Osteno Outcrop | Shells | An Ammonitidan of the family Oxynoticeratidae. | Oxynoticeras specimen |
| Paltechioceras | P. (Plesechioceras) demissum P.(Plesechioceras) delicatum | Burgioli quarry | Shells | An Ammonitidan of the family Echioceratidae. | |
| Paracoroniceras | P. cf. gmuendense | Cava Salnova quarry | Shells | An Ammonitidan of the family Arietitidae. | |
| Paradasyceras | P. stella | Cava Salnova quarry Galli Quarry | Shells | An Ammonitidan of the family Juraphyllitidae. | |
| Parallelodon | P. (Grammatodon) cypriniforme | Cava Salnova quarry Galli Quarry | Shells | A Clam of the family Parallelodontidae. | |
| Pecten | P. (Pseudamusium) diblasii P. (Pseudamusium) hehlii P. (Chlamys) subalpinus P. (Chlamys) textorius P. spp. | Broccatello of Arzo Cava Salnova quarry | Shells | A Scallop of the family Pectinoidae. | Pecten |
| Pholadomya | P. sp. | Cava Salnova quarry | Shells | A Clam of the family Pholadomyidae. | Pholadomya |
| Plagiostoma | P. giganteum P. stabilei P. spp. | Broccatello of Arzo Cava Salnova quarry | Shells | A File Clam of the family Limidae. Identified originally as "Lima (Plagiostoma) gigantea". | Plagiostoma |
| Pleuromya | P. cf. angusta P. saltriensis P. galathea | Cava Salnova quarry Galli Quarry | Shells | A Clam of the family Pleuromyidae. | |
| Pleurotomaria | P. oblita P. anglica P. escheri P. granulatocincta P. hettangiensis P. saltriensis P. cf. gigas P. sp. | Broccatello of Arzo Cava Salnova quarry | Shells | A Snail of the family Pleurotomariidae. | Pleurotomaria |
| Promicroceras | P. planicosta | Burgioli quarry | Shells | An Ammonitidan of the family Eoderoceratidae. | |
| Ptychomphalus | P. expansus | Cava Salnova quarry | Shells | A Snail of the family Eotomariidae. | |
| Pyrgotrochus | P. basilicus P. cf.basilicus | Cava Salnova quarry Galli Quarry | Shells | A Snail of the family Pleurotomariidae. | |
| Schlotheimia | S.(Charmasseiceras?) sp. | Galli Quarry | Shells | An Ammonitidan of the family Schlotheimiidae | |
| Terquemia | T. heberti | Cava Salnova quarry | Shells | A Clam of the family Prospondylidea. | |
| Trochus | T. sp | Broccatello of Arzo Cava Salnova quarry | Shells | A Snail of the family Trochidae. | Trochus |
| Trochotoma | T. subturrita | Cava Salnova quarry | Shells | A Snail of the family Pleurotomariidae. | |
| Uncinoteuthis | U. cuvieri | Osteno | An incomplete specimen consisting of the arm crown only | A coleoid of the family Ostenoteuthidae. | Uncinoteuthis |
| Vermiceras | V. spiratissimum V. cf.spiratissimum | Cava Salnova quarry | Shells | An Ammonitidan of the family Arietitidae. | Vermiceras specimen |
Echinoderms
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Hispidocrinus | H. scalaris | Pradalunga | Multiple ossicles | A Crinoidean, member of the family Isocrininae inside Isocrinida. | |
| Isocrinus | I. tuberculatus I. spp. | Broccatello of Arzo Cava Salnova quarry M. Campo dei Fiori Ubiale-Clanezzo | Multiple ossicles | A Crinoidean, member of the family Isocrininae inside Isocrinida. | |
| Millericrinus | M. cf. adneticus | M. di Grone | Multiple ossicles | A Crinoidean, member of the family Millericrinidae inside Millericrinida. | |
| Miocidaris | M. amalthei | Cava Salnova quarry | Spines & Ambulacrum | An Echinoidean of the family Miocidaridae. | |
| Palaeocoma | P. milleri | Mount Campo dei Fiori Ubiale-Clanezzo | MSNVI 044/017, dorsally-ventrally oriented ophiuroid | An Ophiuridan of the family Ophiodermatidae. Extant tropical species like Ophioderma are benthic predators and scavengers that show the same short spines seen in Palaeocoma. | Palaeocoma milleri |
| Pentacrinites | P. fasciculosus P. tuberculosus P. subangularis P. spp. | Broccatello of Arzo Cava Salnova quarry | Multiple ossicles | A Sea lily of the family Pentacrinitidae. | Pentacrinites |
| Plegiocidaris | P. falsani | Cava Salnova quarry Galli Quarry | Spines & Ambulacrum | An Echinoidean of the family Cidaridae. | Plegiocidaris |
Hemichordata
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Megaderaion | M. sinemuriense | Osteno | Six specimens with soft parts | An acorn worm, member of the family Harrimaniidae inside Enteropneusta. This genus was probably benthic, linked with burrowing systems. | Modern relative, Saccoglossus |
Chondrichthyes
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Hybodontiformes | Indeterminate | Osteno | Teeth | A shark, member of the family Hybodontiformes. | |
| Myriacanthidae | Gen et sp. nov | Osteno | Virtually complete skull with associated parts of the branchial skeleton | A chimaerid, member of the family Myriacanthidae inside Chimaeriformes. | |
| Notidanoides | N. arzoensis | Broccatello of Arzo Cava Salnova quarry | Teeth | A Crassodontidanidae Hexanchiform | |
| Ostenoselache | O. stenosoma | Osteno | One complete articulated specimen | A euselachiian of the family Ostenoselachidae. A genus only found in the Osteno Formation. With an elongated eel-shaped morphology, the specimens found measure at least 240 mm in length, with a neurocranium with a long, complex rostrum. It has been suggested it was an animal able to produce electric shocks, probably to kill prey. | Ostenoselache |
| Palaeospinax | P. pinnai | Osteno | A single specimen in four contiguous pieces and including parts of the dentition and squamation | A shark of the family Palaeospinacidae. | |
| Squaloraja | S. sp. | Osteno | One complete articulated specimen | A chimaerid of the family Squalorajidae. This genus belongs to a lineage of ray-like chimaeras. | Squaloraja |
| Sphenodus | S. helveticus S. cf. alpinus S. stschurowskii | Broccatello of Arzo Cava Salnova quarry | Teeth | An Orthacodontidae Synechodontiform |
Actinopterygii
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Dapedium | D. sp. | Osteno | MSNM V620 -MSNM V630, caudal fin with associated scales | Holostei of the family Dapediidae. | Dapedium |
| Dorsetichthys | D. bechei | Osteno | MSNM V409-MSNM V638: nearly complete articulated skeleton; MSNM V547-575: patch of articulated scales; MSNM V558: two isolated head skeletons; MSNM V572: poorly preserved head skeleton; MSNM V573: isolated caudal and caudal peduncle; MSNM V595: incomplete head skeleton; MSNM V604: patch of articulated scales; MSNM V629: nearly complete articulated skeleton; MSNM V633: incomplete articulated skeleton; MSNM V650: isolated head skeleton, in part and counterpart; MSNM V654: patch of articulated scales. MSNM V658: largely incomplete articulated skeleton; MSNM V662: partially complete articulated skeleton; MSNM V664: nearly complete articulated skeleton. | A member of the family Dorsetichthyidae. It is the main fish recovered on the formation. | Dorsetichthys |
| Furo | F. ("Eugnathus") sp. | Osteno | MSNM V623, an incomplete articulated skeleton; MSNM V634, an incomplete partially articulated skeleton. | A member of the family Furidae. | Furo |
| Ostenolepis | O. marianii | Osteno | MSNM V617, an almost complete, articulated skeleton; MSNM V538, largely incomplete specimen; MSNM V554, incomplete articulated skeleton; MSNM V652, incomplete twisted articulated skeleton; MSNM V618-19, skull & toracic section specimen. Other 3 Specimens | A member of the family Palaeoniscimorpha. Previously referred to Cosmolepis and Pteroniscus. | Ostenolepis |
| Peripeltopleurus | P. jurassicus | Osteno | MSNM V659, almost complete specimen lacking the tail | A member of the family Wushaichthyidae. Represents the only known Jurassic survivor of the order Peltopleuriformes | Peripeltopleurus |
| Pholidolepis | P. teruzzii | Osteno | MSNM V564-V621: almost complete, articulated skeleton. MSNM V529, MSNM V562, MSNM V533, MSNM V565, MSNM V596, MSNM V645, MSNM V648, MSNM V581, MSNM V649: isolated caudals; MSNM V540: incomplete articulated skeleton; MSNM V556, MSNM V559, MSNM V567, MSNM V568, MSNM V579, MSNM V594: isolated head skeleton; MSNM V563: disarticulated and largely incomplete skeleton; MSNM V570: incomplete articulated skeleton; MSNM V613: almost com-plete articulated skeleton; MSNM V625: incomplete articulated skeleton; MSNM V641: complete, articulated skeleton; MSNM V646: incomplete articulated skeleton MSNM V656: isolated pectoral girdle. | A member of the family Pholidophoridae. | Pholidolepis |
Sarcopterygii
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Holophagus | H. cf. gulo | Osteno | MSNM V530, V561, V587, V586, V588, V589, V591, V592, V593, V597, V598, V601, V599, V600, V667, V602, V607, V666, V668, isolated scales; MSNM V543, isolated urohyal. | A Coelacanth of the family Latimeriidae. Isolated 20 mm scales suggests a taxon up to 75 cm long. | Holophagus |
Icthyosaurs
In 2016 new vertebrate remains were discovered in the Salnova quarry, the remains are being studied to understand if it is a new dinosaur or some other creature. Latter has been confirmed to be Marine Diapsid material.
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Ichthyosaurus | I. spp. | Galli Quarry | 3 Vertebrae Imprints | A Neoichthyosaurian of the family Ichthyosauridae. | Ichthyosaurus |
| Temnodontosaurus | T. cf.platyodon | Cava Salnova quarry | Isolated Tooth Isolated Vertebrae | A Neoichthyosaurian of the family Temnodontosauridae. Quoted on the 1880s, specimen that apparently has never been described or figured and whose present repository is unknown | Temnodontosaurus platyodon |
Pterosaurs
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Pterosauria | Indeterminate | Cava Salnova quarry | Isolated Tooth | A Pterosaur, referred originally to "Pterodactylus longirostris" | Bergamodactylus, an older (Triassic) Pterosaur from Italy |
Dinosaurs
| Genus | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Saltriovenator | S. zanellai | Cava Salnova quarry | MSNM V3664, very fragmentary and disarticulated skeleton MSNM V3659, one maxillary or dentary tooth (Referred material) | A potential Ceratosaur. Traces on the bones show that the dinosaur carcass remained exposed to the water-sediment interface for months or years, long enough to first be defleshed by mobile scavengers, then colonized by a microbial community that spanned the bone–water interface, which in turn attracted slow-moving grazers and epibionts. | Saltriovenator holotype |
Algae
Potential Dasycladaceae are seen on intertidal facies, while Characeae are seen at the base.
| Taxa | Species | Locality | Material | Notes | Images |
|---|---|---|---|---|---|
| Cayeuxia | C. spp. | Alpe Perino; Castello Cabiaglio-Orino | Imprints | Calcareous green alga of Udoteaceae; typical of shallow subtidal facies | |
| Thaumatoporella | T. spp. | Alpe Perino Limestone | Imprints | Encrusting calcified alga, typical of Liassic shallow-marine platforms |
Plants
| Genus | Species | Location | Material | Notes | Images |
|---|---|---|---|---|---|
| Brachyphyllum | B. saportum B. spp. | Osteno Dintorni di Moltrasio | Branched Shoots | A coniferophytan of the family Cheirolepidiaceae. | |
| Callialasporites | C. spp. | Castello Cabiaglio-Orino | Pollen | Araucariaceae Conifer | Extant Araucaria. |
| Cerebropollenites | C. macroverrucosus | Castello Cabiaglio-Orino | Pollen | Sciadopityaceae Conifer | Extant Sciadopitys. |
| Classopollis | C. torosus | Castello Cabiaglio-Orino | Pollen | Cheirolepidiaceae Conifer, dominant palynomorph | |
| Clathropteris | C. platyphylla | Carate-Lario | Fronds | A Fern of the family Dipteridaceae | |
| Concavisporites | C. crassexinius | Castello Cabiaglio-Orino | Spores | Trilete spore of Lycophyte affinity | |
| Ctenopteris | C. grandis | Moltrasio | Fronds | A Fern of the family Polypodiaceae | |
| Equisitites | E. bunburyanus | Osteno | Stems | A Horsetail of the family Equisetaceae. | |
| Gliscopollis | C. meyeriana | Castello Cabiaglio-Orino | Pollen | Typical of Jurassic-Cretaceous Cheirolepidiaceae | |
| Granulatisporites | G. punctatus | Castello Cabiaglio-Orino | Spores | Trilete spore of Lycopodiales | |
| Lomatopteris | L. jurensis | Carate-Lario | Fronds | A "Seed Fern", affinities with Cyclopteridaceae. | Lomatopteris jurensis |
| Lycopodiacidites | L. rugulatus | Castello Cabiaglio-Orino | Spores | Trilete spore of Lycopodiaceae | Extant Lycopodium. |
| Microreticulatisporites | M. fuscus | Castello Cabiaglio-Orino | Spores | Trilete spore of probable Lycophyte origin | |
| Otozamites | O. bunburyanus O. bechei O. latior O. brevifolius O. pterophylloides O. sp. | Osteno Moltrasio Carate-Lario | Leaflets | A spermatopsidan, member of Bennettitales. | |
| Pachypteris | P. cf. rhomboidalis | Osteno | Fronds | A "Seed Fern" of the family Corystospermaceae. | |
| Pagiophyllum | P. peregrinum P. sp. | Osteno Carate-Lario | Branched Shoots | A coniferophytan of the family Araucariaceae. | |
| Porcellispora | P. longdonensis | Castello Cabiaglio-Orino | Spores | Affinities with Bryophyta or Marchantiophyta, potentially Riellaceae. | |
| Thinnfeldia | T. rhomboidalis T. obtusa | Carate-Lario | Fronds | A "Seed Fern" of the family Corystospermaceae. | |
| Uvaesporites | U. argenteaeformis | Castello Cabiaglio-Orino | Spores | Trilete spore of Selaginellaceae. | Extant Selaginella. |
| Williamsonia | W. sp. | Osteno | Reproductive Organ | A spermatopsidan, member of Bennettitales. | Williamsonia |
| Widdringtonites | W. baehmanni | Mombello | Branched Shoots | A coniferophytan of the family Araucariaceae. | |
| Zamites | Z. sp. | Osteno | Leaflets | A spermatopsidan, member of Bennettitales. |
See also
- List of fossiliferous stratigraphic units in Italy
- Calcare di Sogno, Toarcian fossiliferous formation of Lombardy
- Rotzo Formation, Pliensbachian fossiliferous formation of Veneto
- Coimbra Formation, Sinemurian fossiliferous formation of Portugal
- Aganane Formation, Pliensbachian Formation of Morocco